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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1360	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus	Rhinolophus coelophyllus		[MSW2] Does not include shamelv, see Hill and Thonglongya (1972:183-186).; [MSW3] euryotis species group. Does not include shameli; see Hill and Thonglongya (1972). Reviewed in part by Yoshiyuki (1990).; [HMW] Rhinolophus coelophyllus Peters, 1867 , “ Moulmein [= Mawlamyine], Burmah [= Burma = Myanmar ] ... Salween valley. ” Rhinolophus coelophyllus is included in the euryotis species group. It is considered to be closely related to R. shameli , based on morphology; the two are often confused, and some records of each were previously attributed to the wrong species. Monotypic .; [batnames2022]  euryotis species group. Does not include shameli; see Hill and Thonglongya (1972). Reviewed in part by Yoshiyuki (1990).; [IUCN] This species is often confused with R. shameli , and some historical records refer to this other taxon (Ith et al. 2011).; [batnames2023]  euryotis species group. Does not include shameli; see Hill and Thonglongya (1972). Reviewed in part by Yoshiyuki (1990).; [batnames2025_1.7] euryotisspecies group. Does not include shameli; see Hill and Thonglongya (1972). Reviewed in part by Yoshiyuki (1990).														coelophyllus	This species is often confused with R. shameli , and some historical records refer to this other taxon (Ith et al. 2011).			coelophyllus 	coelophyllus 			coelophyllus W. C. H. Peters, 1867		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Croslet horseshoe bat	Burma – Malaya	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhinolophus coelophyllus	Burma, Salaween River.	Peters	1867	Proc. Zool. Soc. Lond., 1866:426.	Distribution: Known only from Burma, Thailand, and Malaya.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Croslet horseshoe bat (Peters' horseshoe bat)	Burma – Malaya	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1867	Proc. Zool. Soc. Lond., 1866:426 [1867].	Does not include shamelv, see Hill and Thonglongya (1972:183-186).	W Malaysia, Thailand, Burma.	Burma, Salaween River.		PETERS	1867	Anterior edge of horseshoe narrowly emar ginated, the emargination extended posteriorly as a narrow groove reaching less than halfway to internarial region. Base of well-developed con necting process at most sparsely haired; recessed into a vertical fissure at base of lancet. Postnarial rostral depression of skull moderately deep, elon gate, enclosed by broad, well-developed supraor bital ridges. Size medium (forearm length, 41 -45 mm).	Distribution: Known only from Burma, Thailand, and Malaya.	No subspecies.		58	species	R. coelophyllus	PETERS	1867	Rhinolophus	genus	Rhinolophus coelophyllus				Anterior edge of horseshoe narrowly emar ginated, the emargination extended posteriorly as a narrow groove reaching less than halfway to internarial region. Base of well-developed con necting process at most sparsely haired; recessed into a vertical fissure at base of lancet. Postnarial rostral depression of skull moderately deep, elon gate, enclosed by broad, well-developed supraor bital ridges. Size medium (forearm length, 41 -45 mm).	No subspecies.		59. R. coelophyllus PETERS 1867 [euryotis group].	59	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus coelophyllus	Rhinolophus		coelophyllus	Peters		1866	1867	Proc. Zool. Soc. Lond.	1866		426		Croslet Horseshoe Bat	Burma, Salaween (= Salween) River.	W Malaysia, Thailand, Burma, Laos.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		euryotis species group. Does not include shameli; see Hill and Thonglongya (1972). Reviewed in part by Yoshiyuki (1990).	885887A2FFEC8A0AFF06F534F239DC09	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	321	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFE58A02F892F252FC75D193.xml	Rhinolophus coelophyllus	Rhinolophidae	Rhinolophus	coelophyllus	Peters	1867	Croslet Horseshoe Bat @en | Rhinolophe de Peters @fr | Thai-Hufeisennase @de | Herradura thai @es	Rhinolophus coelophyllus Peters, 1867 , “ Moulmein [= Mawlamyine], Burmah [= Burma = Myanmar ] ... Salween valley. ” Rhinolophus coelophyllus is included in the euryotis species group. It is considered to be closely related to R. shameli , based on morphology; the two are often confused, and some records of each were previously attributed to the wrong species. Monotypic .	