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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1325	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pteropus ualanus	Pteropus mariannus [synonym of]	N/A	Pteropus mariannus [synonym of]	Pteropus mariannus ualanus	Pteropus ualanus	Pteropus ualanus	Pteropus ualanus	Pteropus ualanus	Pteropus ualanus	Pteropus ualanus	Pteropus ualanus	Pteropus ualanus	Pteropus ualanus	Pteropus ualanus		[MSW3] mariannus species group. Often treated as a subspecies of mariannus, but clearly distinct (Corbet and Hill, 1980; K. Helgen, pers. comm.; Flannery, 1995b). See comments under mariannus.; [HMW] Pteropus ualanus Peters, 1883 , “ Insel Kuschai (Oualan) [= Kosrae Island],” Federated States of Micronesia . Pteropus ualanus is in the griseus species group. It is distinct from mariannus but closely related to tonganus and admiralitatum . Monotypic.; [batnames2022]  griseus species group; see Almeida et al. (2014). Often treated as a subspecies of mariannus, but clearly distinct (Corbet and Hill, 1980; K.Helgen, pers. comm.; Flannery, 1995b). See comments under mariannus. ; [IUCN] Pteropus ualanus was originally described as a distinct species (Peters 1883), an arrangement followed by Anderson (1912). The taxa was subsumed as a subspecies of P. mariannus by Kuroda (1938) and treated as such for a number of decades (e.g., Mickleburgh et al. 1992, Koopman 1993, Wiles 2005). Flannery (1995) was among the first to reclassify it as a separate species, but did so without providing supporting evidence. This treatment was provisionally followed by Simmons (2005). Recent genetic analyses have confirmed that P. ualanus is distinct from P. mariannus and other Pteropus (Almeida et al. 2014).; [batnames2023]  griseus species group; see Almeida et al. (2014). Often treated as a subspecies of mariannus, but clearly distinct (Corbet and Hill, 1980; K.Helgen, pers. comm.; Flannery, 1995b). See comments under mariannus. ; [batnames2025_1.7] griseus species group; see Almeida et al. (2014). Often treated as a subspecies of mariannus, but clearly distinct (Corbet and Hill, 1980; K.Helgen, pers. comm.; Flannery, 1995b). See comments under mariannus.									ualensis			ualanus 	ualanus - ualensis	ualensis, ualanus	Pteropus ualanus was originally described as a distinct species (Peters 1883), an arrangement followed by Anderson (1912). The taxa was subsumed as a subspecies of P. mariannus by Kuroda (1938) and treated as such for a number of decades (e.g., Mickleburgh et al. 1992, Koopman 1993, Wiles 2005). Flannery (1995) was among the first to reclassify it as a separate species, but did so without providing supporting evidence. This treatment was provisionally followed by Simmons (2005). Recent genetic analyses have confirmed that P. ualanus is distinct from P. mariannus and other Pteropus (Almeida et al. 2014).	ualanus 	ualanus - ualensis	ualensis, ualanus 	ualensis, ualanus 	ualanus 	ualanus - ualensis	ualensis Finsch, 1881 [nomen nudum]|ualanus W. C. H. Peters, 1883		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Ualan I, E Caroline Is		N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Pteropus ualanus	Pteropus		ualanus	Peters		1883		Ges. Nat. Fr.	1		1		Kosrae Flying Fox	Ualan (= Kosrae; Micronesia)	Kosrae (Micronesia).	CITES – Appendix II. IUCN/SSC Action Plan (1992) – Endangered: Limited Distribution as P. mariannus ualanus. IUCN 2003 – Not listed.	ualensis Finsch, 1881 [nomen nudum].	mariannus species group. Often treated as a subspecies of mariannus, but clearly distinct (Corbet and Hill, 1980; K. Helgen, pers. comm.; Flannery, 1995b). See comments under mariannus.	