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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1347	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Rhinolophus clivosus [synonym of]	Rhinolophus bocharicus [synonym of]	Rhinolophus clivosus [synonym of]	Rhinolophus clivosus bocharicus	Rhinolophus bocharicus	Rhinolophus bocharicus	Rhinolophus bocharicus	Rhinolophus bocharicus	Rhinolophus bocharicus	Rhinolophus bocharicus	Rhinolophus bocharicus	Rhinolophus bocharicus	Rhinolophus bocharicus	Rhinolophus bocharicus		[MSW3] ferrumequinum species group. Included in clivosus by Aellen (1959), but see Hanák (1969), Felten (1977), DeBlase (1980), Gromov and Baranova (1981), Pavlinov and Rossolimo (1987), and Horácek et al. (2000). Apparently does not include rubiginosus, which is here placed in ferrumequinum following Csorba et al. (2003).; [HMW] Rhinolophus bocharicus Kastschenko & Akimov, 1917 , Murgab River , Turkmenistan . Rhinolophus bocharicus is in the ferrumequinum species group based on morphology and genetic evidence and has been variously included as a subspecies of R ferrumequinum or R clivosus . Detailed morphological work concluded that R bocharicus is distinct from both species and that it and R clivosus might have evolved from populations of R ferrumequinum . The name rubiginosus by N. Gubarev in 1941 is included under R ferrumequinum as a synonym instead of a subspecies of R bocharicus . A single specimen attributed to R bocharicus from Iran was actually a misidentifiedjuvenile R ferrumequinum . Monotypic.; [batnames2022]  ferrumequinum species group. Included in clivosus by Aellen (1959), but see HanÃ¡k (1969), Felten (1977), DeBlase (1980), Gromov and Baranova (1981),Pavlinov and Rossolimo (1987), and HorÃ¡cek et al. (2000). Apparently does not include rubiginosus , which is here placed in ferrumequinum following Csorba et al. (2003).; [IUCN] Koopman (1993) included this in synonymy under R. clivosus , whereas others maintain it as a separate and a monotypic species (Simmons 2005, Benda and Vallo 2012, Srinivasulu and Srinivasulu 2012, Benda and Gaisler 2015).; [batnames2023]  ferrumequinum species group. Included in clivosus by Aellen (1959), but see HanÃ¡k (1969), Felten (1977), DeBlase (1980), Gromov and Baranova (1981),Pavlinov and Rossolimo (1987), and HorÃ¡cek et al. (2000). Apparently does not include rubiginosus , which is here placed in ferrumequinum following Csorba et al. (2003).; [batnames2025_1.7] ferrumequinum species group. Included in clivosus by Aellen (1959), but see HanÃ¡k (1969), Felten (1977), DeBlase (1980), Gromov and Baranova (1981), Pavlinov and Rossolimo (1987), HorÃ¡cek et al. (2000), and Uvizl et al. (2024). Uvizl et al. (2024) found no evidence of subspecies within this taxon. Apparently does not include rubiginosus, which is here placed in ferrumequinum following Csorba et al. (2003).											The name rubiginosus by N. Gubarev in 1941 is included under R ferrumequinum as a synonym instead of a subspecies of R bocharicus? No?			bocharicus	Koopman (1993) included this in synonymy under R. clivosus , whereas others maintain it as a separate and a monotypic species (Simmons 2005, Benda and Vallo 2012, Srinivasulu and Srinivasulu 2012, Benda and Gaisler 2015).			bocharicus 	bocharicus 			bocharicus Kastschenko & Akimov, 1918						N/A							Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Turkestan, NE Iran, N Afghanistan																															NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus bocharicus	Rhinolophus		bocharicus	Kastchenko and Akimov		1917		Annu. Mus. Zool. Acad. St. Petersb.	22		221		Central Asian Horseshoe Bat	Turkmenistan, Murgab River.	Kyrgyzstan, W Tajikistan, NE Iran, Uzbekistan, Turkmenistan, Afghanistan, possibly N Pakistan.