NC, C & SE Myanmar , Thailand (including Tarutao I), C Laos , and NW Peninsular Malaysia (including Langkawi I).	Head-body 36-50 mm , tail 16-4-23- 9 mm , ear 15- 1-20 mm , hindfoot 7-8-9- 6 mm , forearm 41-6-47- 8 mm ; weight 6-2-8- 6 g . There is considerable geographic variation in terms of skull measurements. Dorsal pelage varies from grayish brown to dark brown, with some individuals reddish brown (base of hairs paler); ventral pelage is pale buff brown and can be nearly creamy white in some individuals, or pale orange in reddish individuals. Ears are medium-sized. Noseleaf has very short, thick lancet with convex outline; connecting process is arched and proximally inserted into densely haired vertical fissure in lancet; sella is short, with almost parallel or continuously convergent sides and rounded tip; horseshoe is moderately wide (8-2-10- 4 mm ) and has weakly developed lateral leaflets. Lower lip has three mental grooves. Skull is moderately built (zygomatic width is slightly greater than mastoid width); sagittal crest is moderately developed; anterior median swellings are inflated and distinctly elevated above posterior swellings; anterior border of rostrum is shorter than in Shamel’s Horsehsoe Bat (. shameli ); nasal swelling as whole is smaller than in Shamel’s Horsehsoe Bat ; frontal depression is deep and pronounced with well-defined supraorbital ridges, although specimens from central and north Myanmar have less developed depressions and supraorbital ridges; canines are slender. P2 is moderately large and within the tooth row; P3 is small and extruded from tooth row; P and P4 are touching. Chromosomal complement has 2n = 62, FN = 64, and FNa = 60 ( Thailand ).	The Croslet Horseshoe Bat seems to favor limestone karst regions with mixed deciduous forest but can also be found in a variety of other habitats, including deciduous dipterocarp forests, moist evergreen forests, hill evergreen forests, dry evergreen forests, pine forests, rubber plantations, orchards, and extensive agricultural land. Throughout the species’ distribution, it can be found from near sea level up to 1500 m elevation.	No information.	Pregnant females were captured in March, and an immature male was captured in October on Tarutao Island.	Croslet Horseshoe Bats roost in caves and tunnels by day. Calls are a FM/CF/FM shape with a peak F recorded at 76-1-84-8 kHz across Thailand , 75- 80 kHz in Kanchanaburi Province , Thailand , 74-5—74-9 kHz in Myanmar , and 76 kHz in Laos .	Croslet Horseshoe Bats roost in relatively large colonies, often with c.100-150 individuals. Up to 900 bats have been recorded roosting in a single cave.	Classified as Least Concern on The IUCN ed List. The Croslet Horseshoe Bat is relatively widespread and common throughout its distribution and does not seem to face any major threats at present.	Bates, Bumrungsri & Csorba (2008 c) | Bates, Thi Mar-Mar et al. (2004) | Harada, Minezawa et al. (1982) | Harada, Yenbutra, Yosida &Takada (1985) | Hill &Thonglongya (1972) | Hood et al. (1988) | Ith, Soisook et al. (2011) | Patawang et al. (2017) | Struebig et al. (2005)	https://zenodo.org/record/3750086/files/figure.png	88 . Croslet Horseshoe Bat Rhinolophus coelophyllus French: Rhinolophe de Peters / German: Thai-Hufeisennase / Spanish: Herradura thai Taxonomy. Rhinolophus coelophyllus Peters, 1867 , “ Moulmein [= Mawlamyine], Burmah [= Burma = Myanmar ] ... Salween valley. ” Rhinolophus coelophyllus is included in the euryotis species group. It is considered to be closely related to R. shameli , based on morphology; the two are often confused, and some records of each were previously attributed to the wrong species. Monotypic . Distribution . NC, C & SE Myanmar , Thailand (including Tarutao I), C Laos , and NW Peninsular Malaysia (including Langkawi I). Descriptive notes . Head-body 36-50 mm , tail 16-4-23- 9 mm , ear 15- 1-20 