03AD87FAFFA0F64F8CB73E59F67DFE30	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	147	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFF9DF673896C3677FF44F6A2.xml	Pteropus ualanus	Pteropodidae	Pteropus	ualanus	Peters	1883	Roussette de Kosrae @fr | Kosrae-Flughund @de | Zorro volador de Kosrae @es | Ualan Fruit Bat @en	Pteropus ualanus Peters, 1883 , “ Insel Kuschai (Oualan) [= Kosrae Island],” Federated States of Micronesia . Pteropus ualanus is in the griseus species group. It is distinct from mariannus but closely related to tonganus and admiralitatum . Monotypic.	Kosrae I, Micronesia . Specimen collected on Pohnpei (Caroline Is) in the 1800s was incorrectly labeled.	Head-body 187-230 mm (tailless), ear 23-24 mm , hindfoot 42-435 mm , forearm 127-133- 5 mm . Eyes of the Kosrae Flying Fox are small, with brown irises. Body is blackish brown, dorsum is slightly speckled with silvery gray hair, and venter is dark brown, also with speckles. Fore neck sometimes has dark rust-brown band. Mantle, sides of neck, and back of head up to ears are rust-brown or sandy-yellow. Crown is dark coffee to rust-brown, gradually darkening along sides of head to blackish deep brown on chin and throat. Uropatagium is reduced to narrow membrane along inside of lower limbs. Index finger has a claw. Skull is typical pteropine and c. 61 mm long. Dentition is typical pteropine, with short cheekteeth mesiodistally.	Forests and mangroves from sea level up to elevations of ¢. 635 m .	No information.	No information.	Kosrae Flying Foxes usually roost in trees during the day and fly to fruit trees at dusk.	Kosrae Flying Foxes mostly roost in colonies, presumably composed of harems and bachelor groups. Historically, colonies reportedly had 100-400 individuals.	CITES Appendix I. Classified as Vulnerable on The IUCN Red List. There was a severe population collapse in 1927 due to a disease that killed thousands of Kosrae Flying Foxes. In the 1980s, Kosrae Flying Foxes might have been exported to Guam as food. Population is currently stable but not very large, so catastrophic events such as typhoons are threats. Increasing loss of habitats (especially deforestation of mangroves) could also quickly become a problem because it does not occur in any protected areas. The Kosrae Flying Fox islisted in IUCN’s action plan for flying foxes that recommends improved management through control of hunting, education, increased legal protection, and forest management.	Almeida et al. (2014) | Andersen (1912b) | Corbet & Hill (1980) | Flannery (1995a) | Mickleburgh et al. (1992) | Nowak (1999) | Peters (1883) | Rainey & Pierson (1992) | Simmons (2005) | Temminck (1841) | Wiles et al. (2008b)		158. Kosrae Flying Fox Pteropus ualanus French: Roussette de Kosrae / German: Kosrae-Flughund / Spanish: Zorro volador de Kosrae Other common names: Ualan Fruit Bat Taxonomy. Pteropus ualanus Peters, 1883 , “ Insel Kuschai (Oualan) [= Kosrae Island],” Federated States of Micronesia . Pteropus ualanus is in the griseus species group. It is distinct from mariannus but closely related to tonganus and admiralitatum . Monotypic. Distribution. Kosrae I, Micronesia . Specimen collected on Pohnpei (Caroline Is) in the 1800s was incorrectly labeled. Descriptive notes. Head-body 187-230 mm (tailless), ear 23-24 mm , hindfoot 42-435 mm , forearm 127-133- 5 mm . Eyes of the Kosrae Flying Fox are small, with brown irises. Body is blackish brown, dorsum is slightly speckled with silvery gray hair, and venter is dark brown, also with speckles. Fore neck sometimes has dark rust-brown band. Mantle, sides of neck, and back of head up to ears are rust-brown or sandy-yellow. Crown is dark coffee to rust-brown, gradually darkening along sides of head to blackish deep brown on chin and throat. Uropatagium is reduced to narrow membrane along inside of lower limbs. Index finger has a claw. Skull is typical pteropine and c. 61 mm long. Dentition is typical pteropine, with short cheekteeth mesiodistally. Habitat. Forests and mangroves from sea level up to elevations of ¢. 