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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1348	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis	Rhinolophus borneensis		[MSW2] Includes chaseni and importunus; see Hill (1983). Formerly included javanicus, celebensis, madurensis, and parvus; see Goodwin (1979:104) and Hill and Thonglongya (1972:187); but see Hill (1983).; [MSW3] megaphyllus species group. Includes chaseni and importunus; see Hill (1983). Formerly included javanicus, celebensis, madurensis, and parvus (e.g., by Goodwin [1979] and Hill and Thonglongya [1972]) but see Hill (1983) and Kitchener et al. (1995a). Type material discussed by Csorba (2002). Subspecies limits are somewhat unclear, and the relationships of various forms to celebensis and malayanus remains problematic; see Csorba et al. (2003). This complex may include more than one species, see Csorba et al. (2003).; [HMW] Rhinolophus borneensis Peters, 1861 , “ Labuan [Island], Borneo ,” Malaysia . Traditionally included in the megaphyllus species group, but this needs to be investigated further using genetic techniques, as A borneensis seems to be sister to a clade containing members of the pusillus species group. All mainland South-east Asian populations are here considered to belong to A chaseni , while Indonesian populations are tentatively included in A borneensis . The identity of Peninsular Malaysian populations is uncertain as no genetic data have been used for testing, so they are here listed under this species, pending further studies. This species may represent a species complex. Three subspecies are recognized.; [batnames2022]  megaphyllus species group. Does not include chaseni ; see Francis et al. (2010), Kruskop (2011), and Abramov et al. (2018). Includes importunus ; see Hill (1983). Formerly included javanicus , celebensis , madurensis , and parvus (e.g., by Goodwin [1979] and Hill and Thonglongya [1972]) but see Hill (1983) and Kitchener et al.(1995a). Type material discussed by Csorba (2002). Subspecies limits are somewhat unclear, and the relationships of various forms to celebensis and malayanus remains problematic; see Csorba et al. (2003). This complex may include more than one species, seeCsorba et al. (2003).; [MDD2022] previously included R. chaseni; [IUCN] This is most likely a species complex (Csorba et al . 2003). Lekagul and McNeely (1977) synonymized this species with Rhinolophus robinsoni , but robinsoni is now considered a separate species (Simmons 2005). The distribution of R. borneensis may be confused, because of the unresolved taxonomy of the group.; [batnames2023]  megaphyllus species group. Does not include chaseni ; see Francis et al. (2010), Kruskop (2011), and Abramov et al. (2018). Includes importunus ; see Hill (1983). Formerly included javanicus , celebensis , madurensis , and parvus (e.g., by Goodwin [1979] and Hill and Thonglongya [1972]) but see Hill (1983) and Kitchener et al.(1995a). Type material discussed by Csorba (2002). Subspecies limits are somewhat unclear, and the relationships of various forms to celebensis and malayanus remains problematic; see Csorba et al. (2003). This complex may include more than one species, seeCsorba et al. (2003).; [MDD2023] previously included R. chaseni; [MDD2025_2.0] previously included R. chaseni; [batnames2025_1.7] megaphyllusspecies group. Does not include chaseni; see Francis et al. (2010), Kruskop (2011), and Abramov et al. (2018). Includes importunus; see Hill (1983). Formerly included javanicus, celebensis, madurensis, and parvus (e.g., by Goodwin [1979] and Hill and Thonglongya [1972]) but see Hill (1983) and Kitchener et al.(1995a). Type material discussed by Csorba (2002). Subspecies limits are somewhat unclear, and the relationships of various forms to celebensis and malayanus remains problematic; see Csorba et al. (2003). This complex may include more than one species, seeCsorba et al. (2003).; [MDD2025_2.2] previously included R. chaseni				javanicus, celebensis, chaseni	(importunus)	chaseni, importunus, spadix.	