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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1356	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Rhinolophus chiewkweeae	Rhinolophus chiewkweeae	Rhinolophus chiewkweeae	Rhinolophus chiewkweeae	Rhinolophus chiewkweeae	Rhinolophus chiewkweeae	Rhinolophus chiewkweeae	Rhinolophus chiewkweeae	Rhinolophus chiewkweeae		[HMW] Rhinolophus chiewkweeae Yoshiyuki & B. L. Lim, 2005 , “Gunung Ledang, 2°84’N, 102°57’E, Tangkak, Muar,Johore, a submontane dipterocarp forest approximately 1276 m in elevations, Malaysia .” Rhinolophus chiewkweeae is included in the pearsonii species group, and appears to be closely related to R.pearsonii . Monotypic.; [MDD2022] recently described; [IUCN] Rhinolophus chiewkweeae is a member of the pearsoni species group which consists of R. pearsoni and R. yunanensis (Yoshiyuki and Lim 2005). It is orange brown in coloration (both R. pearsoni and R. yunanensis are dark brown) and has intermediate characters to those of R. pearsoni and R. yunanensis , except for its short ears and tail (Yoshiyuki and Lim 2005).; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														chiewkweeae	Rhinolophus chiewkweeae is a member of the pearsoni species group which consists of R. pearsoni and R. yunanensis (Yoshiyuki and Lim 2005). It is orange brown in coloration (both R. pearsoni and R. yunanensis are dark brown) and has intermediate characters to those of R. pearsoni and R. yunanensis , except for its short ears and tail (Yoshiyuki and Lim 2005).			chiewkweeae 	chiewkweeae 			chiewkweeae Yoshiyuki & Lim Boo Liat, 2005						N/A																																								NA																											885887A2FFDE8A3BF8B3F693F72EDD14	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	332	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFFA8A1CFF03FEF1FCCAD36A.xml	Rhinolophus chiewkweeae	Rhinolophidae	Rhinolophus	chiewkweeae	Yoshiyuki & Lim	2005	Chiew Kwee’s Horseshoe Bat @en | Rhinolophe de Chiew Kwee @fr | Chiew-Kwee-Hufeisennase @de | Herradura de Chiew Kwee @es	Rhinolophus chiewkweeae Yoshiyuki & B. L. Lim, 2005 , “Gunung Ledang, 2°84’N, 102°57’E, Tangkak, Muar,Johore, a submontane dipterocarp forest approximately 1276 m in elevations, Malaysia .” Rhinolophus chiewkweeae is included in the pearsonii species group, and appears to be closely related to R.pearsonii . Monotypic.	Malay Peninsula, including Langkawi I.	Head—body 51-5—65 mm, tail 14- 7-19 mm , ear 20- 1-26 mm , hindfoot 12-14- 4 mm , forearm 51-9- 56- 1 mm . Dorsal pelage is orangish brown, while ventral pelage is lighter. Ears are moderately large. Noseleaf has a broadly triangular, long, and nearly parallel straightsided lancet; connecting process starts at the tip of the sella and is low and arched; sella is broadly rounded at the tip and broader at the base; horseshoe is wide (11- 6 mm ) and covers the muzzle, with a distinct median emargination. Lower lip has one mental groove. Baculum is similar to that of Pearson’s Horseshoe Bat (. pearsonii ) and is fairly long (3- 3 mm long), having a deep U-shaped emargination and a thick lateral expansion on the ampulla. Skull is large and robust (zygomatic width greater than mastoid width); anterior median swellings are well developed; lateral swellings are less swollen and the posterior swellings are relatively concave; rostral profile is low; frontal depression appears to be relatively shallow; sagittal and supraorbital crests are well developed. C1 is weak; P2 is relatively large and within the tooth row; P3 is minute and extruded from the tooth row, allowing P2 and P4 to touch.	Typically found in hill and lowland dipterocarp forests, at elevations from around sea level up to 1276 m .	No information.	No information.	Call shape is FM /CF/FM with peak F of 53-6-54-7 kHz, duration of 28-4-63-3 milliseconds, and interpulse interval of 32—289 milliseconds, in eastern Peninsular Malaysia .	No information.	Not assessed on The IUCN Red List. The overall range of Chiew Kwee’s Horseshoe Bat is quite small. Deforestation is likely to be a major threat to the species. Due to reservoirs for hydroelectric schemes, mercury accumulates at high concentrations in this species, which may well be a localized threat.	