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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1371	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens	Rhinolophus eloquens		[MSW2] Includes perauritus; see Koopman (1975:389).; [MSW3] fumigatus species group. Includes perauritus; see Koopman (1975).; [HMW] Rhinolophus hildebrandti [sic] eloquens K. Andersen, 1905 , “ Entebbi [= Entebbe] , Uganda .” Rhinolophus eloquens is in the Junugatus species group and has been included as a subspecies of R hildebrandtii , although it is generally recognized as a distinct species. Rhinolophus eloquens is sister to the rest of the fumigatus group. Two subspecies recognized .; [batnames2022]  fumigatus species group. Includes perauritus; see Koopman (1975). May be conspecific with fumigatus Demos et al. (2019).; [IUCN] The taxonomic relationship between Rhinolophus eloquens and R. brandti needs clarification.; [batnames2023]  fumigatus species group. Includes perauritus; see Koopman (1975). May be conspecific with fumigatus Demos et al. (2019).; [batnames2025_1.7] fumigatus species group. Includes perauritus; see Koopman (1975). May be conspecific with fumigatus Demos et al. (2019).				perauritus		perauritus.	perauritus, eloquens	eloquens, perauritus		eloquens, perauritus		eloquens, perauritus		eloquens, perauritus	The taxonomic relationship between Rhinolophus eloquens and R. brandti needs clarification.	eloquens, perauritus		eloquens, perauritus	eloquens, perauritus	eloquens, perauritus		eloquens Andersen, 1905|perauritus De Beaux, 1922		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		S Sudan – S Somalia – N Tanzania	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhinolophus eloquens	Uganda, Entebbe.	K. Andersen	1905	Ann. Mag. Nat. Hist., ser. 7, 15:74.	Distribution: Ranging from south ern Somalia to eastern Zaire and south to north ern Tanzania (including Pemba and Zanzibar is lands).		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		S Sudan – S Somalia – N Tanzania	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	K. Andersen	1905	Ann. Mag. Nat. Hist., ser. 7, 15:74.	Includes perauritus; see Koopman (1975:389).	Uganda, S Somalia, S Sudan, NE Zaire, Kenya, Rwanda, N Tanzania, Zanzibar and Pemba.	Uganda, Entebbe.		ANDERSEN	1905	Anterior upper premolar greatly reduced and ex truded from toothrow. Fifth metacarpal of wing somewhat longer than fourth or third. Lateral mental grooves obliterated. Size fairly large (fore arm length, 54-60 mm; condylocanine length, 21-23 mm).	Distribution: Ranging from south ern Somalia to eastern Zaire and south to north ern Tanzania (including Pemba and Zanzibar is lands).	Two subspecies are recognized:	R. e. perauritus (southern Somalia), R. e. eloquens (south ern Sudan to northern Tanzania).	57	species	R. eloquens	ANDERSEN	1905	Rhinolophus	genus	Rhinolophus eloquens				Anterior upper premolar greatly reduced and ex truded from toothrow. Fifth metacarpal of wing somewhat longer than fourth or third. Lateral mental grooves obliterated. Size fairly large (fore arm length, 54-60 mm; condylocanine length, 21-23 mm).	Two subspecies are recognized:		45. R. eloquens ANDERSEN 1905 [luctus group].	45	_R. e. eloquens_ Andersen, 1905; _R. e. perauritus_ De Beaux, 1922			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus eloquens	Rhinolophus		eloquens	K. Andersen		1905		Ann. Mag. Nat. Hist., ser. 7	15		74		Eloquent Horseshoe Bat	Uganda, Entebbe.	Uganda, S Somalia, S Sudan, NE Dem. Rep. Congo, Kenya, Rwanda, N Tanzania, Zanzibar and Pemba.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	perauritus De Beaux, 1922.	fumigatus species group. Includes perauritus; see Koopman (1975).	