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		ferrumequinum species group. Included in clivosus by Aellen (1959), but see Hanák (1969), Felten (1977), DeBlase (1980), Gromov and Baranova (1981), Pavlinov and Rossolimo (1987), and Horácek et al. (2000). Apparently does not include rubiginosus, which is here placed in ferrumequinum following Csorba et al. (2003).	885887A2FFD18A36F899F8EEFC67DB93	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	295	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFDF8A39FF6AF7FEF4D7DD69.xml	Rhinolophus bocharicus	Rhinolophidae	Rhinolophus	bocharicus	Kastschenko & Akimov	1918	Bokhara Horseshoe Bat @en | Rhinolophe du Boukhara @fr | Buchara-Hufeisennase @de | Herradura de Bujarâ @es | Central Asian Horseshoe Bat @en	Rhinolophus bocharicus Kastschenko & Akimov, 1917 , Murgab River , Turkmenistan . Rhinolophus bocharicus is in the ferrumequinum species group based on morphology and genetic evidence and has been variously included as a subspecies of R ferrumequinum or R clivosus . Detailed morphological work concluded that R bocharicus is distinct from both species and that it and R clivosus might have evolved from populations of R ferrumequinum . The name rubiginosus by N. Gubarev in 1941 is included under R ferrumequinum as a synonym instead of a subspecies of R bocharicus . A single specimen attributed to R bocharicus from Iran was actually a misidentifiedjuvenile R ferrumequinum . Monotypic.	S Kazakhstan , Uzbekistan , W Kyrgyzstan , Turkmenistan , C & W Tajikistan , and N Afghanistan .	Head-body c . 38-48 mm , tail 22-32 mm , ear 19- 3-24 mm , hindfoot C. 13 mm , forearm 47- 5-53 mm ; weight 9-20 g . Dorsal pelage is pale smoky gray-brown, being darker on shoulders (hairs have whitish bases); venter is whitish gray. There is no orange morph. Males lack axillary tufts. Ears are medium in length. Noseleaf has hastate lancet; connecting process is tall and distinctively rounded in side view ; sella is narrow, with concave sides and blunt tip; and horseshoe is comparatively narrow at 5-4-7- 3 mm wide and does not completely cover muzzle. Lower lip has three very indistinct grooves. Baculum is very similar to that of the Greater Horseshoe Bat ( AE ferrumequinum ) but generally lacks strong protuberances on ventral sides of basal cone, and lancet of shaft is generally longer than in the Greater Horseshoe Bat. Skull is robust (zygomatic width is much larger than mastoid width); nasal swellings are underdeveloped anteriorly but well developed posteriorly; sagittal crest is moderately developed, and frontal depression is deep to very deep; and supraorbital crests are low. P2 is tiny and completely displaced labially or occasionally absent, allowing C1 and P4 to come in contact, and P3 is tiny and displaced labially, so P2 and P4 always touch. Dental formula is 11/2, C 1/1, P 2/3, M 3/3 (x2) = 32 or 11/2, C 1/1, P 1/3, M 3/3 (x2) = 30.	Arid mountain foothills.	The Bokhara Horseshoe Bat is insectivorous, feeding largely on lepidopterans and occasionally coleopterans and other insects. It forages close to the ground and probably around cluttered with vegetation areas.	Births of Bokhara Horseshoe Bats have been recorded in June to earlyJuly.	The Bokhara Horseshoe Bat is nocturnal, foraging primarily at dusk. There seems to be a migration between winter and summer for some individuals traveling from Central Asia to Afghanistan (north to south). Nevertheless, some individuals remain in the same cave throughout the year in Uzbekistan . Bokhara Horseshoe Bats are known to have day roosts in caves and abandoned mine shafts.	