Kumaran , Daud et al. (2013) | Morni et al. (2016) | Pounsin et al. (2018) | Syaripuddin et al. (2014) | Yoshiyuki & Lim (2005) | Zalipah et al. (2019)	https://zenodo.org/record/3750126/files/figure.png	107 . Chiew Kwee’s Horseshoe Bat Rhinolophus chiewkweeae French: Rhinolophe de Chiew Kwee / German: Chiew-Kwee-Hufeisennase I Spanish: Herradura de Chiew Kwee Taxonomy. Rhinolophus chiewkweeae Yoshiyuki & B. L. Lim, 2005 , “Gunung Ledang, 2°84’N, 102°57’E, Tangkak, Muar,Johore, a submontane dipterocarp forest approximately 1276 m in elevations, Malaysia .” Rhinolophus chiewkweeae is included in the pearsonii species group, and appears to be closely related to R.pearsonii . Monotypic. Distribution. Malay Peninsula, including Langkawi I. Descriptive notes. Head—body 51-5—65 mm, tail 14- 7-19 mm , ear 20- 1-26 mm , hindfoot 12-14- 4 mm , forearm 51-9- 56- 1 mm . Dorsal pelage is orangish brown, while ventral pelage is lighter. Ears are moderately large. Noseleaf has a broadly triangular, long, and nearly parallel straightsided lancet; connecting process starts at the tip of the sella and is low and arched; sella is broadly rounded at the tip and broader at the base; horseshoe is wide (11- 6 mm ) and covers the muzzle, with a distinct median emargination. Lower lip has one mental groove. Baculum is similar to that of Pearson’s Horseshoe Bat (. pearsonii ) and is fairly long (3- 3 mm long), having a deep U-shaped emargination and a thick lateral expansion on the ampulla. Skull is large and robust (zygomatic width greater than mastoid width); anterior median swellings are well developed; lateral swellings are less swollen and the posterior swellings are relatively concave; rostral profile is low; frontal depression appears to be relatively shallow; sagittal and supraorbital crests are well developed. C1 is weak; P2 is relatively large and within the tooth row; P3 is minute and extruded from the tooth row, allowing P2 and P4 to touch. Habitat. Typically found in hill and lowland dipterocarp forests, at elevations from around sea level up to 1276 m . Food and Feeding. No information. Breeding. No information. Activity patterns. Call shape is FM /CF/FM with peak F of 53-6-54-7 kHz, duration of 28-4-63-3 milliseconds, and interpulse interval of 32—289 milliseconds, in eastern Peninsular Malaysia . Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The IUCN Red List. The overall range of Chiew Kwee’s Horseshoe Bat is quite small. Deforestation is likely to be a major threat to the species. Due to reservoirs for hydroelectric schemes, mercury accumulates at high concentrations in this species, which may well be a localized threat. Bibliography. Kumaran , Daud et al. (2013), Morni et al. (2016), Pounsin et al. (2018), Syaripuddin et al. (2014), Yoshiyuki & Lim (2005), Zalipah et al. (2019).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus chiewkweeae	Rhinolophus		chiewkweeae	Yoshiyuki & Lim	2005	0	Bull Natn. Sci Mus Tokyo, Ser. A	31(1): 30	Chiew Kwee&apos;s Horseshoe Bat	None.	Gunung Ledang, Tangkak, Muar, Johore, Malaysia	Malaysia	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus chiewkweeae	23	Chiew Kwee's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	chiewkweeae	Yoshiyuki & Boo Liat Lim	2005	0	Rhinolophus_chiewkweeae	Yoshiyuki, M., Lim, B.L. 2005. A new horseshoe bat, Rhinolophus chiewkweeae (Chiroptera, Rhinolophidae), from Malaysia. Bull. Natn. Sci. Mus., Tokyo, Ser. A. 31(1): 30.	https://www.kahaku.go.jp/research/publication/zoology/download/31_1/BNSM_A310105.pdf	NSMT 33472		"Gunung Ledang, 2°84'N, 102°57'E, Tangkak, Muar, Johore, a submontane dipterocarp forest approximately 1276 m in elevations, Malaysia."		102.95	chiewkweeae Yoshiyuki & Boo Liat Lim, 2005	recently described	Yoshiyuki, M., Lim, B.L. 2005. A new horseshoe bat, Rhinolophus chiewkweeae (Chiroptera, Rhinolophidae), from Malaysia. Bull. Natn. Sci. Mus., Tokyo, Ser. A. 31(1): 29-36.	Malaysia	Asia	Indomalaya	DD	0	0	0	Rhinolophus_chiewkweeae	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Rhinolophus chiewkweeae	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	chiewkweeae	Yoshiyuki &; Lim, 2005	Rhinolophus chiewkweeae is a member of the pearsoni species group which consists of R. pearsoni and R. yunanensis (Yoshiyuki and Lim 2005). It is orange brown in coloration (both R. pearsoni and R. yunanensis are dark brown) and has intermediate characters to those of R. pearsoni and R. yunanensis , except for its short ears and tail (Yoshiyuki and Lim 2005).	