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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1334	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus	Rhinolophus acuminatus		[MSW3] pusillus species group. Subspecific allocations of mainland and Philippine populations are uncertain.; [HMW] Rhinolophus acuminatus Peters, 1871 , “ Gadok .” Java , Indonesia . Tentatively included in the megaphyllus species group but its placement there is uncertain; further genetic work is needed. The most recent genetic study placed the species as sister to R. malayanus , forming a clade sister to much of the megaphyllus , pusillus , philippinensis , and macrotis groups. Five subspecies recognized.; [batnames2022]  pusillus species group. Subspecific allocations of mainland and Philippine populations are uncertain.; [batnames2023]  pusillus species group. Subspecific allocations of mainland and Philippine populations are uncertain.; [batnames2025_1.7] pusillus species group. Subspecific allocations of mainland and Philippine populations are uncertain.						audax, calypso, circe, sumatranus.	sumatranus, circe, calypso, acuminatus, audax	acuminatus, audax, calypso, circe, sumatranus		acuminatus, audax, calypso, circe, sumatranus		acuminatus, audax, calypso, circe, sumatranus		acuminatus, audax, calypso, sumatranus, circe		acuminatus, audax, calypso, circe, sumatranus		acuminatus, audax, calypso, sumatranus, circe	acuminatus, audax, calypso, sumatranus, circe, aequalis, aegualis	acuminatus, audax, calypso, circe, sumatranus		acuminatus W. C. H. Peters, 1871|audax Andersen, 1905|calypso Andersen, 1905|sumatranus Andersen, 1905|circe Andersen, 1906|aequalis G. M. Allen, 1922|aegualis Hooper, 1977 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Acuminate horseshoe bat	Thailand – Java – Lombok, Borneo, Palawan	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhinolophus acuminatus	Indonesia, Java.	Peters	1871	Monatsb. Preuss. Akad. Wiss. Berlin, p. 308.	Distribution: Ran ging from Laos through Cambodia, Thailand, Malaya and Sumatra to Borneo and the south western Philippines and to Java and Lombok, also the west Sumatran islands.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Acuminate horseshoe bat	Thailand – Java – Lombok, Borneo, Palawan	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1871	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1871:308.		Borneo, Sumatra (including Nias and Engano Isis), Java, Lombok, and Bali (Indonesia), Palawan (Philippines), Thailand, Laos, Cambodia.	Indonesia, Java.		PETERS	1871	Sella small, without lateral processes. Median groove of horseshoe simple, no papilla at poste rior end. Connecting process pointed, triangular in profile. Periotic bones not enlarged, no narrow ing of median basioccipital. Size relatively large (forearm length, 46-53 mm).	Distribution: Ran ging from Laos through Cambodia, Thailand, Malaya and Sumatra to Borneo and the south western Philippines and to Java and Lombok, also the west Sumatran islands.	Five subspecies are currently recognized:	R. a. sumatranus (Sumatra, Borneo), R. a. circe (Nias is land), R. a. calypso (Engano island), R. a. acuminatus (Ja va), R. a. audax (Bali, Lombok). Subspecific allocations of mainland and Philippine populations are uncertain.	54	species	R. acuminatus	PETERS	1871	Rhinolophus	genus	Rhinolophus acuminatus				Sella small, without lateral processes. Median groove of horseshoe simple, no papilla at poste rior end. Connecting process pointed, triangular in profile. Periotic bones not enlarged, no narrow ing of median basioccipital. Size relatively large (forearm length, 46-53 mm).	Five subspecies are currently recognized:		25. R. acuminatus PETERS 1871 [pusillus group].	25	_R. a. acuminatus_ Peters, 1871; _R. a. audax_ Andersen, 1905; _R. a. calypso_ Andersen, 1905; _R. a. circe_ Andersen, 1906; _R. a. sumatranus_ Andersen, 1905			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus acuminatus	Rhinolophus		acuminatus	Peters		1871		Monatsb. K. Preuss. Akad. Wiss. Berlin	1871		308		Accuminate Horseshoe Bat	Indonesia, Java, Gadok.	