635 m . Food and Feeding. No information. Breeding. No information. Activity patterns. Kosrae Flying Foxes usually roost in trees during the day and fly to fruit trees at dusk. Movements, Home range and Social organization. Kosrae Flying Foxes mostly roost in colonies, presumably composed of harems and bachelor groups. Historically, colonies reportedly had 100-400 individuals. Status and Conservation. CITES Appendix I. Classified as Vulnerable on The IUCN Red List. There was a severe population collapse in 1927 due to a disease that killed thousands of Kosrae Flying Foxes. In the 1980s, Kosrae Flying Foxes might have been exported to Guam as food. Population is currently stable but not very large, so catastrophic events such as typhoons are threats. Increasing loss of habitats (especially deforestation of mangroves) could also quickly become a problem because it does not occur in any protected areas. The Kosrae Flying Fox islisted in IUCN’s action plan for flying foxes that recommends improved management through control of hunting, education, increased legal protection, and forest management. Bibliography. Almeida et al. (2014), Andersen (1912b), Corbet & Hill (1980), Flannery (1995a), Mickleburgh et al. (1992), Nowak (1999), Peters (1883), Rainey & Pierson (1992), Simmons (2005), Temminck (1841), Wiles et al. (2008b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Pteropus ualanus	Pteropus		ualanus	Peters	1883	0	Ges. Nat. Fr.	1:01	Kosrae Flying Fox	 ualensis Finsch, 1881 [ nomen nudum ].	Ualan (= Kosrae; Micronesia)	Kosrae (Micronesia).	Appendix I	Endangered	 griseus species group; see Almeida et al. (2014). Often treated as a subspecies of mariannus, but clearly distinct (Corbet and Hill, 1980; K.Helgen, pers. comm.; Flannery, 1995b). See comments under mariannus. 	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pteropus ualanus	23	Kosrae Flying Fox	Ualan Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTEROPODINI	Pteropus	NA	ualanus	W. Peters	1883	0	Pteropus_ualanus	Peters, W. C. H. (1883). Mittheilung über die von Hrn. Dr. Finsch von den Carolinen-Inseln eingesandten Flederhunde. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1883, 1.	https://www.biodiversitylibrary.org/item/35577#page/11/mode/1up	AM B.4456, ZMB 5788, ZMB 5788a-d, ZMB 5792a-b, ZMB 5789, ZMB 5791, ZMB 5798, ZMB 5808, ZMB 5809 [syntypes]		"Insel Kuschai (Oualan) [= Kosrae Island]," Federated States of Micronesia.			ualensis Finsch, 1881 [nomen nudum]|ualanus W. Peters, 1883	NA	NA	Micronesia	Oceania	Australasia/Oceania	EN	0	0	0	Pteropus_ualanus	0	sciname match	Pteropus_ualanus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136531	Pteropus ualanus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Pteropus	ualanus	Peters, 1883	Pteropus ualanus was originally described as a distinct species (Peters 1883), an arrangement followed by Anderson (1912). The taxa was subsumed as a subspecies of P. mariannus by Kuroda (1938) and treated as such for a number of decades (e.g., Mickleburgh et al. 1992, Koopman 1993, Wiles 2005). Flannery (1995) was among the first to reclassify it as a separate species, but did so without providing supporting evidence. This treatment was provisionally followed by Simmons (2005). Recent genetic analyses have confirmed that P. ualanus is distinct from P. mariannus and other Pteropus (Almeida et al. 2014).	20000000	Pteropus ualanus	Endangered	B1ab(iii)	2020	2019-09-18 00:00:00 UTC	3.1	English	Pteropus ualanus is assessed as Endangered under criterion B1ab(iii) because it is restricted to a single location, has a small geographic range (extent of occurrence of 130 km²; ca 110 km² of this is land) making it vulnerable to a single catastrophic event (e.g., infectious disease or typhoon), and has low-levels of ongoing habitat degradation and loss. A survey in 2013–2014 estimated a population size of 1,500–2,250 mature individuals.	