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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L133	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus	Chalinolobus picatus		[MSW2] Subgenus Chalinolobus.; [MSW3] Reviewed by Van Deusen and Koopman (1971).; [HMW] Scotophilus picatus Gould, 1852 , Capt. Sturt’s Depot, New South Wales , Australia . This species is monotypic.; [batnames2022] Reviewed by Van Deusen and Koopman (1971).; [batnames2023] Reviewed by Van Deusen and Koopman (1971).; [batnames2025_1.7] Reviewed by Van Deusen and Koopman (1971).														picatus				picatus 	picatus			picatus (J. Gould, 1852)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Little pied bat	S Queensland – New South Wales	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Chalinolobus picatus	Australia, New South Wales, Capt. Sturt's Depot.	Gould	1852	Mamm. Aust., ser. 4, Vol. 3, pl. 43.	Distribution: Confined to dryer interior areas of southeastern Australia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Little pied bat	C, S Queensland – Victoria	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Gould	1852	Mamm. Aust., pt. 4, vol. 3, pl. 43.	Subgenus Chalinolobus.	NW New South Wales, C and S Queensland, and South Australia (Australia).	Australia, New South Wales, Capt. Sturt's Depot.		GOULD	1852	Size relatively small (forearm length, 31-36 mm). Supraorbital swellings of skull poorly developed. Post-orbital constriction well developed. No median crest on braincase. Inner upper incisor bicuspid. Outer upper incisor of medium length. Antero-internal cusp present on last upper premolar. Fur black with lateral and posterior areas of ventral side white.	Distribution: Confined to dryer interior areas of southeastern Australia.	No subspecies.		124	species	C. picatus	GOULD	1852	Chalinolobus	subgenus	Chalinolobus picatus				Size relatively small (forearm length, 31-36 mm). Supraorbital swellings of skull poorly developed. Post-orbital constriction well developed. No median crest on braincase. Inner upper incisor bicuspid. Outer upper incisor of medium length. Antero-internal cusp present on last upper premolar. Fur black with lateral and posterior areas of ventral side white.	No subspecies.		4. C. picatus (GOULD 1852).	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Chalinolobus picatus	Chalinolobus		picatus	Gould	y	1852		Mamm. Aust.	pt. 4	vol. 3	pl. 43		Little Pied Bat	Australia, New South Wales, Capt. Sturt's Depot.	NW New South Wales, C and S Queensland, and South Australia (Australia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		Reviewed by Van Deusen and Koopman (1971).	4C3D87E8FFDE6A62FA8792801F8DB838	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	795	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFDE6A62FA8792801F8DB838.xml	Chalinolobus picatus	Vespertilionidae	Chalinolobus	picatus		1852	Chalinolobe pie @fr | Kleine Lappenfledermaus @de | Calinolobopequeno @es | Pied Bat @en	Scotophilus picatus Gould, 1852 , Capt. Sturt’s Depot, New South Wales , Australia . This species is monotypic.	E Australia , in SE Queensland , NW New South Wales , and E South Australia .	Head-body 39-6-50 mm, tail 29-39 mm, ear 7-5-11-3 mm, forearm 31-36 mm; weight 3-8 g. The Little Pied Bat is the smallest species of wattled bat. Dorsal pelage is glossy black; ventral pelage is glossy black faintly washed with gray, except the margin between ventral pelage and membranes, which is white and makes a V-shape stretching from forearms down to pubic region where it converges, extending onto base of ventral portion of membranes; long dorsal fur extends onto much of uropatagium, grading into a fringe of brownish black hairs. Facial skin, ears, arms and membranes are blackish. Ears are short and broad with ribbing on inner surface, and have small fleshy lobes on lower margin close to lips, and extending under eyes; tragus is pointed inward, rounded, and stubby. A fleshy lobe at corner of mouth is very poorly developed compared with congeners. Uropatagium extends to end oftail. Skull has less pronounced supraorbital swellings than in congeners; there is intermediate contrast between interorbital and intertemporal widths, compared to the Hoary Wattled Bat ( C. nigrogriseus ) and the Chocolate Wattled Bat ( C. morio ); there is no sagittal crest. I* has a posterior cusp; I’ is ¢.40% of height of I? above cingulum; P* is minute but present; P* has an anterointernal cusp; lower molars are myotodont.	Little Pied Bats seem to reach their greatest abundance in mallee, mixed woodlands, mulga ( Acacia , Fabaceae ), and riverine open forest communities around permanent or semi-permanent water bodies, although they have also been reported in dry open forest, open woodlands, chenopod shrublands, Callitris ( Cupressaceae ) forest, and Casuarina pauper ( Casuarinaceae ) woodlands. Recorded at elevations of 40-400 m.	Little Pied Bats are fast fliers and are highly maneuverable, changing direction often and flying close to vegetation while foraging. Foraging bats have been observed gleaning off walls with a distinctive up-and-down sweeping action; they also hunted aerially, with many swoops and dives. One stomach contained only moths.	A pregnant female with two well-developed fetuses was taken in late October. Lactating females were reported in late October, November, and December.	