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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1344	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Rhinolophus luctus [synonym of]	Rhinolophus luctus beddomei	Rhinolophus beddomei	Rhinolophus beddomei	Rhinolophus beddomei	Rhinolophus beddomei	Rhinolophus beddomei	Rhinolophus beddomei	Rhinolophus beddomei	Rhinolophus beddomei	Rhinolophus beddomei	Rhinolophus beddomei		[MSW3] trifoliatus species group. Distinct from luctus; see Topál and Csorba (1992), Bates and Harrison (1997), and Hendrichsen et al. (2001a).; [HMW] Rhinolophus beddomei K. Andersen, 1905 , “ Wynaad [= Wayanad], Mysore [= Mysuru ], S[outh] . India .” Rhinolophus beddomei is included in the trifoliatus species group and appears to be close to R. trifoliatus , R.sedulus , and R. luctus. Two subspecies are recognized.; [batnames2022]  trifoliatus species group. Distinct from luctus ; see Top&aacute;l and Csorba (1992), Bates and Harrison (1997), and Hendrichsen et al. (2001a).; [IUCN] This species belongs to trifoliatus species group. Earlier, the taxa beddomei Andersen 1905 and sobrinus Andersen, 1918 were included under Rhinolophus luctus Temminck, 1835 (Ellerman and Morrison-Scott 1951, Corbet and Hill 1992, Koopman 1993). Here Rhinolophus beddomei Andersen, 1905 is treated as a distinct species (TopÃ¡l and Csorba 1992, Bates and Harrison 1997, Simmons 2005, Srinivasulu and Srinivasulu 2012).; [batnames2023]  trifoliatus species group. Distinct from luctus ; see Top&aacute;l and Csorba (1992), Bates and Harrison (1997), and Hendrichsen et al. (2001a).; [batnames2025_1.7] trifoliatus species group. Distinct from luctus; see Top&aacute;l and Csorba (1992), Bates and Harrison (1997), and Hendrichsen et al. (2001a).								beddomei, sobrinus		beddomei, sobrinus		beddomei, sobrinus		beddomei, sobrinus	This species belongs to trifoliatus species group. Earlier, the taxa beddomei Andersen 1905 and sobrinus Andersen, 1918 were included under Rhinolophus luctus Temminck, 1835 (Ellerman and Morrison-Scott 1951, Corbet and Hill 1992, Koopman 1993). Here Rhinolophus beddomei Andersen, 1905 is treated as a distinct species (TopÃ¡l and Csorba 1992, Bates and Harrison 1997, Simmons 2005, Srinivasulu and Srinivasulu 2012).	beddomei, sobrinus		beddomei, sobrinus	beddomei, sobrinus	beddomei, sobrinus		beddomei Andersen, 1905|sobrinus Andersen, 1918						N/A																																								_R. b. beddomei_ Andersen, 1905; _R. b. sobrinus_ Andersen, 1918			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus beddomei	Rhinolophus		beddomei	K. Andersen		1905		Ann. Mag. Nat. Hist., ser. 7	16		253		Bedomme's Horseshoe Bat	India, Madras [= Kerala], Wynaad.	S India, Sri Lanka.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	sobrinus K. Andersen, 1918.	trifoliatus species group. Distinct from luctus; see Topál and Csorba (1992), Bates and Harrison (1997), and Hendrichsen et al. (2001a).	885887A2FFE88A01F8B0F2B9F98EDFED	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	330	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFFC8A1AF8B1FD56F286D6A0.xml	Rhinolophus beddomei	Rhinolophidae	Rhinolophus	beddomei	K. Andersen	1905	Rhinolophede Beddome @fr | Beddome-Wollhaar-Hufeisennase @de | Herradura de Beddome @es | Bedomme's Horseshoe Bat @en	Rhinolophus beddomei K. Andersen, 1905 , “ Wynaad [= Wayanad], Mysore [= Mysuru ], S[outh] . India .” Rhinolophus beddomei is included in the trifoliatus species group and appears to be close to R. trifoliatus , R.sedulus , and R. luctus. Two subspecies are recognized.	R. b. beddomei K. Andersen, 1905 — W & E India ( Maharashtra , Karnataka , Kerala , and Andhra Pradesh ). R.b. sobrinus K Andersen, 1918 - Sri Lanka .	