The biology of Pteropus ualanus is poorly known. Recent surveys (Hayes and Engbring in press) reveal it is likely a habitat generalist, occurring in mangroves and small islands along the coast, in lowland and montane rainforest to the summits of the highest peaks, and in agroforest. Colony size is highly variable, ranging from two to more than a thousand individuals. Most bats depart the colony shortly before sunset and return shortly after dawn, indicating that most foraging occurs at night, but the species may forage during the day as well because some individuals arrive and depart throughout the day, including the midday hours. It reportedly feeds on palms and breadfruit trees (Kittlitz 1836) and undoubtedly feeds on the fruits and flowers of other tree species. Little information is available on reproduction, but young have been observed in June (Hayes and Engbring in press). Recent surveys by Hayes (Hayes and Engbring in press) revealed that most individuals depart colonies shortly before sunset and return shortly after dawn, but some arrive and depart throughout the day. Most individuals at the Yen Yen Island colony fly a few kilometers over Lelu Harbor to and from distant destinations on the main island.	Pteropus ualanus is vulnerable to four major threats: overharvesting, habitat loss and degradation, severe typhoons, and infectious disease. Even though the residents of Kosrae do not eat or hunt P. ualanus (Sarfert 1919, Mickleburgh et al. 1992, Wiles and Brooke 2009), overharvesting remains a potential threat. Sizable numbers of P. ualanus were apparently killed for export to Guam and the Commonwealth of the Northern Mariana Islands during the 1980s, resulting in a noticeable decline in bat abundance on the island (Rainey 1990, Mickleburgh et al. 1992, Wiles 1992). CITES restrictions enacted in 1989 likely reduced most harvest of P. ualanus (Wiles and Brooke 2009), but poaching and smuggling of about 100 bats at a time at the Yeseng and Utwa colonies continued to occur every few years through 2008 by hunters from Guam and Saipan (Stoney Alokoa, pers. comm.). It is unlikely that this more recent level of harvest has had a significant impact on the population, but this could change if hunting pressure should ever increase. Habitat degradation and loss is likely a relatively minor issue due to Kosrae’s rugged topography and relatively small human population (Anonymous 2018). However, some habitat is being lost due to minor deforestation of rainforest habitat (Anonymous 2018), deforestation of mangroves at a rate of 10% per decade (Hauff et al. 2006), and rapid coastal erosion (Ramsay et al. 2013). Mangrove forests are an important habitat for P. ualanus and may further decline in the future with the progression of sea level rise associated with climate change. Although a variety of invasive weeds and vines are present, none yet appear to threaten forest integrity on the island (Anonymous 2018). Large typhoons have caused mass mortality events in several species of bats (e.g., Flannery 1989, Pierson and Rainey 1992, Wiles and Brooke 2009, O’Shea et al. 2016). Pteropus ualanus survived previous typhoons that damaged the forest on Kosrae in c 1780, 1874, 1891, and 1905 (Spennemann 2004). Typhoons are rare events on Kosrae and the island has not been struck by a severe typhoon since 1905. However, climate change is likely to increase the intensity of typhoons in the tropical western Pacific, thus storm-related impacts to P. ualanus could increase in the future. An infectious disease could again dramatically reduce the population, as it did during 1926–1927 (Coultas 1931), but no subsequent outbreaks have ever been reported. Mass mortality from infectious diseases is extremely rare in island populations of Pteropus , and there are no known cases of all individuals on an island being eliminated (Flannery 1989, Mickleburgh et al. 1992, Pierson and Rainey 1992, O’Shea et al. 2016). The pathogen(s) causing epidemics in other species of Pteropus may be introduced, with domesticated animals or humans implicated as their reservoir (Flannery 1989, Pierson and Rainey 1992, O’Shea et al. 2016).	