Bokhara Horseshoe Bats roost in colonies of several hundred (up to 600) individuals and rarely in small groups. They have been recorded roosting commonly with Geoffroy’s Myotis ( Myotis emarginatus) and occasionally with Greater Horseshoe Bats and Lesser Myotis ( Myotis blythii ). Female and male Bokhara Horseshoe Bats segregate into maternal and non-matemal colonies during the breeding season. Males move back to form mixed colonies after young can fly and forage on their own.	Classified as Least Concern on The IUCN Red List. There are no major risks to the Bokhara Horseshoe Bat, but habitat loss and cave destruction and modification might be threats in some regions.	Bailey et al. (2016) | Benda & Gaisler (2015) | Benda, Aulagnier et al. (2008) | Benda, Faizolâhi et al. (2012) | Benda, Hanâk & Ôervenÿ (2011) | Bobrinski étal. (1944) | Csorba étal. (2003) | Gubarev (1941) | Hanâk (1969) | Kastschenko & Akimov (1917) | Strelkov (1971) | Thomas (1997)	https://zenodo.org/record/3749968/files/figure.png	30 . Bokhara Horseshoe Bat Rhinolophus bocharicus French: Rhinolophe du Boukhara / German: Buchara-Hufeisennase / Spanish: Herradura de Bujarâ Other common names: Central Asian Horseshoe Bat Taxonomy. Rhinolophus bocharicus Kastschenko & Akimov, 1917 , Murgab River , Turkmenistan . Rhinolophus bocharicus is in the ferrumequinum species group based on morphology and genetic evidence and has been variously included as a subspecies of R ferrumequinum or R clivosus . Detailed morphological work concluded that R bocharicus is distinct from both species and that it and R clivosus might have evolved from populations of R ferrumequinum . The name rubiginosus by N. Gubarev in 1941 is included under R ferrumequinum as a synonym instead of a subspecies of R bocharicus . A single specimen attributed to R bocharicus from Iran was actually a misidentifiedjuvenile R ferrumequinum . Monotypic. Distribution. S Kazakhstan , Uzbekistan , W Kyrgyzstan , Turkmenistan , C & W Tajikistan , and N Afghanistan . Descriptive notes. Head-body c . 38-48 mm , tail 22-32 mm , ear 19- 3-24 mm , hindfoot C. 13 mm , forearm 47- 5-53 mm ; weight 9-20 g . Dorsal pelage is pale smoky gray-brown, being darker on shoulders (hairs have whitish bases); venter is whitish gray. There is no orange morph. Males lack axillary tufts. Ears are medium in length. Noseleaf has hastate lancet; connecting process is tall and distinctively rounded in side view ; sella is narrow, with concave sides and blunt tip; and horseshoe is comparatively narrow at 5-4-7- 3 mm wide and does not completely cover muzzle. Lower lip has three very indistinct grooves. Baculum is very similar to that of the Greater Horseshoe Bat ( AE ferrumequinum ) but generally lacks strong protuberances on ventral sides of basal cone, and lancet of shaft is generally longer than in the Greater Horseshoe Bat. Skull is robust (zygomatic width is much larger than mastoid width); nasal swellings are underdeveloped anteriorly but well developed posteriorly; sagittal crest is moderately developed, and frontal depression is deep to very deep; and supraorbital crests are low. P2 is tiny and completely displaced labially or occasionally absent, allowing C1 and P4 to come in contact, and P3 is tiny and displaced labially, so P2 and P4 always touch. Dental formula is 11/2, C 1/1, P 2/3, M 3/3 (x2) = 32 or 11/2, C 1/1, P 1/3, M 3/3 (x2) = 30. Habitat. Arid mountain foothills. Food and Feeding. The Bokhara Horseshoe Bat is insectivorous, feeding largely on lepidopterans and occasionally coleopterans and other insects. It forages close to the ground and probably around cluttered with vegetation areas. Breeding. Births of Bokhara Horseshoe Bats have been recorded in June to earlyJuly. Activity patterns. The Bokhara Horseshoe Bat is nocturnal, foraging primarily at dusk. There seems to be a migration between winter and summer for some individuals traveling from Central Asia to Afghanistan (north to south). Nevertheless, some