Little Pied Bats are nocturnal, leaving the day roost around dusk and returning around dawn. During the day, they roost in trees, caves, buildings, and abandoned mines. Cave roosts are usually warm and dry, with ambient roost temperatures sometimes over 40°C. Tree roosts have been recorded in hollows in C. pauper , mulga, and large eucalypts; the bats prefer hollows in larger mature trees with dead limbs, or dead and fallen trees that have left a hollowed stump. Hollows are usually large with variably sized entrances. Call shape is a shallow FM/QCF sweep.	Little Pied Bats may roost singly or in colonies of up to 80, but usually in groups of under ten. Roosts are switched nearly every day, but remain in the same general area. Members of a colony roosting in a clump of small, dead mulga near Bourke, New South Wales , commuted 17 km to forage in a dry creek bed, returning in the morning, making a round trip of 34 km per night.	Classified as Near Threatened on The IUCN Red List. The Little Pied Bat is generally uncommon throughout much of its distribution but can be locally common around permanent or semi-permanent water. It may be disappearing from the central portion of its range due to agricultural activities. The species cannot survive well in fragmented habitats and requires large areas of ideal habitat. With this in mind, habitat fragmentation and destruction are the greatest threats, especially since large-scale habitat loss has occurred and continues across the eastern portion of the range in Queensland . In New South Wales , ¢.10% ofits overall distribution has been cleared for cotton over the past 50 years, and the process continues today. In wellconnected small remnants of native habitat, the species currently persists, but populations seem to be decreasing throughout its range. It is also threatened to a lesser extent by roost disturbance, timber harvesting, and the changing offire regimes.	Churchill (2008) | Dominelli (2000) | Ellis & Pennay (2008) | Ford et al. (2008) | Pennay & Freeman (2005) | Pennay et al. (2011) | Richards (1979) | Ryan (1966) | Schulz et al. (1994) | Van Deusen & Koopman (1971) | Young & Ford (2000)	https://zenodo.org/record/6397922/files/figure.png	72. Little Pied Bat Chalinolobus picatus French: Chalinolobe pie / German: Kleine Lappenfledermaus / Spanish: Calinolobo pequeno Other common names: Pied Bat Taxonomy. Scotophilus picatus Gould, 1852 , Capt. Sturt’s Depot, New South Wales , Australia . This species is monotypic. Distribution. E Australia , in SE Queensland , NW New South Wales , and E South Australia . Descriptive notes. Head-body 39-6-50 mm, tail 29-39 mm, ear 7-5-11-3 mm, forearm 31-36 mm; weight 3-8 g. The Little Pied Bat is the smallest species of wattled bat. Dorsal pelage is glossy black; ventral pelage is glossy black faintly washed with gray, except the margin between ventral pelage and membranes, which is white and makes a V-shape stretching from forearms down to pubic region where it converges, extending onto base of ventral portion of membranes; long dorsal fur extends onto much of uropatagium, grading into a fringe of brownish black hairs. Facial skin, ears, arms and membranes are blackish. Ears are short and broad with ribbing on inner surface, and have small fleshy lobes on lower margin close to lips, and extending under eyes; tragus is pointed inward, rounded, and stubby. A fleshy lobe at corner of mouth is very poorly developed compared with congeners. Uropatagium extends to end oftail. Skull has less pronounced supraorbital swellings than in congeners; there is intermediate contrast between interorbital and intertemporal widths, compared to the Hoary Wattled Bat ( C. nigrogriseus ) and the Chocolate Wattled Bat ( C. morio ); there is no sagittal crest. I* has a posterior cusp; I’ is ¢.40% of height of I? above cingulum; P* is minute but present; P* has an anterointernal cusp; lower molars are myotodont. Habitat. Little Pied Bats seem to reach their greatest abundance in mallee, mixed woodlands, mulga ( Acacia , Fabaceae ), and riverine open forest communities around permanent or semi-permanent water bodies, although they have also been reported in dry open forest, open woodlands, chenopod shrublands, Callitris ( Cupressaceae ) forest, and Casuarina pauper ( Casuarinaceae ) woodlands. Recorded at elevations of 40-400 m. Food and Feeding. Little Pied Bats are fast fliers and are highly maneuverable, changing direction often and flying close to vegetation while foraging. Foraging bats have been observed gleaning off walls with a distinctive up-and-down sweeping action; they also hunted aerially, with many swoops and dives. One stomach contained only moths. Breeding. A pregnant female with two well-developed fetuses was taken in late October. Lactating females were reported in late October, November, and December. Activity patterns. Little Pied Bats are nocturnal, leaving the day roost around dusk and returning around dawn. During the day, they roost in trees, caves, buildings, and abandoned mines. Cave roosts are usually warm and dry, with ambient roost temperatures sometimes over 40°C. Tree roosts have been recorded in hollows in C. pauper , mulga, and large eucalypts; the bats prefer hollows in larger mature trees with dead limbs, or dead and fallen trees that have left a hollowed stump. Hollows are usually large with variably sized entrances. Call shape is a