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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1380	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Rhinolophus landeri [synonym of]	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis	Rhinolophus guineensis		[MSW2] Originally described as a subspecies of R. landeri, but see Bohme and Hutterer (1979) who demonstrated that it was a separate species.; [MSW3] landeri species group. Originally described as a subspecies of landeri, but see Böhme and Hutterer (1979), who demonstrated that it is a separate species.; [HMW] Rhinolophus landeri guineensis Eisentraut , 1960 , near Tahiré, base of Kelesi Plateau , 500 m , Guinea . Rhinolophus guineensis is in the landeri species group based on morphological data, although genetic data are needed to confirm this placement. It was originally described as a subspecies of R landeri but is now recognized as a distinct species. Monotypic.; [batnames2022]  landeri species group. Originally described as a subspecies of landeri, but see Böhme and Hutterer (1979), who demonstrated that it is aseparate species.; [batnames2023]  landeri species group. Originally described as a subspecies of landeri, but see Böhme and Hutterer (1979), who demonstrated that it is aseparate species.; [batnames2025_1.7] landerispecies group. Originally described as a subspecies of landeri, but see Böhme and Hutterer (1979), who demonstrated that it is aseparate species.														guineensis				guineensis	guineensis			guineensis Eisentraut, 1960						N/A					Distribution: Known only from Senegal, Guinea, and Sierra Leone.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Senegal – Sierra Leone; ref. 4.30	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Eisentraut	1960	Stygg. Beitr. Naturk., 39:1.	Originally described as a subspecies of R. landeri, but see Bohme and Hutterer (1979) who demonstrated that it was a separate species.	Guinea, Sierra Leone, Liberia.	Guinea, Tahir6 (foot of Kelesi Plateau).		EISENTRAUT	1960	Connecting process triangular in profile. First phalanx of fourth digit of wing shortened. Size relatively large (forearm length, 44-49 mm), but skull relatively small (total length, 20-22 mm; maxillary toothrow length, 7.2-7.8 mm).	Distribution: Known only from Senegal, Guinea, and Sierra Leone.	No subspecies.		55	species	R. guineensis	EISENTRAUT	1960	Rhinolophus	genus	Rhinolophus guineensis				Connecting process triangular in profile. First phalanx of fourth digit of wing shortened. Size relatively large (forearm length, 44-49 mm), but skull relatively small (total length, 20-22 mm; maxillary toothrow length, 7.2-7.8 mm).	No subspecies.		27. R. guineensis EISENTRAUT 1960 [pusillus group].	27	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus guineensis	Rhinolophus		guineensis	Eisentraut		1960		Stuttg. Beitr. Naturk.	39		1		Guinean Horseshoe Bat	Guinea, Tahiré (foot of Kelesi Plateau).	Senegal, Guinea, Sierra Leone, Liberia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		landeri species group. Originally described as a subspecies of landeri, but see Böhme and Hutterer (1979), who demonstrated that it is a separate species.	885887A2FFC58A23FF55FB73F7BBDD0B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	282	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFCC8A2DF84EFAC8FEA3DB68.xml	Rhinolophus guineensis	Rhinolophidae	Rhinolophus	guineensis	Eisentraut	1960	Guinean Horseshoe Bat @en | Rhinolophede Guinée @fr | Guinea-Hufeisennase @de | Herradura de Guinea @es | SenegalHorseshoe Bat @en | Other commonnames @en	Rhinolophus landeri guineensis Eisentraut , 1960 , near Tahiré, base of Kelesi Plateau , 500 m , Guinea . Rhinolophus guineensis is in the landeri species group based on morphological data, although genetic data are needed to confirm this placement. It was originally described as a subspecies of R landeri but is now recognized as a distinct species. Monotypic.	