individuals remain in the same cave throughout the year in Uzbekistan . Bokhara Horseshoe Bats are known to have day roosts in caves and abandoned mine shafts. Movements, Home range and Social organization. Bokhara Horseshoe Bats roost in colonies of several hundred (up to 600) individuals and rarely in small groups. They have been recorded roosting commonly with Geoffroy’s Myotis ( Myotis emarginatus) and occasionally with Greater Horseshoe Bats and Lesser Myotis ( Myotis blythii ). Female and male Bokhara Horseshoe Bats segregate into maternal and non-matemal colonies during the breeding season. Males move back to form mixed colonies after young can fly and forage on their own. Status and Conservation. Classified as Least Concern on The IUCN Red List. There are no major risks to the Bokhara Horseshoe Bat, but habitat loss and cave destruction and modification might be threats in some regions. Bibliography. Bailey et al. (2016), Benda & Gaisler (2015), Benda, Aulagnier et al. (2008), Benda, Faizolâhi et al. (2012), Benda, Hanâk & Ôervenÿ (2011), Bobrinski étal. (1944), Csorba étal. (2003), Gubarev (1941), Hanâk (1969), Kastschenko & Akimov (1917), Strelkov (1971), Thomas (1997).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus bocharicus	Rhinolophus		bocharicus	Kastschenko & Akimov	1917	0	Annu. Mus. Zool. Acad. St. Petersb.	1.0701	Central Asian Horseshoe Bat	None.	Turkmenistan, Murgab River.	Kyrgyzstan, W Tajikistan, NE Iran, Uzbekistan, Turkmenistan, Afghanistan, possibly N Pakistan.	Not listed.	Least Concern	 ferrumequinum species group. Included in clivosus by Aellen (1959), but see HanÃ¡k (1969), Felten (1977), DeBlase (1980), Gromov and Baranova (1981),Pavlinov and Rossolimo (1987), and HorÃ¡cek et al. (2000). Apparently does not include rubiginosus , which is here placed in ferrumequinum following Csorba et al. (2003).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus bocharicus	23	Bokhara Horseshoe Bat	Central Asian Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	bocharicus	Kastschenko & Akimov	1917	0	Rhinolophus_bocharicus	Katschenko, N. F., & Akimov, M. P. (1917). Rhinolophus bocharicus sp. n. EÅ¾egodnik ZoologiÄeskago MuzeÃ¢ Akademii Nauk, 22, 221.	https://www.biodiversitylibrary.org/item/34689#page/431/mode/1up			Murgab River, Turkmenistan.			bocharicus Kastschenko & Akimov, 1917	NA	NA	Kazakhstan|Uzbekistan|Kyrgyzstan|Turkmenistan|Tajikistan|Afghanistan|Iran?	Asia	Palearctic	LC	0	0	0	Rhinolophus_bocharicus	0	sciname match	Rhinolophus_bocharicus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19526	Rhinolophus bocharicus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	bocharicus	Kastschenko &; Akimov, 1917	Koopman (1993) included this in synonymy under R. clivosus , whereas others maintain it as a separate and a monotypic species (Simmons 2005, Benda and Vallo 2012, Srinivasulu and Srinivasulu 2012, Benda and Gaisler 2015).	20000000	Rhinolophus bocharicus	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Though a reasonably widespread species, it is uncommon in certain parts of its range but is believed to be stable hence listed as Least Concern.	Distributed in desertified mountain foothills. Inhabits caves and deserted mine galleries. Females during pregnancy and lactation period form colonies separate from males. Usually those colonies reach several hundred in number, rarely small groups. Delivery occurs in June-beginning of July. After young are able to fly and feed independently, males move to form mixed colonies. Those mixed colonies are often common roosts with Myotis emarginatus , sometimes with Rhinolophus ferrumequinum , and/or Myotis blythi . Mating occurs after lactation period or during winter. The species hibernates in deep caves. Preys at dusk, flying low above the ground, feed mainly on butterflies, sometimes on beetles andother insects.	