885887A2FFD98A3FF887FCC8F71ED6C6	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	291	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFC38A25F89FFEFEF4B9D56F.xml	Rhinolophus eloquens	Rhinolophidae	Rhinolophus	eloquens	K. Andersen	1905	Eloquent Horseshoe Bat @en | Rhinolophe éloquent @fr | Lautfreudige Hufeisennase @de | Herradura elocuente @es	Rhinolophus hildebrandti [sic] eloquens K. Andersen, 1905 , “ Entebbi [= Entebbe] , Uganda .” Rhinolophus eloquens is in the Junugatus species group and has been included as a subspecies of R hildebrandtii , although it is generally recognized as a distinct species. Rhinolophus eloquens is sister to the rest of the fumigatus group. Two subspecies recognized .	A e. eloquens K Andersen, 1905 - S South Sudan , NE & SE DR Congo , Uganda , Rwanda , Kenya , and Tanzania (including Pemba and Unguja Is in Zanzibar Archipelago); possibly also S Ethiopia , extending as far as Malawi and Mozambique . R e. perauritus De Beaux, 1922 — S Somalia .	Head—body c . 68-72 mm , tail 20-45 mm , ear 21—38 mm , hindfoot 12—14 mm , forearm 53—63 mm ; weight 13-40 g . Dorsal pelage is generally pale brown (hairs are dark gray basally); venter is paler. There is no orange morph. Males lack axillary tufts. Ears are medium in length (44—51% of forearm length). Noseleaf has long subtriangular lancet, with slightly concave sides and rounded tip; connecting process is large and rounded, terminating near tip of sella; sella is covered with longish hairs and is broad and rounded at top, with upper two-thirds being parallel-sided; and horseshoe is medium in width (114—12- 3 mm ), covers muzzle, has lateral leaflets, and has deep median emargination. Lower lip has one groove. Wings and uropatagium are brown to dark grayish brown. Baculum is trumpet-shaped, with straight, cylindrical shaft. Skull is robust, with sturdy zygomatic arches (zygomatic width is much wider than mastoid width); nasal swellings are relatively high, being longer than they are broad; frontal depression is shallow to relatively deep; and sagittal crest is comparatively prominent anteriorly. Dental formula is commonly 11/2, C 1/1, P 1/2, M 3/3 (x2) = 28, or rarely 11/2, C 1/1, P 2/2, M 3/3 (x2) = 30,11/2, C 1/1, P 2/3, M 3/3 (x2) = 32, or 11/2, C 1/1, P 1/3, M 3/3 = 30. P2 is very small and completely displaced labially or completely absent, allowing full contact ofC1 and P4, and P3 is usually absent but, if present, is fully displaced labially and lies under cingula of P2 and P4, which are in contact.	Various semiarid savanna, mesic woodland savanna, and montane forest habitats, including Acacia ( Fabaceae )- Commiphora ( Burseraceae ) brushland/thicket and mosaics of evergreen brushland and secondary Acacia savanna, at elevations up to 1500 m .	No information.	No information.	The Eloquent Horseshoe Bat roosts in caves. There might be more information available in papers that misidentified it as Hildebrandt’s Horseshoe Bat {hildebrandtii ).	No information.	Classified as Least Concern on The IUCN Red List. The Eloquent Horseshoe Bat has a relatively wide distribution, but littie is known about its ecology. It seems to be most threatened by general habitat loss from agricultural expansion.	ACR (201'8) | Cotterill (2013b) | Csorba et al. (2003) | Koopman (1975) | Monadjem, Taylor , Jacobs & Cotterill (2017d) | Thorn & Kerbis Peterhans (2009) | Taylor et al. (2012)	https://zenodo.org/record/3749954/files/figure.png	23 . Eloquent Horseshoe Bat Rhinolophus eloquens French: Rhinolophe éloquent /German: Lautfreudige Hufeisennase /Spanish: Herradura elocuente Taxonomy. Rhinolophus hildebrandti [sic] eloquens K. Andersen, 1905 , “ Entebbi [= Entebbe] , Uganda .” Rhinolophus eloquens is in the Junugatus species group and has been included as a subspecies of R hildebrandtii , although it is generally recognized as a distinct species. Rhinolophus eloquens is sister to the rest of the fumigatus group. Two subspecies recognized . Subspecies and Distribution. A e. eloquens K Andersen, 1905 - S South Sudan , NE & SE DR Congo , Uganda , Rwanda , Kenya , and Tanzania (including Pemba and Unguja Is in Zanzibar Archipelago); possibly also S Ethiopia , extending as far as Malawi and Mozambique . R e. perauritus De Beaux, 1922 — S Somalia . Descriptive notes. Head—body c . 