Known from scattered records in S Senegal (Niokolo-Koba National Park and Diattacounda ), SW & SE Guinea (near Kindia, Simandou Range, and Mt Nimba), C Sierra Leone , N Liberia (including Liberian Mt Nimba ), and E Ivory Coast ; it is potentially present in Gambia in patches of relict forest.	Head-body c. 49-57 mm , tail 23-30 mm , ear 17-22 mm , hindfoot 8-10 mm , forearm 44-50 mm ; weight 8-11 g . The Guinean Horseshoe Bat is similar to Lander’s Horseshoe Bat ( AE . landeri ), but is larger in craniodental dimensions with wider horseshoe. Pelage is variable, with dorsum varying from brown to grayish brown, orangish brown, or pale reddish brown (individual hairs with darker tips than bases); venter is always slightly paler. Axillary tufts of males are generally white, but there is one case in which they were reddish brown. Ears are comparatively short (38—47% of forearm length) and have 11—12 internal folds. Noseleaf has hastate lancet; connecting process is subtriangular, with pointed tip; sella is naked and parallel-sided; horseshoe is narrow at 8-3-9- 3 mm ; there are no lateral leaflets; and median emargination is present. Lower lip has one groove. Wings and uropatagium are dark gray, and first phalanx of fourth finger is relatively short. Skull is medium built; zygomatic arches are moderately broad; zygomatic width is slightly larger than mastoid width; palatal bridge is 26-31% of C 1- M 3 length; P2 is comparatively large and is in tooth row, being slightly displaced labially; C1 and P4 are well separated; P3 is very small to virtually vestigial and is fully displaced labially; and P 2 and P4 are in contact.	Primarily highland regions with submontane to montane forests in Guinea , Liberia , and Ivory Coast and, to a lesser extent, lowland rainforest and forest savanna mosaic habitats in Senegal and Guinea . In Makeni, Sierra Leone , Guinean Horseshoe Bats have been recorded in disturbed areas with small patches of relict forest. They are generally found at higher elevations and have been recorded at elevations of 1400 m and above. On Guinean and Liberian Mt Nimba, they have been recorded in montane grasslands at 900-1600 m , where they are considerably common during dry seasons.	No information.	Single embryos in Guinean Horseshoe Bats have been recorded in December in the Simandou Range, Guinea , and March near Kindia, Guinea .	Guinean Horseshoe Bats are nocturnal. Two torpid individuals were captured on Mount Nimba, Guinea . They have been recorded roosting in caves throughout their distribution, although two individuals were found in hollow trees in Senegal . They have also been recorded roosting in mines. Call shape is FM/CF/ FM , with F component of 85 kHz in Guinea , which is significantly lower than in Lander’s Horseshoe Bat.	Roosts of the Guinean Horseshoe Bat are often shared with other species of Rhinolophus , Lissonycteris , Nycteris , and Hipposideros.	Classified as Vulnerable on The IUCNed List. The Guinean Horseshoe Bat has a constrained area of occupancy (probably less than 2000 km 2), severely fragmented distribution, and continuous decline in extent and quality of its forest and cave habitats. Major threats include deforestation resulting from logging operations, conversion of land to agricultural use, and mining activities. There is also limited threat of overhunting for bushmeat.	