Habitat loss and artificial cave modification are threats, but are not thought to pose a major risk to the species at present.	There are no data on population size.	Stable	Distributed in Central Asia from south Turkmenistan to north Afghanistan (Csorba et al.  2003, Benda and Vallo 2012, Srinivasulu and Srinivasulu 2012, Benda and Gaisler 2015).		Terrestrial	There are no conservation measures applied to the species.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		bocharicus	Kastschenko & Akimov	1917	0	Annu. Mus. Zool. Acad. St. Petersb.	1.070139	Central Asian Horseshoe Bat	None.	Turkmenistan, Murgab River.	Kyrgyzstan, W Tajikistan, NE Iran, Uzbekistan, Turkmenistan, Afghanistan, possibly N Pakistan.	Not listed.	Least Concern	 ferrumequinum species group. Included in clivosus by Aellen (1959), but see HanÃ¡k (1969), Felten (1977), DeBlase (1980), Gromov and Baranova (1981),Pavlinov and Rossolimo (1987), and HorÃ¡cek et al. (2000). Apparently does not include rubiginosus , which is here placed in ferrumequinum following Csorba et al. (2003).	Rhinolophus bocharicus	1004662	23	Bokhara Horseshoe Bat	Central Asian Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	bocharicus	Kastschenko & Akimov	1917	0	Rhinolophus_bocharicus	Katschenko, N. F., & Akimov, M. P. (1917). Rhinolophus bocharicus sp. n. EÅ¾egodnik ZoologiÄeskago MuzeÃ¢ Akademii Nauk, 22, 221.	https://www.biodiversitylibrary.org/item/34689#page/431/mode/1up			Murgab River, Turkmenistan.			bocharicus Kastschenko & Akimov, 1917	NA	NA				Kazakhstan|Uzbekistan|Kyrgyzstan|Turkmenistan|Tajikistan|Afghanistan|Iran?	Asia	Palearctic	LC	0	0	0	Rhinolophus_bocharicus	0	sciname match	Rhinolophus_bocharicus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_bocharicus	1004662	23	Bokhara Horseshoe Bat	Central Asian Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	bocharicus	Kastschenko & Akimov	0	Rhinolophus bocharicus	Kastschenko, N.F. and Akimov, M.N. 1918. _Rhinolophus bocharicus_ sp. n. Annuaire du MusÃ©e Zoologique de l'AcadÃ©mie des Sciences de Russie 22(1-3):221-223.	https://www.biodiversitylibrary.org/page/8482836	ZIN S. 8702	lectotype		Murgab River, Turkmenistan.	39.65	66.96667	NA	NA				Kazakhstan|Uzbekistan|Kyrgyzstan|Turkmenistan|Tajikistan|Afghanistan|Iran?	Asia	Palearctic	LC	0	0	0	Rhinolophus_bocharicus	0	sciname match	Rhinolophus_bocharicus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		bocharicus	Kastschenko & Akimov	1917	0	Annu. Mus. Zool. Acad. St. Petersb.	1.070139	Central Asian Horseshoe Bat	None.	Turkmenistan, Murgab River	Kyrgyzstan, W Tajikistan, NE Iran, Uzbekistan, Turkmenistan, Afghanistan, possibly N Pakistan	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19526/21983564/' target='_blank'>Least Concern</a>	ferrumequinum species group. Included in clivosus by Aellen (1959), but see HanÃ¡k (1969), Felten (1977), DeBlase (1980), Gromov and Baranova (1981), Pavlinov and Rossolimo (1987), HorÃ¡cek et al. (2000), and Uvizl et al. (2024). Uvizl et al. (2024) found no evidence of subspecies within this taxon. Apparently does not include rubiginosus, which is here placed in ferrumequinum following Csorba et al. (2003).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus bocharicus; Rhinolophus bocharicus; Rhinolophus bocharicus; Rhinolophus bocharicus; Rhinolophus bocharicus; Rhinolophus bocharicus; The name rubiginosus by N. Gubarev in 1941 is included under R ferrumequinum as a synonym instead of a subspecies of R bocharicus? No?; bocharicus; Bokhara Horseshoe Bat; Rhinolophe du Boukhara; Buchara-Hufeisennase; Herradura de Bujarâ; Central Asian Horseshoe Bat; Bokhara Horseshoe Bat; Central Asian Horseshoe Bat; Central Asian Horseshoe Bat; Central Asian Horseshoe Bat; R. bocharicus