Thailand; Laos; Cambodia; Peninsular Malaysia and Sabah; Borneo; Sumatra (including Nias and Engano Isls); Java, Krakatau, Lombok, and Bali (Indonesia); Palawan, Balabac, Busuanga (Philippines).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	audax K. Andersen, 1905; calypso K. Andersen, 1905; circe K. Andersen, 1906; sumatranus K. Andersen, 1905.	pusillus species group. Subspecific allocations of mainland and Philippine populations are uncertain.	885887A2FFE68A03F84FEFCAF52ADA38	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	310	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFD08A36FF1EF12BF74DD2B6.xml	Rhinolophus acuminatus	Rhinolophidae	Rhinolophus	acuminatus	Peters	1871	Acuminate Horseshoe Bat @en | Rhinolophe acuminò @fr | Spitzsattel-Hufeisennase @de | Herradura acuminado @es	Rhinolophus acuminatus Peters, 1871 , “ Gadok .” Java , Indonesia . Tentatively included in the megaphyllus species group but its placement there is uncertain; further genetic work is needed. The most recent genetic study placed the species as sister to R. malayanus , forming a clade sister to much of the megaphyllus , pusillus , philippinensis , and macrotis groups. Five subspecies recognized.	R . a. acuminatus Peters, 1871 - Krakatau I andJava . R.a. audax K. Andersen, 1905 — Bali and Lombok Is. . a. calypso K. Andersen, 1905 — Enggano I. R. a. circe K. Andersen, 1906 — Nias I. R. a. sumatranus K Andersen, 1905 - W Sumatra and N Borneo. Subspecific identity of mainland populations in SE Asia, as well as the Philippines (Negros, Busuanga, Palawan , and Balabac Is) is uncertain, although mainland populations resemble individuals fromJava or Lombok.	Head—body 48—52 mm , tail 18- 9—31 mm , ear 16-7—21- 4 mm , hindfoot 9-10- 9 mm , forearm 44—53 mm ; weight 9- 6-16 g . There are two color morphs: either grayish brown dorsally with hairs tipped light gray or silver, and grayish brown ventrally; or grayish brown tinged with russet dorsally, and pale reddish brown ventrally. Ears are small to medium-sized. Noseleaf has concave-sided lancet that becomes almost parallel-sided toward tip; connecting process is broad and triangular, with bluntly to sharply pointed tip; sella is variable between subspecies, being nearly parallel-sided to distinctly narrowed centrally, but with rounded tip consistently ; horseshoe does not completely cover muzzle (7-8—10- 2 mm wide ), has visible lateral leaflets, and has wide median emargination. Lower lip has three mental grooves. Baculum (based on specimens of subspecies calypso) has dorso-ventrally higher-than-wide basal cone; ventral notch on proximal margin of base is extremely deep, narrow, and divergent distally; shaft is slightly bent dorsally (although its ventral profile looks nearly straight in lateral view); there is slight constriction medially in dorsal view and also at very tip of bone. Skull is broad, with short rostrum (zygomatic width is significantly broader than mastoid width); anterior median swellings are considerably inflated while posterior swellings are less developed and shorter; sagittal crest is comparatively strong; frontal depression is shallow and very short; supraorbital crests are low with sharp ridges, combining at point in front of center of orbital cavity . P2 is moderately to well developed and generally within tooth row or slightly displaced, which separates C1 from P4; P3 is of medium size and displaced from tooth row, allowing P2 and P4 usually to touch, although not always . Chromosomal complement has 2n = 62 and FN = 60 ( Thailand ).	Lowland dipterocarp forests, primary and secondary tropical forests, bamboo thickets, and some urban areas. Recorded at elevations of 60—250 m in the Philippines and at c. 1000—1700 m in Borneo.	Acuminate Horseshoe Bats are insectivorous. They are typically slow and maneuverable fliers that appear to forage in the subcanopy, several meters from the ground.	A pregnant and a lactating female were captured in June on Palawan.	