ACR (2018) | Böhme & Hutterer (1978) | Csorba et al. (2003) | Decher et al. (2016) | Denys et al. (2013) | Eisentraut (1960b) | ahr (2008d, 2013d) | ahr & Ebigbo (2003) | Grubb et al. (1998) | Konstantinov et al. (2000) | Monadjem , Richards & Denys (2016) | Rosevear (1965) | Taylor, Macdonald et al. (2018) | Weber & ahr (2007b)	https://zenodo.org/record/3749920/files/figure.png	5 . Guinean Horseshoe Bat Rhinolophus guineensis French: Rhinolophe de Guinée I German: Guinea-Hufeisennase / Spanish: Herradura de Guinea Other common names: Senegal Horseshoe Bat Taxonomy. Rhinolophus landeri guineensis Eisentraut , 1960 , near Tahiré, base of Kelesi Plateau , 500 m , Guinea . Rhinolophus guineensis is in the landeri species group based on morphological data, although genetic data are needed to confirm this placement. It was originally described as a subspecies of R landeri but is now recognized as a distinct species. Monotypic. Distribution. Known from scattered records in S Senegal (Niokolo-Koba National Park and Diattacounda ), SW & SE Guinea (near Kindia, Simandou Range, and Mt Nimba), C Sierra Leone , N Liberia (including Liberian Mt Nimba ), and E Ivory Coast ; it is potentially present in Gambia in patches of relict forest. Descriptive notes. Head-body c. 49-57 mm , tail 23-30 mm , ear 17-22 mm , hindfoot 8-10 mm , forearm 44-50 mm ; weight 8-11 g . The Guinean Horseshoe Bat is similar to Lander’s Horseshoe Bat ( AE . landeri ), but is larger in craniodental dimensions with wider horseshoe. Pelage is variable, with dorsum varying from brown to grayish brown, orangish brown, or pale reddish brown (individual hairs with darker tips than bases); venter is always slightly paler. Axillary tufts of males are generally white, but there is one case in which they were reddish brown. Ears are comparatively short (38—47% of forearm length) and have 11—12 internal folds. Noseleaf has hastate lancet; connecting process is subtriangular, with pointed tip; sella is naked and parallel-sided; horseshoe is narrow at 8-3-9- 3 mm ; there are no lateral leaflets; and median emargination is present. Lower lip has one groove. Wings and uropatagium are dark gray, and first phalanx of fourth finger is relatively short. Skull is medium built; zygomatic arches are moderately broad; zygomatic width is slightly larger than mastoid width; palatal bridge is 26-31% of C 1- M 3 length; P2 is comparatively large and is in tooth row, being slightly displaced labially; C1 and P4 are well separated; P3 is very small to virtually vestigial and is fully displaced labially; and P 2 and P4 are in contact. Habitat. Primarily highland regions with submontane to montane forests in Guinea , Liberia , and Ivory Coast and, to a lesser extent, lowland rainforest and forest savanna mosaic habitats in Senegal and Guinea . In Makeni, Sierra Leone , Guinean Horseshoe Bats have been recorded in disturbed areas with small patches of relict forest. They are generally found at higher elevations and have been recorded at elevations of 1400 m and above. On Guinean and Liberian Mt Nimba, they have been recorded in montane grasslands at 900-1600 m , where they are considerably common during dry seasons. Food and Feeding. No information. Breeding. Single embryos in Guinean Horseshoe Bats have been recorded in December in the Simandou Range, Guinea , and March near Kindia, Guinea . Activity patterns. Guinean Horseshoe Bats are nocturnal. Two torpid individuals were captured on Mount Nimba, Guinea . They have been recorded roosting in caves throughout their distribution, although two individuals were found in hollow trees in Senegal . They have also been recorded roosting in mines. Call shape is FM/CF/ FM , with F component of 85 kHz in Guinea , which is significantly lower than in Lander’s Horseshoe Bat. Movements, Home range and Social organization. Roosts of the Guinean Horseshoe Bat are often shared with other species of Rhinolophus , Lissonycteris , Nycteris , and Hipposideros. Status and Conservation. Classified as Vulnerable on The IUCNed List. The Guinean Horseshoe Bat has a constrained area of occupancy (probably less than 2000 km 2), severely fragmented distribution, and continuous decline in extent and quality of its forest and cave habitats. Major threats include