Head-body 65-75 mm , tail 37-48 mm , ear 21-34 mm , hindfoot 12- 1-18 mm , forearm 54-9—64- 3 mm . Fur is long and woolly ; dorsal and ventral pelage is dark brown or black (hairs with paler tips). Ears are blackish and of medium length. Noseleaf is dark brown or blackish, and has long and pointed or rounded lancet; connecting process is relatively low and broadly rounded; sella is broad and widely rounded off with large circular lateral lappets on either side of base; horseshoe is very wide ( 11 mm ), covering muzzle, and has wide median emargination. Lower lip has one mental groove. Skull is robust (zygomatic breadth is larger than mastoid breadth); anterior median swellings are elongated but do not expand anteriorly to upper edge of nasal orifice, formed by a protruding bony rim; posterior swellings are more laterally situated; frontal depression is deep and long; supraorbital crests are sharp; sagittal crest is very well developed; canines are strong but relatively short. P2 is medium-sized and within tooth row or slightly displaced; P4 is small to medium-sized and extruded from the tooth row to a variable degree; P2 and P4 are almost in contact. Chromosomal complement has 2n = 32 and FN = 60.	Mainly found in dense dry and tropical moist forests. Elevational range is from sea level up to elevations of 1070 m .	Beddome’s Woolly Horseshoe Bat feeds on a variety of insects especially beetles and termites. It flies low over the ground while foraging, and will forage close to vegetation.	A pregnant female with one embryo was collected in mid-January in Sri Lanka . Young have been reported with females in Kerala , India , in mid-May , and in Sri Lanka in mid-March and late April. Females may not breed until two or three years old.	Beddome’s Woolly Horseshoe Bat roosts by day in small caves, dilapidated buildings, tree hollows, wells, culverts, and old and unused tunnels, or clings from overhanging rocky outcrops. It emerges late in the evening from its roosts to forage through the night. Call shape is FM/CF/FM with a peak F recorded at 38-5—38-7 kHz and durations of 48-2—58 milliseconds in the Western Ghats of India ; another study in the Western Ghats reported peak frequencies of 41-7-43-3 kHz (mean 42-8 kHz) and durations of 1-4-51-5 milliseconds (mean 25-2 milliseconds). A recording inside a cave in Sri Lanka produced peak frequencies of 41—62-5 kHz but this huge range must be an error, perhaps including more than one species.	Beddome’s Woolly Horseshoe Bat roosts alone or in pairs, or sometimes in groups of three or four individuals.	Classified as Least Concern on TheIUCNRed List. Beddome’s Woolly Horseshoe Bat is relatively widespread and common throughout much of its distribution. However, in Sri Lanka the species is regionally listed as Vulnerable according to the National Red List for the country , and is mostly restricted to old-growth forests with small populations. The species appears to be threatened most by deforestation for logging and agricultural expansion.	Bates & Harrison (1997) | Csorba et al. (2003) | Edirisinghe et al. (2016) | Koubfnovà et al. (2010) | Kusuminda et al. (2013) | Molur et al. (2002) | Naidu & Gururaj (1984) | Raghuram et al. (2014) | Rao et al. (2004) | Sinha (1973) | Soisook, Niyomwan et al. (2010) | Soisook, Struebig et al. (2015) | Srinivasulu & Molur (2008b) | Srinivasulu & Srinivasulu (2012) | Topàl & Csorba (1992) | Volleth et al. (2017) | Wordley et al. (2014)	https://zenodo.org/record/3750118/files/figure.png	103 . Beddome’s Woolly Horseshoe Bat Rhinolophus beddomei French: Rhinolophe de Beddome / German: Beddome-Wollhaar-Hufeisennase / Spanish: Herradura de Beddome Other common names: Bedomme's Horseshoe Bat Taxonomy. Rhinolophus beddomei K. Andersen, 1905 , “ Wynaad [= Wayanad], Mysore [= Mysuru ], S[outh] . India .” Rhinolophus beddomei is included in the trifoliatus species group and appears to be close to R. trifoliatus , R.sedulus , and R. luctus. Two subspecies are recognized. Subspecies and Distribution. R. b. beddomei K. Andersen, 1905 — W & E India ( Maharashtra , Karnataka , Kerala , and Andhra Pradesh ). R.b. sobrinus K Andersen, 1918 - Sri Lanka . Descriptive notes. Head-body 65-75 mm , tail 37-48 mm , ear 21-34 mm , hindfoot 12- 1-18 mm , forearm 54-9—64- 3 mm . Fur is long and woolly ; dorsal and ventral pelage is dark brown or black (hairs with paler tips). Ears are blackish and of medium length. Noseleaf is dark brown or blackish, and has long and pointed or rounded lancet; connecting process is relatively low and broadly rounded; sella is broad and widely rounded off with large circular lateral lappets on either side of base; horseshoe is very wide ( 11 mm ), covering muzzle, and has wide median emargination. Lower lip has one mental groove. Skull is robust (zygomatic breadth is larger than mastoid breadth); anterior median swellings are elongated but do not expand anteriorly to upper edge of nasal orifice, formed by a protruding bony rim; posterior swellings are more laterally situated; frontal depression is deep and long; supraorbital crests are sharp; sagittal crest is very well developed; canines are strong but relatively short. P2 is medium-sized and within tooth row or slightly displaced; P4 is small to medium-sized and extruded from the tooth row to a variable degree; P2 and P4 are almost in contact. Chromosomal complement has 2n = 32 and FN = 60. Habitat. Mainly found in dense dry and tropical moist forests. Elevational range is from sea level up to elevations of 1070 m . Food and Feeding. Beddome’s Woolly Horseshoe Bat feeds on a variety of insects especially beetles and termites. It flies low over the ground while foraging, and will forage close to vegetation. Breeding. A pregnant female with one embryo was collected in mid-January in Sri Lanka . Young have been reported with females in Kerala , India , in mid-May , and in Sri Lanka in mid-March and late April. Females may not breed until two or three years old. Activity patterns. Beddome’s Woolly Horseshoe Bat roosts by day in small caves, dilapidated buildings, tree hollows, wells, culverts, and old and unused tunnels, or clings from overhanging rocky outcrops. It emerges late in the evening from its roosts to forage through the night. Call shape is FM/CF/FM with a peak F recorded at 38-5—38-7 kHz and durations of 48-2—58 milliseconds in the Western Ghats of India ; another study in the Western Ghats reported peak frequencies of 41-7-43-3 kHz (mean 42-8 kHz) and durations of 1-4-51-5 milliseconds (mean 25-2 milliseconds). A recording inside a cave in Sri Lanka produced peak frequencies of 41—62-5 kHz but this huge range must be an error, perhaps including more than one species. Movements, Home range and Social organization. Beddome’s Woolly Horseshoe Bat roosts alone or in pairs, or sometimes in groups of three or four individuals. Status and Conservation. Classified as Least Concern on TheIUCNRed List. Beddome’s Woolly Horseshoe Bat is relatively widespread and common throughout much of its distribution. However, in Sri Lanka the species is regionally listed as Vulnerable according to the National Red List for the country , and is mostly restricted to old-growth forests with small populations. The species appears to be threatened most by deforestation for logging and agricultural expansion. Bibliography. Bates & Harrison (1997), Csorba eta /. (2003), Edirisinghe