mm , hindfoot 7-8-9- 6 mm , forearm 41-6-47- 8 mm ; weight 6-2-8- 6 g . There is considerable geographic variation in terms of skull measurements. Dorsal pelage varies from grayish brown to dark brown, with some individuals reddish brown (base of hairs paler); ventral pelage is pale buff brown and can be nearly creamy white in some individuals, or pale orange in reddish individuals. Ears are medium-sized. Noseleaf has very short, thick lancet with convex outline; connecting process is arched and proximally inserted into densely haired vertical fissure in lancet; sella is short, with almost parallel or continuously convergent sides and rounded tip; horseshoe is moderately wide (8-2-10- 4 mm ) and has weakly developed lateral leaflets. Lower lip has three mental grooves. Skull is moderately built (zygomatic width is slightly greater than mastoid width); sagittal crest is moderately developed; anterior median swellings are inflated and distinctly elevated above posterior swellings; anterior border of rostrum is shorter than in Shamel’s Horsehsoe Bat (. shameli ); nasal swelling as whole is smaller than in Shamel’s Horsehsoe Bat ; frontal depression is deep and pronounced with well-defined supraorbital ridges, although specimens from central and north Myanmar have less developed depressions and supraorbital ridges; canines are slender. P2 is moderately large and within the tooth row; P3 is small and extruded from tooth row; P and P4 are touching. Chromosomal complement has 2n = 62, FN = 64, and FNa = 60 ( Thailand ). Habitat. The Croslet Horseshoe Bat seems to favor limestone karst regions with mixed deciduous forest but can also be found in a variety of other habitats, including deciduous dipterocarp forests, moist evergreen forests, hill evergreen forests, dry evergreen forests, pine forests, rubber plantations, orchards, and extensive agricultural land. Throughout the species’ distribution, it can be found from near sea level up to 1500 m elevation. Food and Feeding. No information. Breeding. Pregnant females were captured in March, and an immature male was captured in October on Tarutao Island. Activity patterns. Croslet Horseshoe Bats roost in caves and tunnels by day. Calls are a FM/CF/FM shape with a peak F recorded at 76-1-84-8 kHz across Thailand , 75- 80 kHz in Kanchanaburi Province , Thailand , 74-5—74-9 kHz in Myanmar , and 76 kHz in Laos . Movements, Home range and Social organization. Croslet Horseshoe Bats roost in relatively large colonies, often with c.100-150 individuals. Up to 900 bats have been recorded roosting in a single cave. Status and Conservation. Classified as Least Concern on The IUCN ed List. The Croslet Horseshoe Bat is relatively widespread and common throughout its distribution and does not seem to face any major threats at present. Bibliography. Bates, Bumrungsri & Csorba (2008 c ), Bates, Thi Mar-Mar et al. (2004), Harada, Minezawa et al. (1982), Harada, Yenbutra, Yosida &Takada (1985), Hill &Thonglongya (1972), Hood et al. (1988), Ith, Soisook et al. (2011), Patawang et al. (2017), Struebig et al. (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus coelophyllus	Rhinolophus		coelophyllus	Peters	1867	0	Proc. Zool. Soc. Lond.	1873:06:00	Croslet Horseshoe Bat	None.	Burma, Salaween (= Salween) River.	W Malaysia, Thailand, Burma, Laos.	Not listed.	Least Concern	 euryotis species group. Does not include shameli; see Hill and Thonglongya (1972). Reviewed in part by Yoshiyuki (1990).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus coelophyllus	23	Croslet Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	coelophyllus	W. Peters	1867	0	Rhinolophus_coelophyllus	Peters, W. C. H. (1867). On some Mammalia collected by Capt. A.C. Beavan, C.M.Z.S., at Moulmein, Burmah. Proceedings of the Zoological Society London, 1867 [for 1866], 426.	https://www.biodiversitylibrary.org/item/90954#page/516/mode/1up	ZMB 3143		"Moulmein [= Mawlamyine], Burmah [= Burma = Myanmar] ... Salween valley."			