deforestation resulting from logging operations, conversion of land to agricultural use, and mining activities. There is also limited threat of overhunting for bushmeat. Bibliography. ACR (2018), Böhme & Hutterer (1978), Csorba et al. (2003), Decher et al. (2016), Denys et al. (2013), Eisentraut (1960b), ahr (2008d, 2013d), ahr & Ebigbo (2003), Grubb et al. (1998), Konstantinov et al. (2000), Monadjem , Richards & Denys (2016), Rosevear (1965), Taylor, Macdonald et al. (2018), Weber & ahr (2007b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus guineensis	Rhinolophus		guineensis	Eisentraut	1960	0	Stuttg. Beitr. Naturk.	######	Guinean Horseshoe Bat	None.	Guinea, Tahiré (foot of Kelesi Plateau).	Senegal, Guinea, Sierra Leone, Liberia.	Not listed.	Endangered	 landeri species group. Originally described as a subspecies of landeri, but see Böhme and Hutterer (1979), who demonstrated that it is aseparate species.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus guineensis	23	Guinean Horseshoe Bat	Senegal Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	guineensis	Eisentraut	1960	0	Rhinolophus_landeri_guineensis	Eisentraut, M. (1960). Zwei neue Rhinolophiden aus Guinea. Stuttgarter Beiträge zur Naturkunde, 39, 1.	https://www.biodiversitylibrary.org/page/34022203#page/109/mode/1up	SMNS 6103		near Tahiré, base of Kelesi Plateau, 500 m, Guinea.			guineensis Eisentraut, 1960	NA	NA	Senegal|Guinea|Sierra Leone|Liberia|Côte d'Ivoire|Gambia?|Guinea-Bissau?	Africa	Afrotropic	EN	0	0	0	Rhinolophus_guineensis	0	sciname match	Rhinolophus_guineensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19542	Rhinolophus guineensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	guineensis	Eisentraut, 1960		20000000	Rhinolophus guineensis	Endangered	B2ab(i,ii,iii,v)	2020	2019-06-18 00:00:00 UTC	3.1	English	Rhinolophus guineensis is listed as Endangered under criterion B2 as its roost distribution is severely fragmented into approximately 20 cave/mine roosts that are critical resources where the bats roost for nearly 12 hours out of every day; these roosts are under great pressure. |Its area of occupancy (AOO) is estimated to be approximately 80 km² based on these 20 cave/mine roosts. Its extent of occurrence (EOO), AOO, habitat, and population are all projected to continue to decline due to ongoing threats from forest loss and degradation and cave/mine loss, degradation, and disturbance.	This species has been recorded from montane tropical moist forest, and to a lesser extent from moist savanna (Koopman 1989, Koopman et al.  1995, Grubb et al.  1998, Weber and Fahr 2007, Denys et al.  2013, Monadjem et al.  2016) or high-elevation grasslands (Decher et al.  2010, Denys et al.  2013, Monadjem et al.  2016). It appears to use riverine forests and forested ravines (Decher et al.  2010, Decher et al.  2015). Animals are usually found roosting in caves or mine adits, however, two were found in hollow trees (Böhme and Hutterer 1978, Grubb et al.  1998, Weber and Fahr 2007, Monadjem et al.  2016). In the Simandou Mountains of Guinea, this species was captured near large boulders, which it may have been using to roost (Decher et al.  2015). Pregnant females have been found in March and December.	Major threats to this species include deforestation resulting from logging operations, the conversion of land to agricultural use, and mining activities. The destruction of old mine adits threatens their roosts. There is also a limited threat of overhunting for the bushmeat trade.	Little information is available on the population abundance of this bat although its global population is thought to be in decline due to forest, cave, and mine degradation and loss. This species was the least numerous of four species of Rhinolophus and Hipposideros recorded roosting in caves and mine adits at 1,500 m asl at Mt Nimba, Guinea (Monadjem, unpublished data), although other surveys in the area have found R. guineensis at higher relative abundances (Monadjem et al. 2016). Studies generally capture few individuals (&lt;10) (Decher et al. 2010, Decher et al. 2015), although Denys et al. (2013) found 41 individuals from two different sites on the Guinean side of Mt. Nimba.	