The Acuminate Horseshoe Bat is nocturnal and roosts during the day, primarily in caves and hollow trees, but it has also been reported roosting in buildings and drainage manifolds- Call shape is FM/CF/FM with a peak F reported at 86-90 kHz (males) and 93—95 kHz (females) in Laos , 90 kHz in Vietnam , 80 kHz in Thailand , and 88—90 kHz in Sabah , Malaysia .	The Acuminate Horseshoe Bat has been reported roosting in small to moderate-sized colonies. A roost in a monastery in Myanmar contained c .100 individuals.	Classified as Least Concern on T / ie IUCN Red List. The Acuminate Horseshoe Bat is apparently common throughout much of its wide range, and does not currently seem to be facing any major threats.	Bates, Thi Mar-Mar et al. (2004) | Csorba et al. (2003) | Esselstyn , Widmann & Heaney (2004) | Francis (2008a) | Harada, Minezawa et al. (1982) | Heaney et al. (1998) | Hood et al. (1988) | Kruskop (2013a) | Matveev (2005) | Rosell-Ambal, McKinnon & Esselstyn (2008) | Schedvin et al. (1994) | Stoffberg et al. (2010) | Zhang Lin et al. (2018)	https://zenodo.org/record/3750030/files/figure.png	62 . Acuminate Horseshoe Bat Rhinolophus acuminatus French: Rhinolophe acuminò / German: Spitzsattel-Hufeisennase / Spanish: Herradura acuminado Taxonomy. Rhinolophus acuminatus Peters, 1871 , “ Gadok .” Java , Indonesia . Tentatively included in the megaphyllus species group but its placement there is uncertain; further genetic work is needed. The most recent genetic study placed the species as sister to R. malayanus , forming a clade sister to much of the megaphyllus , pusillus , philippinensis , and macrotis groups. Five subspecies recognized. Subspecies and Distribution. R . a. acuminatus Peters, 1871 - Krakatau I andJava . R.a. audax K. Andersen, 1905 — Bali and Lombok Is. . a. calypso K. Andersen, 1905 — Enggano I. R. a. circe K. Andersen, 1906 — Nias I. R. a. sumatranus K Andersen, 1905 - W Sumatra and N Borneo. Subspecific identity of mainland populations in SE Asia, as well as the Philippines (Negros, Busuanga, Palawan , and Balabac Is) is uncertain, although mainland populations resemble individuals fromJava or Lombok. Descriptive notes. Head—body 48—52 mm , tail 18- 9—31 mm , ear 16-7—21- 4 mm , hindfoot 9-10- 9 mm , forearm 44—53 mm ; weight 9- 6-16 g . There are two color morphs: either grayish brown dorsally with hairs tipped light gray or silver, and grayish brown ventrally; or grayish brown tinged with russet dorsally, and pale reddish brown ventrally. Ears are small to medium-sized. Noseleaf has concave-sided lancet that becomes almost parallel-sided toward tip; connecting process is broad and triangular, with bluntly to sharply pointed tip; sella is variable between subspecies, being nearly parallel-sided to distinctly narrowed centrally, but with rounded tip consistently ; horseshoe does not completely cover muzzle (7-8—10- 2 mm wide ), has visible lateral leaflets, and has wide median emargination. Lower lip has three mental grooves. Baculum (based on specimens of subspecies calypso) has dorso-ventrally higher-than-wide basal cone; ventral notch on proximal margin of base is extremely deep, narrow, and divergent distally; shaft is slightly bent dorsally (although its ventral profile looks nearly straight in lateral view); there is slight constriction medially in dorsal view and also at very tip of bone. Skull is broad, with short rostrum (zygomatic width is significantly broader than mastoid width); anterior median swellings are considerably inflated while posterior swellings are less developed and shorter; sagittal crest is comparatively strong; frontal depression is shallow and very short; supraorbital crests are low with sharp ridges, combining at point in front of center of orbital cavity . P2 is moderately to well developed and generally within tooth row or slightly displaced, which separates C1 from P4; P3 is of medium size and displaced from tooth row, allowing P2 and P4 usually to touch, although not always . Chromosomal complement has 2n = 62 and FN = 60 ( Thailand ). Habitat. Lowland dipterocarp forests, primary and secondary tropical forests, bamboo thickets, and some urban areas. Recorded at elevations of 60—250 m in the Philippines and at c. 1000—1700 m in Borneo. Food and Feeding. Acuminate Horseshoe Bats are insectivorous. They are typically slow and maneuverable fliers that appear to forage in the subcanopy, several meters from the ground. Breeding. A pregnant and a lactating female were captured in June on Palawan. Males with strong musky odor were collected in October in Vietnam , suggesting this was the mating seas © ®- Activity patterns. The Acuminate Horseshoe Bat is nocturnal and roosts during the day, primarily in caves and hollow trees, but it has also been reported roosting in buildings and drainage manifolds- Call shape is FM/CF/FM with a peak F reported at 86-90 kHz (males) and 93—95 kHz (females) in Laos , 90 kHz in Vietnam , 80 kHz in Thailand , and 88—90 kHz in Sabah , Malaysia . Movements, Home range and Social organization. The Acuminate Horseshoe Bat has been reported roosting in small to moderate-sized colonies. A roost in a monastery in Myanmar contained c .100 individuals. Status and Conservation. Classified as Least Concern on T / ie IUCN Red List. The Acuminate Horseshoe Bat is apparently common throughout much of its wide range, and does not currently seem to be facing any major threats. Bibliography. Bates, Thi Mar-Mar et al. (2004), Csorba et al. (2003), Esselstyn , Widmann & Heaney (2004), Francis (2008a), Harada, Minezawa et al. (1982), Heaney et al. (1998), Hood et al. (1988), Kruskop (2013a), Matveev (2005), Rosell-Ambal, McKinnon & Esselstyn (2008), Schedvin et al. (1994), Stoffberg et al. (2010), Zhang Lin et al. (2018).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus acuminatus	Rhinolophus		acuminatus	Peters	1871	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1876:08:00	Accuminate Horseshoe Bat	<b> audax </b>K. Andersen, 1905;<b> calypso </b>K. Andersen, 1905;<b> circe </b>K. Andersen, 1906; <b> sumatranus </b>K. Andersen, 1905	Indonesia, Java, Gadok.	Thailand; Laos; Cambodia; Peninsular Malaysia and Sabah; Borneo; Sumatra (including Nias and Engano Isls); Java, Krakatau, Lombok, and Bali (Indonesia); Palawan, Balabac, Busuanga (Philippines).	Not listed.	Least Concern	 pusillus species group. Subspecific allocations of mainland and Philippine populations are uncertain.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus acuminatus	23	Acuminate Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	acuminatus	W. Peters	1871	0	Rhinolophus_acuminatus	Peters, W. C. H. (1871). Ãœber die Gattungen und Arten der Hufeisennasen, Rhinolophi. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1871, 308.	https://www.biodiversitylibrary.org/item/112818#page/322/mode/1up	ZMB 2548 [syntypes]		"Gadok." Java, Indonesia.			acuminatus W. Peters, 1871|audax K. Andersen, 1905|calypso K. Andersen, 1905|sumatranus K. Andersen, 1905|circe K. Andersen, 1906	NA	NA	Myanmar|Vietnam|Laos|Cambodia|Thailand|Malaysia|Indonesia|Philippines	Asia	Indomalaya	LC	0	0	0	Rhinolophus_acuminatus	0	sciname match	Rhinolophus_acuminatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19520	Rhinolophus acuminatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	acuminatus	Peters, 1871		20000000	Rhinolophus acuminatus	Least Concern		2019	2018-08-30 00:00:00 UTC	3.1	English	The species is listed as Least Concern because it is widespread and unconsidered for conservation priority throughout the distributional range, which includes protected areas, and lack of major threats although disturbance to the roosting sites occasionally appeared.	Little is known about habitat and ecology of the species. An observation and echolocation recording in southern Vietnam suggest that the species inhabits caves, tunnel and old buildings, forages in different habitats ranging from secondary to primary forests (Esselstyn et al. 2004. Thong et al. , unpublished data). The known roosts of the species have been found from 60-250 m asl in Philippines and Vietnam (Esselstyn et al. 2004, Rosell-Ambal et al. 2008). It occurs in lowland dipterocarp forest in Vietnam, Cambodia and Borneo (Payne et al. 1985, Thong et al. unpublished data).	