Pteropus ualanus experienced a major decline in abundance in 1926-1927, when an unidentified, possibly introduced disease decimated the population (Coultas 1931). William Coultas collected vertebrate specimens on Kosrae from 15 January to 11 June 1931 and travelled extensively in the interior of the island and climbed every major peak. He stated that P. ualanus was “Virtually extinct on Kusai [former name of the island]. In 1926 or 27, an epidemic broke out among these mammals which practically wiped out the race. There were only two living specimens remaining on the island that I know of, when I left” (Coultas 1931). In 1983, several colonies of 100-400 bats each were noted on the island and the species was considered locally common (Wiles and Payne 1986, Mickleburgh et al. 1992). Hunting and commercial export of bats to Guam and the Commonwealth of the Northern Mariana Islands during the 1980s was extensive enough to produce a noticeable decline in sightings island wide by the late 1980s, but population size was never assessed during this period (Wiles and Payne 1986, Rainey 1990, Mickleburgh et al. 1992, Wiles 1992). Based on searches for colonies in 2013–2014, the global population is currently estimated at 1,500–2,250 mature individuals (based on an estimated 2,000–3,000 total individuals [Hayes and Engbring in press] and assuming that 75% of all animals are mature). Surveys revealed a minimum of 1,765 individuals inhabiting six colonies on the island, including: 1,038 in mangroves just west of Utwa village, 698 on Yen Yen Island, a small island just north of Lelu village, 20 at the summit of Mt. Matunte, south of Tafunsak village, five on a ridge about 1.2 km south-southeast of the summit of Mt. Oma, at Yeseng, southwest of Malem village, two on a ridge 0.9 km southeast of Mt. Oma, west of Malem village, and two on Srukames, a tiny island just north of Walung village. These counts were conservative because they did not include individuals hidden in vegetation, remaining in the colony after sunset, or in other colonies that may have existed (at least one additional colony of several hundred bats reportedly occurred near Malem, Stoney Alokoa pers. comm.). Thus, the population likely exceeds 2,000 total individuals and may exceed 3,000 total individuals.	Stable	Pteropus ualanus is restricted to the island of Kosrae, where it has an extent of occurrence (EOO) of 131 km², with 110 km² of this being land area of the Federated States of Micronesia. It occurs throughout the island, including the highest peaks (up to 634 m), coastal mangroves, and small satellite islands (Hayes and Engbring in press).	<span class="datalabel1">The residents of Kosrae do not eat P. ualanus ;and the species is not hunted for local consumption (Sarfert 1919, Mickleburgh et al. 1992, Wiles and Brooke 2009). Significant numbers of bats were reportedly harvested by hunters and commercially exported to Guam and the Commonwealth of the Northern Mariana Islands during the 1980s (Rainey 1990, Mickleburgh et al. 1992), although available statistics for the trade revealed only small numbers shipped to these islands (Wiles and Payne 1986, Stinson et al. 1992, Wiles 1992). Transhipment of bats through Pohnpei may account for some of this discrepancy (Wiles and Payne 1986). CITES restrictions enacted in 1989 presumably reduced the threat of continuing commercial trade of the species (Wiles and Brooke 2009), however, poaching and smuggling of about 100 bats at a time by hunters from Guam and Saipan continued to occur every few years through 2008 (Stoney Alokoa pers. comm.).	