68-72 mm , tail 20-45 mm , ear 21—38 mm , hindfoot 12—14 mm , forearm 53—63 mm ; weight 13-40 g . Dorsal pelage is generally pale brown (hairs are dark gray basally); venter is paler. There is no orange morph. Males lack axillary tufts. Ears are medium in length (44—51% of forearm length). Noseleaf has long subtriangular lancet, with slightly concave sides and rounded tip; connecting process is large and rounded, terminating near tip of sella; sella is covered with longish hairs and is broad and rounded at top, with upper two-thirds being parallel-sided; and horseshoe is medium in width (114—12- 3 mm ), covers muzzle, has lateral leaflets, and has deep median emargination. Lower lip has one groove. Wings and uropatagium are brown to dark grayish brown. Baculum is trumpet-shaped, with straight, cylindrical shaft. Skull is robust, with sturdy zygomatic arches (zygomatic width is much wider than mastoid width); nasal swellings are relatively high, being longer than they are broad; frontal depression is shallow to relatively deep; and sagittal crest is comparatively prominent anteriorly. Dental formula is commonly 11/2, C 1/1, P 1/2, M 3/3 (x2) = 28, or rarely 11/2, C 1/1, P 2/2, M 3/3 (x2) = 30,11/2, C 1/1, P 2/3, M 3/3 (x2) = 32, or 11/2, C 1/1, P 1/3, M 3/3 = 30. P2 is very small and completely displaced labially or completely absent, allowing full contact ofC1 and P4, and P3 is usually absent but, if present, is fully displaced labially and lies under cingula of P2 and P4, which are in contact. Habitat. Various semiarid savanna, mesic woodland savanna, and montane forest habitats, including Acacia ( Fabaceae )- Commiphora ( Burseraceae ) brushland/thicket and mosaics of evergreen brushland and secondary Acacia savanna, at elevations up to 1500 m . Food and Feeding. No information. Breeding. No information. Activity patterns. The Eloquent Horseshoe Bat roosts in caves. There might be more information available in papers that misidentified it as Hildebrandt’s Horseshoe Bat {hildebrandtii ). Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Eloquent Horseshoe Bat has a relatively wide distribution, but littie is known about its ecology. It seems to be most threatened by general habitat loss from agricultural expansion. Bibliography. ACR (201'8), Cotterill (2013b), Csorba et al. (2003), Koopman (1975), Monadjem, Taylor , Jacobs & Cotterill (2017d), Thorn & Kerbis Peterhans (2009), Taylor et al. (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus eloquens	Rhinolophus		eloquens	K. Andersen	1905	0	Ann. Mag. Nat. Hist.	ser. 7, 15: 74	Eloquent Horseshoe Bat	<b> perauritus </b>De Beaux, 1922.	Uganda, Entebbe.	Uganda, S Somalia, S Sudan, NE Dem. Rep. Congo, Kenya, Rwanda, N Tanzania, Zanzibar and Pemba.	Not listed.	Least Concern	 fumigatus species group. Includes perauritus; see Koopman (1975). May be conspecific with fumigatus Demos et al. (2019).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus eloquens	23	Eloquent Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	eloquens	K. Andersen	1905	0	Rhinolophus_Hildebranti_eloquens	Andersen, K. (1905). Further descriptions of new Rhinolophi from Africa. Annals and Magazine of Natural History, ser. 7, 15, 74.	https://www.biodiversitylibrary.org/item/63423#page/88/mode/1up	BM 1899.8.4.4		"Entebbi [= Entebbe], Uganda."			eloquens K. Andersen, 1905|perauritus De Beaux, 1922	NA	NA	South Sudan|Democratic Republic of the Congo|Uganda|Rwanda|Kenya|Tanzania|Somalia|Ethiopia?|Malawi?|Mozambique?	