borneensis, spadix, importunus, chaseni	borneensis, chaseni, importunus, spadix		borneensis, importunus, spadix		borneensis, chaseni, importunus, spadix		borneensis, spadix, importunus	This is most likely a species complex (Csorba et al . 2003). Lekagul and McNeely (1977) synonymized this species with Rhinolophus robinsoni , but robinsoni is now considered a separate species (Simmons 2005). The distribution of R. borneensis may be confused, because of the unresolved taxonomy of the group.	borneensis, importunus, spadix		borneensis, spadix, importunus 	borneensis, spadix, importunus 	borneensis, importunus, spadix		borneensis W. C. H. Peters, 1861|spadix G. S. Miller, 1901|importunus Chasen, 1939		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Bornean horseshoe bat	Cambodia, Borneo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhinolophus borneensis	Malaysia, Sabah, Labuan Isl.	Peters	1861	Monatsb. Preuss. Akad. Wiss. Berlin, p. 709.	Distribution: Confined to a small area of southern Indo-China, Borneo and nearby islands, and Java.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Bornean horseshoe bat	Kampuchea, ? Vietnam, Borneo, Java; ref. 4.12	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1861	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1861:709.	Includes chaseni and importunus; see Hill (1983). Formerly included javanicus, celebensis, madurensis, and parvus; see Goodwin (1979:104) and Hill and Thonglongya (1972:187); but see Hill (1983).	Borneo; Labuan and Banguey Isis (Malaysia); Java, Karimata Isis and South Natuna Isis (Indonesia); Cambodia; Vietnam.	Malaysia, Sabah, Labuan Isl.		PETERS	1861	Sella rounded on top. Lancet more or less hastate. Supraorbital width of skull less than width of nasal swellings which are not enlarged. Palatal bridge relatively short. Middle lower premolar extruded from toothrow. Supraorbital crests in front of mid-orbit. Braincase relatively inflated. Size relatively small (forearm length, 41-47 mm; total length of skull, 18-20 mm).	Distribution: Confined to a small area of southern Indo-China, Borneo and nearby islands, and Java.	There are four currently recognized sub species (HILL 1983).	R. b. borneensis (Borneo), R. b. spadix (South Natunas and Karimata in the South China Sea), R. b. importunus (Java), R. b. chaseni (Cambodia and extreme southern Vietnam).	52	species	R. borneensis	PETERS	1861	Rhinolophus	genus	Rhinolophus borneensis				Sella rounded on top. Lancet more or less hastate. Supraorbital width of skull less than width of nasal swellings which are not enlarged. Palatal bridge relatively short. Middle lower premolar extruded from toothrow. Supraorbital crests in front of mid-orbit. Braincase relatively inflated. Size relatively small (forearm length, 41-47 mm; total length of skull, 18-20 mm).	There are four currently recognized sub species (HILL 1983).		4. R. borneensis PETERS 1861 [ferrumequium group].	4	_R. b. borneensis_ Peters, 1861; _R. b. importunus_ Chasen, 1939; _R. b. spadix_ Miller, 1901			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus borneensis	Rhinolophus		borneensis	Peters		1861		Monatsb. K. Preuss. Akad. Wiss. Berlin	1861		709		Bornean Horseshoe Bat	Malaysia, N Borneo, Sabah, Labuan Isl.	Borneo; Labuan and Banguey Isls (Malaysia); Java, Karimata Isls, and South Natuna Isls (Indonesia); Cambodia, Laos, and Vietnam.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	chaseni Sanborn, 1939; importunus Chasen, 1939; spadix Miller, 1901.	megaphyllus species group. Includes chaseni and importunus; see Hill (1983). Formerly included javanicus, celebensis, madurensis, and parvus (e.g., by Goodwin [1979] and Hill and Thonglongya [1972]) but see Hill (1983) and Kitchener et al. (1995a). Type material discussed by Csorba (2002). Subspecies limits are somewhat unclear, and the relationships of various forms to celebensis and malayanus remains problematic; see Csorba et al. (2003). This complex may include more than one species, see Csorba et al. (2003).	