coelophyllus W. Peters, 1867	NA	NA	Myanmar|Thailand|Laos|Malaysia	Asia	Indomalaya	LC	0	0	0	Rhinolophus_coelophyllus	0	sciname match	Rhinolophus_coelophyllus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19532	Rhinolophus coelophyllus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	coelophyllus	Peters, 1867	This species is often confused with R. shameli , and some historical records refer to this other taxon (Ith et al. 2011).	20000000	Rhinolophus coelophyllus	Least Concern		2020	2019-08-06 00:00:00 UTC	3.1	English	<p>Listed as Least Concern as the species is relatively widespread, although never common, there are no major threats at present (although cave disturbance could pose a threat to local populations in limestone karst areas), and it occurs in a number of protected areas.</p>	<p>The species appears to favour limestone karst areas in mixed deciduous forest, but is also found in deciduous dipterocarp, moist evergreen, hill evergreen, dry evergreen and pine forests. It also appears quite tolerant of highly disturbed habitats, also being found in rubber plantations, orchards and extensive agricultural lands. In Thailand, the species has been found at altitudes ranging from near sea level to 1,500 m (Ith et al. 2011).</p>	<p>There are no major threats to this relatively widespread species, although disturbance of caves may pose a threat to local populations in limestone karst areas.</p>	<p>The species is locally common in Thailand, a population there may have contained 1,000 individuals (S. Bumrungsri pers. comm.). Its population status in other countries is not known but it is not thought to be common anywhere.</p>	Stable	<p>This species ranges from northern Myanmar to northern Peninsular Malaysia, including central Thailand and western Lao PDR (Ith et al. 2011, Thomas et al. 2013). </p>		Terrestrial	It occurs in a number of protected areas.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		coelophyllus	Peters	1867	0	Proc. Zool. Soc. Lond.	1873:06:00	Croslet Horseshoe Bat	None.	Burma, Salaween (= Salween) River.	W Malaysia, Thailand, Burma, Laos.	Not listed.	Least Concern	 euryotis species group. Does not include shameli; see Hill and Thonglongya (1972). Reviewed in part by Yoshiyuki (1990).	Rhinolophus coelophyllus	1004670	23	Croslet Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	coelophyllus	W. Peters	1867	0	Rhinolophus_coelophyllus	Peters, W. C. H. (1867). On some Mammalia collected by Capt. A.C. Beavan, C.M.Z.S., at Moulmein, Burmah. Proceedings of the Zoological Society London, 1867 [for 1866], 426.	https://www.biodiversitylibrary.org/item/90954#page/516/mode/1up	ZMB 3143		"Moulmein [= Mawlamyine], Burmah [= Burma = Myanmar] ... Salween valley."			coelophyllus W. Peters, 1867	NA	NA				Myanmar|Thailand|Laos|Malaysia	Asia	Indomalaya	LC	0	0	0	Rhinolophus_coelophyllus	0	sciname match	Rhinolophus_coelophyllus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_coelophyllus	1004670	23	Croslet Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	coelophyllus	W. C. H. Peters	0	Rhinolophus coelophyllus	Peters, W.C.H. 1867-04. On some Mammalia collected by Capt. A. C. Beavan, C.M.Z.S., at Moulmein, Burmah. Proceedings of the Zoological Society of London 1866(3):426-429.	https://www.biodiversitylibrary.org/page/28627918	ZMB 3143	holotype		"Moulmein [= Mawlamyine], Burmah [= Burma = Myanmar] ... Salween valley."			NA	NA				Myanmar|Thailand|Laos|Malaysia	Asia	Indomalaya	LC	0	0	0	Rhinolophus_coelophyllus	0	sciname match	Rhinolophus_coelophyllus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		coelophyllus	Peters	1867	0	Proc. Zool. Soc. Lond.	1873:06:00	Croslet Horseshoe Bat	None.	Burma, Salaween (= Salween) River.	W Malaysia, Thailand, Burma, Laos.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19532/21980746/' target='_blank'>Least Concern</a>	euryotisspecies group. Does not include shameli; see Hill and Thonglongya (1972). Reviewed in part by Yoshiyuki (1990).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus coelophyllus; Rhinolophus coelophyllus; Rhinolophus coelophyllus; Rhinolophus coelophyllus; Rhinolophus coelophyllus; Rhinolophus coelophyllus; coelophyllus; Croslet Horseshoe Bat; Rhinolophe de Peters; Thai-Hufeisennase; Herradura thai; Croslet Horseshoe Bat; Croslet Horseshoe Bat; Croslet Horseshoe Bat; R. coelophyllus