Decreasing	This West African bat has been patchily recorded from fewer than 20 localities (cave/mine roosts) in southern Senegal, Guinea, Sierra Leone, Liberia and Côte d'Ivoire. Rhinolophus guineensis is generally a highland species recorded at elevations of 900–1,500 m asl and over (Grubb et al. 1998, Denys et al. 2013, Decher et al. 2015, Monadjem et al. 2016), although it has also been found as low as 600 m asl (Decher et al. 2015). It is suspected to be present in Gambia in remnant patches of forest (Grubb et al. 1998) and in Guinea-Bissau.		Terrestrial	This species has been recorded from the Mount Nimba World Heritage Site (a portion of which is currently being mined on the Guinea side) and the "Massif du Ziama" Biosphere Reserve, both in Guinea. It has also been recorded from a few state forests ("Forêts Classées") in Guinea. There is a need to conserve remaining areas of suitable habitat for this species. Further studies are needed to better determine the species range.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		guineensis	Eisentraut	1960	0	Stuttg. Beitr. Naturk.	39:01:00	Guinean Horseshoe Bat	None.	Guinea, Tahiré (foot of Kelesi Plateau).	Senegal, Guinea, Sierra Leone, Liberia.	Not listed.	Endangered	 landeri species group. Originally described as a subspecies of landeri, but see Böhme and Hutterer (1979), who demonstrated that it is aseparate species.	Rhinolophus guineensis	1004689	23	Guinean Horseshoe Bat	Senegal Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	guineensis	Eisentraut	1960	0	Rhinolophus_landeri_guineensis	Eisentraut, M. (1960). Zwei neue Rhinolophiden aus Guinea. Stuttgarter Beiträge zur Naturkunde, 39, 1.	https://www.biodiversitylibrary.org/page/34022203#page/109/mode/1up	SMNS 6103		near Tahiré, base of Kelesi Plateau, 500 m, Guinea.			guineensis Eisentraut, 1960	NA	NA				Senegal|Guinea|Sierra Leone|Liberia|Côte d'Ivoire|Gambia?|Guinea-Bissau?	Africa	Afrotropic	EN	0	0	0	Rhinolophus_guineensis	0	sciname match	Rhinolophus_guineensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_guineensis	1004689	23	Guinean Horseshoe Bat	Senegal Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	guineensis	Eisentraut	0	Rhinolophus landeri guineensis	Eisentraut, M. 1960-06-01. Zwei neue Rhinolophiden aus Guinea. Stuttgarter Beiträge zur Naturkunde 39:1-7.	https://www.biodiversitylibrary.org/page/34022203	SMNS 6103	holotype		near Tahiré, base of Kelesi Plateau, 500 m, Guinea.			NA	NA				Senegal|Guinea|Sierra Leone|Liberia|Cote d'Ivoire|Gambia?|Guinea-Bissau?	Africa	Afrotropic	EN	0	0	0	Rhinolophus_guineensis	0	sciname match	Rhinolophus_guineensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		guineensis	Eisentraut	1960	0	Stuttg. Beitr. Naturk.	39:01:00	Guinean Horseshoe Bat	None.	Guinea, Tahiré (foot of Kelesi Plateau).	Senegal, Guinea, Sierra Leone, Liberia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19542/21980043/' target='_blank'>Endangered</a>	landerispecies group. Originally described as a subspecies of landeri, but see Böhme and Hutterer (1979), who demonstrated that it is aseparate species.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus guineensis; Rhinolophus guineensis; Rhinolophus guineensis; Rhinolophus guineensis; Rhinolophus guineensis; Rhinolophus guineensis; guineensis; Guinean Horseshoe Bat; Rhinolophede Guinée; Guinea-Hufeisennase; Herradura de Guinea; SenegalHorseshoe Bat; Other commonnames; Guinean Horseshoe Bat; Senegal Horseshoe Bat; Guinean Horseshoe Bat; Guinean Horseshoe Bat; R. guineensis