885887A2FFE98A0FFF6FF0DBF22AD0C8	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	309	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFD18A36F899F8EEFC67DB93.xml	Rhinolophus borneensis	Rhinolophidae	Rhinolophus	borneensis	Peters	1861	Bornean Horseshoe Bat @en | Rhinolophe de Bornéo @fr | Borneo-Hufeisennase @de | Herradura de Borneo @es	Rhinolophus borneensis Peters, 1861 , “ Labuan [Island], Borneo ,” Malaysia . Traditionally included in the megaphyllus species group, but this needs to be investigated further using genetic techniques, as A borneensis seems to be sister to a clade containing members of the pusillus species group. All mainland South-east Asian populations are here considered to belong to A chaseni , while Indonesian populations are tentatively included in A borneensis . The identity of Peninsular Malaysian populations is uncertain as no genetic data have been used for testing, so they are here listed under this species, pending further studies. This species may represent a species complex. Three subspecies are recognized.	. b. borneensis Meters, 1861 — Borneo, as well as nearby Banggi, Labuan , and Karimata Is . R. b. importunus Chasen, 1939 — C & SWJava, including Nusakambanagan I off S Java . . b. spadix G. S. Miller, 1901 — South Natuna Is. Taxonomic identity of specimens collected in NW Peninsular Malaysia is uncertain. Also, either this species or the Sulawesi Horseshoe Bat (. celebensis ) was recently captured in Lampung Province, Sumatra, but in that study the two species were not differentiated.	Tail 21-26 mm , ear 17-19 mm , forearm 40-44 mm ; weight 7-8- 5 g . Dorsal pelage is dark brown (brown morph) to bright reddish brown (orange morph), while ventral pelage is paler. Ears are medium-sized and brown; noseleaf too is brown. Noseleaf has moderately long, nearly cuneate lancet; connecting process is high and rounded, higher than sella; sella is narrow with slight constriction above midpoint and parallel or nearly parallel lateral margins; horseshoe is wide (c.8- 7 mm ), nearly covers whole muzzle, has a relatively deep median emargination, and has visible lateral leaflets. Lower lip has three mental grooves. Skull is small but relatively robustly built ( zygomatic width generally larger than mastoid width, occasionally subequal); zygomatic arch is highly arched; anterior median swellings are bulbous and longer than they are wide; lateral and posterior swellings are well developed; sagittal crest is weakly to moderately developed; frontal depression is moderately developed; supraorbital ridges are prominent and more or less sharp. P2 is reduced but almost always within tooth row, separating C1 and P4; P3 is moderate to small in size and usually completely displaced labially, nearly always allowing P2 and P4 to touch (rarely slightly separated).	Primary and secondary tropical forests at elevations from sea level up to C . 1000 m . Recorded in dipterocarp forest in Sarawak , Borneo.	The Bornean Horseshoe Bat feeds on insects and uses perches to feed on once the insect is captured.	No information.	Bornean Horseshoe Bats are nocturnal, foraging at night and roosting during the day. They are known to roost in cave systems in Borneo as well as in hollow bamboos and trees, among young leaves of bananas, and in rock crevices. Call shape is FM/CF/FM and the peak F has been recorded at 81-8 kHz in Sabah , Borneo.	Bornean Horseshoe Bats are known to roost in large colonies of several hundred individuals.	Classified as Least Concern on The IUCN Red List. The Bornean Horseshoe Bat has a wide distribution and is considered common at least in Borneo, which comprises most of its range. It is rare in some regions, especially in Java . Populations are probably most threatened by habitat loss and fragmentation due to deforestation and agricultural expansion.	