Africa	Afrotropic	LC	0	0	0	Rhinolophus_eloquens	0	sciname match	Rhinolophus_eloquens	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19539	Rhinolophus eloquens	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	eloquens	K. Andersen, 1905	The taxonomic relationship between Rhinolophus eloquens and R. brandti needs clarification.	20000000	Rhinolophus eloquens	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category	This species is associated with caves in savanna habitats. It is not known if it can persist in disturbed habitats.	This species is threatened by the general conversion of its habitat to agricultural use.	Little information is available on the population abundance or size of this species.	Unknown	This species is endemic to Central and Eastern Africa. It has been recorded from southern Sudan, eastern Democratic Republic of the Congo, Rwanda, Uganda, Kenya, Somalia and the islands of Zanzibar and Pemba in Tanzania.		Terrestrial	There appear to be no direct conservation measures in place. It is not known if the species is present in any protected areas. Further research needed into the distribution, natural history and threats to this bat.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		eloquens	K. Andersen	1905	0	Ann. Mag. Nat. Hist.	ser. 7, 15: 74	Eloquent Horseshoe Bat	<b> perauritus </b>De Beaux, 1922.	Uganda, Entebbe.	Uganda, S Somalia, S Sudan, NE Dem. Rep. Congo, Kenya, Rwanda, N Tanzania, Zanzibar and Pemba.	Not listed.	Least Concern	 fumigatus species group. Includes perauritus; see Koopman (1975). May be conspecific with fumigatus Demos et al. (2019).	Rhinolophus eloquens	1004680	23	Eloquent Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	eloquens	K. Andersen	1905	0	Rhinolophus_Hildebranti_eloquens	Andersen, K. (1905). Further descriptions of new Rhinolophi from Africa. Annals and Magazine of Natural History, ser. 7, 15, 74.	https://www.biodiversitylibrary.org/item/63423#page/88/mode/1up	BM 1899.8.4.4		"Entebbi [= Entebbe], Uganda."			eloquens K. Andersen, 1905|perauritus De Beaux, 1922	NA	NA				South Sudan|Democratic Republic of the Congo|Uganda|Rwanda|Kenya|Tanzania|Somalia|Ethiopia?|Malawi?|Mozambique?	Africa	Afrotropic	LC	0	0	0	Rhinolophus_eloquens	0	sciname match	Rhinolophus_eloquens	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_eloquens	1004680	23	Eloquent Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	eloquens	Andersen	0	Rhinolophus Hildebrandti eloquens	Andersen, K.C. 1905-01-01. Further descriptions of new _Rhinolophi_ from Africa. Annals and Magazine of Natural History (7)15(85):70-76.	https://www.biodiversitylibrary.org/page/19243592	BMNH:Mamm:1899.8.4.4	holotype	https://data.nhm.ac.uk/object/f7dd9086-cc93-4759-b8d9-24f8d88db755	"Entebbi [= Entebbe], Uganda."			NA	NA				South Sudan|Democratic Republic of the Congo|Uganda|Rwanda|Kenya|Tanzania|Somalia|Ethiopia?|Malawi?|Mozambique?	Africa	Afrotropic	LC	0	0	0	Rhinolophus_eloquens	0	sciname match	Rhinolophus_eloquens	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		eloquens	K. Andersen	1905	0	Ann. Mag. Nat. Hist.	ser. 7, 15: 74	Eloquent Horseshoe Bat	perauritus De Beaux, 1922.	Uganda, Entebbe.	Uganda, S Somalia, S Sudan, NE Dem. Rep. Congo, Kenya, Rwanda, N Tanzania, Zanzibar and Pemba.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19539/21979320/' target='_blank'>Least Concern</a>	fumigatus species group. Includes perauritus; see Koopman (1975). May be conspecific with fumigatus Demos et al. (2019).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus eloquens; Rhinolophus eloquens; Rhinolophus eloquens; Rhinolophus eloquens; Rhinolophus eloquens; Rhinolophus eloquens; eloquens; perauritus; eloquens; perauritus; perauritus; eloquens; perauritus; Eloquent Horseshoe Bat; Rhinolophe éloquent; Lautfreudige Hufeisennase; Herradura elocuente; Eloquent Horseshoe Bat; Eloquent Horseshoe Bat; Eloquent Horseshoe Bat; R. eloquens