Terrestrial	Few conservation actions have been developed for P. ualanus. The species was listed on Appendix I of CITES in 1989. The large Utwa colony, which comprises perhaps half of the known population, is located within the Utwe Biosphere Reserve, which was established in 2005 by the United Nations Educational, Scientific and Cultural Organization (UNESCO) Man and the Biosphere Programme (MAB). Regular monitoring of the population, research, and public awareness projects should be implemented.	Oceanian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Pteropus		ualanus	Peters	1883	0	Ges. Nat. Fr.	1:01	Kosrae Flying Fox	 ualensis Finsch, 1881 [ nomen nudum ].	Ualan (= Kosrae; Micronesia)	Kosrae (Micronesia).	Appendix I	Endangered	 griseus species group; see Almeida et al. (2014). Often treated as a subspecies of mariannus, but clearly distinct (Corbet and Hill, 1980; K.Helgen, pers. comm.; Flannery, 1995b). See comments under mariannus. 	Pteropus ualanus	1004506	23	Kosrae Flying Fox	Ualan Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTEROPODINI	Pteropus	NA	ualanus	W. Peters	1883	0	Pteropus_ualanus	Peters, W. C. H. (1883). Mittheilung über die von Hrn. Dr. Finsch von den Carolinen-Inseln eingesandten Flederhunde. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1883, 1.	https://www.biodiversitylibrary.org/item/35577#page/11/mode/1up	AM B.4456, ZMB 5788, ZMB 5788a-d, ZMB 5792a-b, ZMB 5789, ZMB 5791, ZMB 5798, ZMB 5808, ZMB 5809 [syntypes]		"Insel Kuschai (Oualan) [= Kosrae Island]," Federated States of Micronesia.			ualensis Finsch, 1881 [nomen nudum]|ualanus W. Peters, 1883	NA	NA				Micronesia	Oceania	Australasia/Oceania	EN	0	0	0	Pteropus_ualanus	0	sciname match	Pteropus_ualanus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pteropus_ualanus	1004506	23	Kosrae Flying Fox	Ualan Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteropodini	Pteropus	NA	ualanus	W. C. H. Peters	0	Pteropus ualanus	Peters, W.C.H. 1883. Ãœber die von Herrn Dr. Finsch von den Carolinen-Inseln eingesandten Flederhunde. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1883:1-2.	https://www.biodiversitylibrary.org/page/8789293	AM B.4456, RMNH.MAM.21688, ZMB 5788a, ZMB 5788b, ZMB 5788c, ZMB 5788d, ZMB 5792a, ZMB 5792b, ZMB 5788, ZMB 5789, ZMB 5791, ZMB 5798, ZMB 5808, ZMB 5809, ZMB 86715	syntypes	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.21688.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.21688.b	"Insel Kuschai (Oualan) [= Kosrae Island]," Federated States of Micronesia.			NA	NA				Micronesia	Oceania (Continent)	Oceania (Biorealm)	EN	0	0	0	Pteropus_ualanus	0	sciname match	Pteropus_ualanus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Pteropus		ualanus	Peters	1883	0	Ges. Nat. Fr.	1:01	Kosrae Flying Fox	ualensis Finsch, 1881 [nomen nudum].	Ualan (= Kosrae; Micronesia)	Kosrae (Micronesia).	<a href='https://cites.org/eng/app/appendices.php'>Appendix I</a>	<a href='https://www.iucnredlist.org/species/136531/21979719/' target='_blank'>Endangered</a>	griseus species group; see Almeida et al. (2014). Often treated as a subspecies of mariannus, but clearly distinct (Corbet and Hill, 1980; K.Helgen, pers. comm.; Flannery, 1995b). See comments under mariannus.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pteropus ualanus; Pteropus ualanus; Pteropus ualanus; Pteropus ualanus; Pteropus ualanus; Pteropus ualanus; ualensis; ualensis; ualensis; ualanus; Roussette de Kosrae; Kosrae-Flughund; Zorro volador de Kosrae; Ualan Fruit Bat; Kosrae Flying Fox; Ualan Fruit Bat; Kosrae Flying Fox; Kosrae Flying Fox; P. ualanus