Corbet & Hill (1992) | Csorba, Fuisz & Kelen (1997) | Csorba, Ujhelyi & Thomas (2003) | Francis et al. (2010) | Hall (1996) | Huang, J. C .C. et al. (2014) | Hutson, Kingston & rancis (2008a) | Kruskop (2011) | Lim et al. (2017) | Mohd-Azlan et al. (2008) | Rahman et al. (2011)	https://zenodo.org/record/3750026/files/figure.png	60 . Bornean Horseshoe Bat Rhinolophus borneensis French: Rhinolophe de Bornéo / German : Borneo-Hufeisennase / Spanish : Herradura de Borneo Taxonomy. Rhinolophus borneensis Peters, 1861 , “ Labuan [Island], Borneo ,” Malaysia . Traditionally included in the megaphyllus species group, but this needs to be investigated further using genetic techniques, as A borneensis seems to be sister to a clade containing members of the pusillus species group. All mainland South-east Asian populations are here considered to belong to A chaseni , while Indonesian populations are tentatively included in A borneensis . The identity of Peninsular Malaysian populations is uncertain as no genetic data have been used for testing, so they are here listed under this species, pending further studies. This species may represent a species complex. Three subspecies are recognized. Subspecies and Distribution. . b. borneensis Meters, 1861 — Borneo, as well as nearby Banggi, Labuan , and Karimata Is . R. b. importunus Chasen, 1939 — C & SWJava, including Nusakambanagan I off S Java . . b. spadix G. S. Miller, 1901 — South Natuna Is. Taxonomic identity of specimens collected in NW Peninsular Malaysia is uncertain. Also, either this species or the Sulawesi Horseshoe Bat (. celebensis ) was recently captured in Lampung Province, Sumatra, but in that study the two species were not differentiated. Descriptive notes. Tail 21-26 mm , ear 17-19 mm , forearm 40-44 mm ; weight 7-8- 5 g . Dorsal pelage is dark brown (brown morph) to bright reddish brown (orange morph), while ventral pelage is paler. Ears are medium-sized and brown; noseleaf too is brown. Noseleaf has moderately long, nearly cuneate lancet; connecting process is high and rounded, higher than sella; sella is narrow with slight constriction above midpoint and parallel or nearly parallel lateral margins; horseshoe is wide (c.8- 7 mm ), nearly covers whole muzzle, has a relatively deep median emargination, and has visible lateral leaflets. Lower lip has three mental grooves. Skull is small but relatively robustly built ( zygomatic width generally larger than mastoid width, occasionally subequal); zygomatic arch is highly arched; anterior median swellings are bulbous and longer than they are wide; lateral and posterior swellings are well developed; sagittal crest is weakly to moderately developed; frontal depression is moderately developed; supraorbital ridges are prominent and more or less sharp. P2 is reduced but almost always within tooth row, separating C1 and P4; P3 is moderate to small in size and usually completely displaced labially, nearly always allowing P2 and P4 to touch (rarely slightly separated). Habitat. Primary and secondary tropical forests at elevations from sea level up to C . 1000 m . Recorded in dipterocarp forest in Sarawak , Borneo. Food and Feeding. The Bornean Horseshoe Bat feeds on insects and uses perches to feed on once the insect is captured. Breeding. No information. Activity patterns. Bornean Horseshoe Bats are nocturnal, foraging at night and roosting during the day. They are known to roost in cave systems in Borneo as well as in hollow bamboos and trees, among young leaves of bananas, and in rock crevices. Call shape is FM/CF/FM and the peak F has been recorded at 81-8 kHz in Sabah , Borneo. Movements, Home range and Social organization. Bornean Horseshoe Bats are known to roost in large colonies of several hundred individuals. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Bornean Horseshoe Bat has a wide distribution and is considered common at least in Borneo, which comprises most of its range. It is rare in some regions, especially in Java . Populations are probably most threatened by habitat loss and fragmentation due to deforestation and agricultural expansion. Bibliography. Corbet & Hill (1992), Csorba, Fuisz & Kelen (1997), Csorba, Ujhelyi & Thomas (2003), Francis et al. (2010), Hall (1996), Huang, J. C .C. et al. (2014), Hutson, Kingston & rancis (2008a), Kruskop (2011), Lim et al. (2017), Mohd-Azlan et al. (2008), Rahman et al. (2011).