There are no major threats to this species although disturbance of caves is a localized threat.	The species was found in groups of up to ca.500 individuals in southern Vietnam (Thong et al. unpublished data). It is likely that the species is locally common (Esselstyn et al. 2004, Heaney et al. 1998, Rosell-Ambal et al. 2008).	Unknown	There are records from Myanmar to Indonesia, Malaysia (Borneo Sabah) and the Philippines (Csorba et al. 2003, Rosell-Ambal et al. 2008).	There is no information about either use or trade of this species throughout its distributional range.	Terrestrial	The species occurs in protected areas within its range.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		acuminatus	Peters	1871	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1876:08:00	Accuminate Horseshoe Bat	<b> audax </b>K. Andersen, 1905;<b> calypso </b>K. Andersen, 1905;<b> circe </b>K. Andersen, 1906; <b> sumatranus </b>K. Andersen, 1905	Indonesia, Java, Gadok.	Thailand; Laos; Cambodia; Peninsular Malaysia and Sabah; Borneo; Sumatra (including Nias and Engano Isls); Java, Krakatau, Lombok, and Bali (Indonesia); Palawan, Balabac, Busuanga (Philippines).	Not listed.	Least Concern	 pusillus species group. Subspecific allocations of mainland and Philippine populations are uncertain.	Rhinolophus acuminatus	1004652	23	Acuminate Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	acuminatus	W. Peters	1871	0	Rhinolophus_acuminatus	Peters, W. C. H. (1871). Ãœber die Gattungen und Arten der Hufeisennasen, Rhinolophi. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1871, 308.	https://www.biodiversitylibrary.org/item/112818#page/322/mode/1up	ZMB 2548 [syntypes]		"Gadok." Java, Indonesia.			acuminatus W. Peters, 1871|audax K. Andersen, 1905|calypso K. Andersen, 1905|sumatranus K. Andersen, 1905|circe K. Andersen, 1906	NA	NA				Myanmar|Vietnam|Laos|Cambodia|Thailand|Malaysia|Indonesia|Philippines	Asia	Indomalaya	LC	0	0	0	Rhinolophus_acuminatus	0	sciname match	Rhinolophus_acuminatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_acuminatus	1004652	23	Acuminate Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	acuminatus	W. C. H. Peters	0	Rhinolophus acuminatus	Peters, W.C.H. 1871. Hr. W. Peters las über die Gattungen und Arten der Hufeisennasen, Rhinolophi. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1871:301-332.	https://www.biodiversitylibrary.org/page/36627558	ZMB 2548	syntypes		"Gadok." Java, Indonesia.			NA	NA				Myanmar|Vietnam|Laos|Cambodia|Thailand|Malaysia|Indonesia|Philippines	Asia	Indomalaya	LC	0	0	0	Rhinolophus_acuminatus	0	sciname match	Rhinolophus_acuminatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		acuminatus	Peters	1871	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1876:08:00	Accuminate Horseshoe Bat	audax K. Andersen, 1905; calypso K. Andersen, 1905; circe K. Andersen, 1906; sumatranus K. Andersen, 1905	Indonesia, Java, Gadok.	Thailand; Laos; Cambodia; Peninsular Malaysia and Sabah; Borneo; Sumatra (including Nias and Engano Isls); Java, Krakatau, Lombok, and Bali (Indonesia); Palawan, Balabac, Busuanga (Philippines).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19520/21974227/' target='_blank'>Least Concern</a>	pusillus species group. Subspecific allocations of mainland and Philippine populations are uncertain.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus acuminatus; Rhinolophus acuminatus; Rhinolophus acuminatus; Rhinolophus acuminatus; Rhinolophus acuminatus; Rhinolophus acuminatus; acuminatus; audax; calypso; circe; sumatranus; acuminatus; audax; calypso; circe; sumatranus; audax; calypso; circe; sumatranus; acuminatus; audax; calypso; sumatranus; circe; Acuminate Horseshoe Bat; Rhinolophe acuminò; Spitzsattel-Hufeisennase; Herradura acuminado; Acuminate Horseshoe Bat; Accuminate Horseshoe Bat; Accuminate Horseshoe Bat; R. acuminatus