et al. (2016), Koubfnovà et al. (2010), Kusuminda et al. (2013), Molur et al. (2002), Naidu & Gururaj (1984), Raghuram et al. (2014), Rao et al. (2004), Sinha (1973), Soisook, Niyomwan et al. (2010), Soisook, Struebig et al. (2015), Srinivasulu & Molur (2008b), Srinivasulu & Srinivasulu (2012),Topàl & Csorba (1992), Volleth et al. (2017), Wordley et al. (2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus beddomei	Rhinolophus		beddomei	K. Andersen	1905	0	Ann. Mag. Nat. Hist.	ser. 7, 16: 253	Bedomme's Horseshoe Bat	<b> sobrinus </b>K. Andersen, 1918.	India, Madras [= Kerala], Wynaad.	S India, Sri Lanka, Thailand.	Not listed.	Least Concern	 trifoliatus species group. Distinct from luctus ; see Top&aacute;l and Csorba (1992), Bates and Harrison (1997), and Hendrichsen et al. (2001a).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus beddomei	23	Beddome's Woolly Horseshoe Bat	Bedomme's Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	beddomei	K. Andersen	1905	0	Rhinolophus_beddomei	Andersen, K. (1905). On bats of the Rhinolophus philppinensis group, with descriptions of five new species. The Annals and magazine of natural history; zoology, botany, and geology, ser. 7, 16, 253.	https://www.biodiversitylibrary.org/page/19215295#page/273/mode/1up	BM 1882.3.3.1		"Wynaad [= Wayanad], Mysore [= Mysuru], S[outh]. India."			beddomei K. Andersen, 1905|sobrinus K. Andersen, 1918	NA	NA	India|Sri Lanka	Asia	Indomalaya	LC	0	0	0	Rhinolophus_beddomei	0	sciname match	Rhinolophus_beddomei	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	40023	Rhinolophus beddomei	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	beddomei	K. Andersen, 1905	This species belongs to trifoliatus species group. Earlier, the taxa beddomei Andersen 1905 and sobrinus Andersen, 1918 were included under Rhinolophus luctus Temminck, 1835 (Ellerman and Morrison-Scott 1951, Corbet and Hill 1992, Koopman 1993). Here Rhinolophus beddomei Andersen, 1905 is treated as a distinct species (TopÃ¡l and Csorba 1992, Bates and Harrison 1997, Simmons 2005, Srinivasulu and Srinivasulu 2012).	20000000	Rhinolophus beddomei	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern because, it is widely but patchily distributed being uncommon in occurrence in its range, and although its habitat is under some threat, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species is primarily a found in dense dry and tropical moist forests. It roosts either as solitary animals or in pairs in caves, old temples, old forts, dilapidated buildings, old unused bunglows, large trees with hollows, wells, old and unused tunnels (B. Srinivasulu and C. Srinivasulu pers. obs. 2016). Seen cohabiting caves with Rhinolopus lepidus and Rhinopoma hardwickii (B. Srinivasulu and C. Srinivasulu pers. obs. 2017). This is a low flyer and feeds on a variety of insects especially beetles and termites (Bates and Harrison 1997).	As this species is a deep forest dweller no direct threats as such to the species or its habitat are recorded. The forests where this species has been recorded are protected and are in good condition.	Though this is a widely distributed species, the overall population is considered to be relatively small because of its great dependence on forest habitats and vulnerability to habitat destruction (Molur et al. 2002). Individuals with a single pup have been observed (Bhargavi Srinivasulu pers. obs. 2017).	Stable	This species is endemic to South Asia. It is presently known from India (Andhra Pradesh, Karnataka, Kerala and Maharashtra) and Sri Lanka (Central, North Central, Southern and Western provinces) in South Asia (Molur et al. 2002, Srinivasulu and Srinivasulu 2012, Srinivasulu et al. 2015). It has been recorded from sea level to an elevation of around 800 m asl. The extent of occurrence is greater than 20,000 kmÂ² and the area of occupancy is greater than 2,000 kmÂ² (Molur et al. 2002).		