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus borneensis	Rhinolophus		borneensis	Peters	1861	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1872:49:00	Bornean Horseshoe Bat	<b> chaseni </b>Sanborn, 1939;<b> importunus </b>Chasen, 1939;<b> spadix </b>Miller, 1901.	Malaysia, N Borneo, Sabah, Labuan Isl.	Borneo; Labuan and Banguey Isls; NW Peninsular Malaysia (Malaysia); Java, Karimata Isls, and South Natuna Isls (Indonesia)	Not listed.	Least Concern	 megaphyllus species group. Does not include chaseni ; see Francis et al. (2010), Kruskop (2011), and Abramov et al. (2018). Includes importunus ; see Hill (1983). Formerly included javanicus , celebensis , madurensis , and parvus (e.g., by Goodwin [1979] and Hill and Thonglongya [1972]) but see Hill (1983) and Kitchener et al.(1995a). Type material discussed by Csorba (2002). Subspecies limits are somewhat unclear, and the relationships of various forms to celebensis and malayanus remains problematic; see Csorba et al. (2003). This complex may include more than one species, seeCsorba et al. (2003).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus borneensis	23	Bornean Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	borneensis	W. Peters	1861	0	Rhinolophus_Borneensis	Peters, W. C. H. (1861). Ãœber die von Hrn. F. Jagor bisher auf Malacca, Borneo, Java und den Philippinen gesammelten Säugethiere aus den Ordnungen der Halbaffen, Pelzflatterer und Flederthiere. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1861, 709.	https://www.biodiversitylibrary.org/item/118969#page/99/mode/1up	ZMB 2533.1, ZMB 2533.2 [syntypes]		"Labuan [Island], Borneo," Malaysia.			borneensis W. Peters, 1861|spadix G. S. Miller, 1901|importunus Chasen, 1939	previously included R. chaseni	Francis, C. M., & Barrett, P. (2008). A field guide to the mammals of South-East Asia. New Holland Publishers.	Indonesia|Malaysia|Brunei	Asia	Indomalaya	LC	0	0	0	Rhinolophus_borneensis	0	sciname match	Rhinolophus_borneensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19527	Rhinolophus borneensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	borneensis	Peters, 1861	This is most likely a species complex (Csorba et al . 2003). Lekagul and McNeely (1977) synonymized this species with Rhinolophus robinsoni , but robinsoni is now considered a separate species (Simmons 2005). The distribution of R. borneensis may be confused, because of the unresolved taxonomy of the group.	20000000	Rhinolophus borneensis	Least Concern		2020	2018-08-04 00:00:00 UTC	3.1	English	Listed as Least Concern as the species is widespread and fairly common, and there are no known major threats.	This species inhabits primary and secondary forest. Diurnal roosts include hollow bamboos, young leaves of bananas, hollow trees and rock crevices. Payne et al. (1985) mention that the species roosts in caves, sometimes in colonies of several hundred individuals. It uses perches to feed.	There are no major threats to this species and it does seem to tolerate some disturbance of its habitat.	The species is relatively common in Borneo, but rare in most other places. Recorded from Gunung Halimun, West Java as rare by Suyanto (2003). Borissenko and Kruskop (2003) noted that this is one of the most common forest bats in southern Viet Nam.	Unknown	This species is known from all of Borneo (although it has yet to be confirmed in Brunei) Viet Nam, Cambodia, Lao PDR, the Malay Peninsula (Simmons 2005, Shazali et al. 2018),from several locations in Java, including Yogyakarta, Nusakambanagan Island just off south central Java, Labuan and Banguey Islands, Karimata Islands, and South Natuna Islands, and Sumatra, Indonesia. It is suspected to occur in Thailand but has yet to be confirmed there.		Terrestrial	It is recorded in many protected areas across its distribution in Southeast Asia. Further taxonomic studies are required to examine specimens that are found in Vietnam and the many Indonesian islands due to disjunct distribution to determine the distribution limits of the species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		borneensis	Peters	1861	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1872:49:00	Bornean Horseshoe Bat	<b> chaseni </b>Sanborn, 1939;<b> importunus </b>Chasen, 1939;<b> spadix </b>Miller, 1901.	