80000000	Rhinolophus chiewkweeae	Data Deficient		2020	2019-07-20 00:00:00 UTC	3.1	English	Rhinolophus chiewkweeae is listed as Data Deficient it is only known from 7 records in Peninsular Malaysia and there is very limited data on the species distribution, population status, ecological requirements, and life history.	Little data has been reported on the habitat for the species and no roosts have been identified. However, on the east coast of Peninsular Malaysia the species has been captured in secondary diptocarp forests at an elevation of 155 m (Morni et al.   2016).	Threats to the species are unknown. However, it can be assumed that continued deforestation and changing fire regimes in the region are impacting the species given it has been captured in forests.	The status of the global population of the species is unknown, but is suspected to be in decline due to forest loss and broader environmental degradation throughout Peninsular Malaysia. Where encountered, the population density for the species is likely low as capture rates have been &lt;10% (Morni et al. 2016, Jayaraj et al. 2013).	Unknown	Rhinolophus chiewkweeaeis currently recognized as an endemic to Peninsular Malaysia and the holotype was collected at Gunung Ledang, Johor, Peninsular Malaysia in 1998 (Yoshiyuki and Lim 2005, Morni et al.     2016). It is only known from seven localities, six of which were from the west coast (Yoshiyuki and Lim 2005, Jayaraj et al.     2013) with the latest being from the east coast at Sungai Buweh, Tasik Kenyir, Terengganu (Morni et al.     2016).		Terrestrial	The species is known from protected areas in Peninsular Malaysia (Jayaraj et al. 2013, Murni et al. 2016). Strengthened protection and management of forests would benefit the species and other forest bats. Additional research is needed on the species population, ecology, life history, and threats.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		chiewkweeae	Yoshiyuki & Lim	2005	0	Bull Natn. Sci Mus Tokyo, Ser. A	31(1): 30	Chiew Kwee&apos;s Horseshoe Bat	None.	Gunung Ledang, Tangkak, Muar, Johore, Malaysia	Malaysia	Not listed.	Data Deficient		Rhinolophus chiewkweeae	1004668	23	Chiew Kwee's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	chiewkweeae	Yoshiyuki & Boo Liat Lim	2005	0	Rhinolophus_chiewkweeae	Yoshiyuki, M., Lim, B.L. 2005. A new horseshoe bat, Rhinolophus chiewkweeae (Chiroptera, Rhinolophidae), from Malaysia. Bull. Natn. Sci. Mus., Tokyo, Ser. A. 31(1): 30.	https://www.kahaku.go.jp/research/publication/zoology/download/31_1/BNSM_A310105.pdf	NSMT 33472		"Gunung Ledang, 2°84'N, 102°57'E, Tangkak, Muar, Johore, a submontane dipterocarp forest approximately 1276 m in elevations, Malaysia."		102.95	chiewkweeae Yoshiyuki & Boo Liat Lim, 2005	recently described	Yoshiyuki, M., Lim, B.L. 2005. A new horseshoe bat, Rhinolophus chiewkweeae (Chiroptera, Rhinolophidae), from Malaysia. Bull. Natn. Sci. Mus., Tokyo, Ser. A. 31(1): 29-36.				Malaysia	Asia	Indomalaya	DD	0	0	0	Rhinolophus_chiewkweeae	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_chiewkweeae	1004668	23	Chiew Kwee's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	chiewkweeae	Yoshiyuki & Lim Boo Liat	0	Rhinolophus chiewkweeae	Yoshiyuki, M. and Lim, B.L. 2005-03-22. A new horseshoe bat, _Rhinolophus chiewkweeae_ (Chiroptera, Rhinolophidae), from Malaysia. Bulletin of the National Science Museum, Tokyo (A)31(1):29-36.	http://ci.nii.ac.jp/naid/110004665380	NSMT-M 33472	holotype		"Gunung Ledang, 2°84'N, 102°57'E, Tangkak, Muar, Johore, a submontane dipterocarp forest approximately 1276 m in elevations, Malaysia."	2.34	102.57	recently described	Yoshiyuki, M., Lim, B.L. 2005. A new horseshoe bat, Rhinolophus chiewkweeae (Chiroptera, Rhinolophidae), from Malaysia. Bull. Natn. Sci. Mus., Tokyo, Ser. A. 31(1): 29-36.				Malaysia	Asia	Indomalaya	DD	0	0	0	Rhinolophus_chiewkweeae	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		chiewkweeae	Yoshiyuki & Lim	2005	0	Bull Natn. Sci Mus Tokyo, Ser. A	31(1): 30	Chiew Kwee&apos;s Horseshoe Bat	None.	Gunung Ledang, Tangkak, Muar, Johore, Malaysia	Malaysia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84372474/84372528/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus chiewkweeae; Rhinolophus chiewkweeae; Rhinolophus chiewkweeae; Rhinolophus chiewkweeae; Rhinolophus chiewkweeae; chiewkweeae; Chiew Kwee’s Horseshoe Bat; Rhinolophe de Chiew Kwee; Chiew-Kwee-Hufeisennase; Herradura de Chiew Kwee; Chiew Kwee's Horseshoe Bat; Chiew Kwee&apos;s Horseshoe Bat; R. chiewkweeae