Terrestrial	Although there are no direct conservation measures in place, the species has been recorded from a number of protected areas in India like Chinnar Wildlife Sanctuary in Kerala, Gundla Brahmeshwaram Wildlife Sanctuary in Andhra Pradesh, and Brahmagiri Wildlife Sanctuary in Karnataka and Parambikulam Tiger Reserve in Kerala (B. Srinivasulu, pers. obs. 2016). Further studies are needed to know the true distribution extent, abundance, reproduction and ecology of this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		beddomei	K. Andersen	1905	0	Ann. Mag. Nat. Hist.	ser. 7, 16: 253	Bedomme's Horseshoe Bat	<b> sobrinus </b>K. Andersen, 1918.	India, Madras [= Kerala], Wynaad.	S India, Sri Lanka, Thailand.	Not listed.	Least Concern	 trifoliatus species group. Distinct from luctus ; see Top&aacute;l and Csorba (1992), Bates and Harrison (1997), and Hendrichsen et al. (2001a).	Rhinolophus beddomei	1004659	23	Beddome's Woolly Horseshoe Bat	Bedomme's Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	beddomei	K. Andersen	1905	0	Rhinolophus_beddomei	Andersen, K. (1905). On bats of the Rhinolophus philppinensis group, with descriptions of five new species. The Annals and magazine of natural history; zoology, botany, and geology, ser. 7, 16, 253.	https://www.biodiversitylibrary.org/page/19215295#page/273/mode/1up	BM 1882.3.3.1		"Wynaad [= Wayanad], Mysore [= Mysuru], S[outh]. India."			beddomei K. Andersen, 1905|sobrinus K. Andersen, 1918	NA	NA				India|Sri Lanka	Asia	Indomalaya	LC	0	0	0	Rhinolophus_beddomei	0	sciname match	Rhinolophus_beddomei	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_beddomei	1004659	23	Beddome's Woolly Horseshoe Bat	Bedomme's Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	beddomei	Andersen	0	Rhinolophus Beddomei	Andersen, K.C. 1905-08-01. On the bats of the _Rhinolophus philippinensis_ group, with descriptions of five new species. Annals and Magazine of Natural History (7)16(92):243-257.	https://www.biodiversitylibrary.org/page/19214880	BMNH:Mamm:1882.3.3.1	holotype	https://data.nhm.ac.uk/object/8742d0fc-cd6d-4a1c-b647-4df97b60a2b4	"Wynaad [= Wayanad], Mysore [= Mysuru], S[outh]. India."			NA	Srinivasulu, A., Srinivasulu, B., Kusuminda, T., Amarasinghe, C., Ukuwela, K. D., Karunarathna, M., ... & Srinivasulu, C. (2023). Taxonomic status of the Lesser Woolly Horseshoe bats (Chiroptera, Rhinolophidae, Rhinolophus beddomei) in peninsular India and Sri Lanka. Zootaxa, 5301(2), 199-218.				India|Sri Lanka	Asia	Indomalaya	LC	0	0	0	Rhinolophus_beddomei	0	sciname match	Rhinolophus_beddomei	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		beddomei	K. Andersen	1905	0	Ann. Mag. Nat. Hist.	ser. 7, 16: 253	Bedomme's Horseshoe Bat	sobrinus K. Andersen, 1918.	India, Madras [= Kerala], Wynaad.	S India, Sri Lanka, Thailand.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/40023/22061859/' target='_blank'>Least Concern</a>	trifoliatus species group. Distinct from luctus; see Top&aacute;l and Csorba (1992), Bates and Harrison (1997), and Hendrichsen et al. (2001a).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus beddomei; Rhinolophus beddomei; Rhinolophus beddomei; Rhinolophus beddomei; Rhinolophus beddomei; Rhinolophus beddomei; beddomei; sobrinus; beddomei; sobrinus; sobrinus; beddomei; sobrinus; Rhinolophede Beddome; Beddome-Wollhaar-Hufeisennase; Herradura de Beddome; Bedomme's Horseshoe Bat; Beddome's Woolly Horseshoe Bat; Bedomme's Horseshoe Bat; Bedomme's Horseshoe Bat; Bedomme's Horseshoe Bat; R. beddomei