Malaysia, N Borneo, Sabah, Labuan Isl.	Borneo; Labuan and Banguey Isls; NW Peninsular Malaysia (Malaysia); Java, Karimata Isls, and South Natuna Isls (Indonesia)	Not listed.	Least Concern	 megaphyllus species group. Does not include chaseni ; see Francis et al. (2010), Kruskop (2011), and Abramov et al. (2018). Includes importunus ; see Hill (1983). Formerly included javanicus , celebensis , madurensis , and parvus (e.g., by Goodwin [1979] and Hill and Thonglongya [1972]) but see Hill (1983) and Kitchener et al.(1995a). Type material discussed by Csorba (2002). Subspecies limits are somewhat unclear, and the relationships of various forms to celebensis and malayanus remains problematic; see Csorba et al. (2003). This complex may include more than one species, seeCsorba et al. (2003).	Rhinolophus borneensis	1004663	23	Bornean Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	borneensis	W. Peters	1861	0	Rhinolophus_Borneensis	Peters, W. C. H. (1861). Ãœber die von Hrn. F. Jagor bisher auf Malacca, Borneo, Java und den Philippinen gesammelten Säugethiere aus den Ordnungen der Halbaffen, Pelzflatterer und Flederthiere. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1861, 709.	https://www.biodiversitylibrary.org/item/118969#page/99/mode/1up	ZMB 2533.1, ZMB 2533.2 [syntypes]		"Labuan [Island], Borneo," Malaysia.			borneensis W. Peters, 1861|spadix G. S. Miller, 1901|importunus Chasen, 1939	previously included R. chaseni	Francis, C. M., & Barrett, P. (2008). A field guide to the mammals of South-East Asia. New Holland Publishers.				Indonesia|Malaysia|Brunei	Asia	Indomalaya	LC	0	0	0	Rhinolophus_borneensis	0	sciname match	Rhinolophus_borneensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_borneensis	1004663	23	Bornean Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	borneensis	W. C. H. Peters	0	Rhinolophus Borneensis	Peters, W.C.H. 1861. Hr. W. Peters berichtet über die von Hrn. F. Jagor bisher auf Malacca, Borneo, Java und den Philippinen gesammelten Säugethiere aus den Ordnungen der Halbaffen, Pelzflatterer und Flederthiere. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1861:706-712.	https://www.biodiversitylibrary.org/page/38646576	ZMB 2533.1, ZMB 2533.2	syntypes		"Labuan [Island], Borneo," Malaysia.			previously included R. chaseni	Francis, C. M. (2008). A field guide to the mammals of South-East Asia. New Holland Publishers.				Indonesia|Malaysia|Brunei	Asia	Indomalaya	LC	0	0	0	Rhinolophus_borneensis	0	sciname match	Rhinolophus_borneensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		borneensis	Peters	1861	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1872:49:00	Bornean Horseshoe Bat	chaseni Sanborn, 1939; importunus Chasen, 1939; spadix Miller, 1901.	Malaysia, N Borneo, Sabah, Labuan Isl.	Borneo; Labuan and Banguey Isls; NW Peninsular Malaysia (Malaysia); Java, Karimata Isls, and South Natuna Isls (Indonesia)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19527/21982599/' target='_blank'>Least Concern</a>	megaphyllusspecies group. Does not include chaseni; see Francis et al. (2010), Kruskop (2011), and Abramov et al. (2018). Includes importunus; see Hill (1983). Formerly included javanicus, celebensis, madurensis, and parvus (e.g., by Goodwin [1979] and Hill and Thonglongya [1972]) but see Hill (1983) and Kitchener et al.(1995a). Type material discussed by Csorba (2002). Subspecies limits are somewhat unclear, and the relationships of various forms to celebensis and malayanus remains problematic; see Csorba et al. (2003). This complex may include more than one species, seeCsorba et al. (2003).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus borneensis; Rhinolophus borneensis; Rhinolophus borneensis; Rhinolophus borneensis; Rhinolophus borneensis; Rhinolophus borneensis; borneensis; chaseni; importunus; spadix; borneensis; importunus; spadix; chaseni; importunus; spadix; borneensis; spadix; importunus; Bornean Horseshoe Bat; Rhinolophe de Bornéo; Borneo-Hufeisennase; Herradura de Borneo; Bornean Horseshoe Bat; Bornean Horseshoe Bat; Bornean Horseshoe Bat; R. borneensis