shallow FM/QCF sweep. Movements, Home range and Social organization. Little Pied Bats may roost singly or in colonies of up to 80, but usually in groups of under ten. Roosts are switched nearly every day, but remain in the same general area. Members of a colony roosting in a clump of small, dead mulga near Bourke, New South Wales , commuted 17 km to forage in a dry creek bed, returning in the morning, making a round trip of 34 km per night. Status and Conservation. Classified as Near Threatened on The IUCN Red List. The Little Pied Bat is generally uncommon throughout much of its distribution but can be locally common around permanent or semi-permanent water. It may be disappearing from the central portion of its range due to agricultural activities. The species cannot survive well in fragmented habitats and requires large areas of ideal habitat. With this in mind, habitat fragmentation and destruction are the greatest threats, especially since large-scale habitat loss has occurred and continues across the eastern portion of the range in Queensland . In New South Wales , ¢.10% ofits overall distribution has been cleared for cotton over the past 50 years, and the process continues today. In wellconnected small remnants of native habitat, the species currently persists, but populations seem to be decreasing throughout its range. It is also threatened to a lesser extent by roost disturbance, timber harvesting, and the changing offire regimes. Bibliography. Churchill (2008), Dominelli (2000), Ellis & Pennay (2008), Ford et al. (2008), Pennay & Freeman (2005), Pennay et al. (2011), Richards (1979), Ryan (1966), Schulz et al. (1994), Van Deusen & Koopman (1971), Young & Ford (2000).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Chalinolobus picatus	Chalinolobus		picatus	Gould	1852	1	Mamm. Aust.	pt. 4, vol. 3, pl. 43	Little Pied Bat	None.	Australia, New South Wales, Capt. Sturt's Depot.	NW New South Wales, C and S Queensland, and South Australia (Australia).	Not listed.	Near Threatened	Reviewed by Van Deusen and Koopman (1971).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Chalinolobus picatus	23	Little Pied Bat	Pied Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Chalinolobus	NA	picatus	Gould	1852	1	Scotophilus_picatus	Gould, J. (1852). The mammals of Australia, Vol. 3. PI. 43 (with text).	https://www.biodiversitylibrary.org/item/197034#page/181/mode/1up	BM 1853.10.22.33		Capt. Sturt's Depot, New South Wales, Australia.			picatus (Gould, 1852)	NA	NA	Australia	Oceania	Australasia/Oceania	NT	0	0	0	Chalinolobus_picatus	0	sciname match	Chalinolobus_picatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	4422	Chalinolobus picatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Chalinolobus	picatus	(Gould, 1852)		20000000	Chalinolobus picatus	Near Threatened	A2c+4c	2020	2019-07-29 00:00:00 UTC	3.1	English	<p>Chalinolobus picatus is listed as Near Threatened (nearly meets VU A2c+4c) as the total number of adults individuals is inferred to have undergone significant decline of an estimated 25–29% over three generations (16.8 years; GL = 5.6 yrs; Pacifici et al. 2013) due to substantial habitat conversion and range contraction in parts of its range in New South Wales and Queensland.</p>	<p>This species occurs in eucalypt and acacia-dominated open woodlands, semi-arid tall shrublands, dry sclerophyll forests, and Araucaria notophyll vine forests (Duncan et al. 1999). It is found in subterranean roosts, tree hollows, and abandoned buildings. Colonies range from a few individuals to up to ca 50 animals (Freeman and Pennay 2001). Generation length is likely to be around four to six years (Woinarski et al. 2014, Pacifici et al. 2013).</p>	This species is threatened by habitat loss (Woinarski et al . 2014). Other potential threats include: roost disturbance, harvesting of timber in State forest lands, and changing fire regimes (Duncan et al. 1999).	<p>Currently, there is no reliable estimate of the species global population size, but it is assumed to be greater than 20,000 mature individuals given its very large area of occurrence. Roost sites typically contain fewer than 100 individuals (Woinarski et al. 2014) and numerical analyses of reporting rates indicates that the Little Pied Bat accounts for about 7% of bats captured annually within range in New South Wales (Pennay et al. 2011). The population is inferred to be declining due to habitat loss caused by widespread land clearing throughout its range. Since 2006 (roughly 3 generations), over 40,385 km²<sup> </sup>of vegetation was cleared in NSW and Queensland, with much of the most intense clearing occurring in the range of the Little Pied Bat (DPIE 2019, Queensland Government 2019). The extent of habitat loss over the past three generations, whilst substantial in area, is insufficient to meet criteria A given the Little Pied Bat’s very large extent of occurrence ca 977,000 km² (Woinarski et al. 2014). Future rates of decline are expected to be similar with no evidence of decrease in land clearance rates.</p>	Decreasing	<p>This species is found in a broad arc in semi-arid habitats from coastal central Queensland, through the central- and south-west of the state and in drier habitats of inland south-eastern Queensland. The range of the species continues through central and western New South Wales and north-eastern South Australia (Ford et al. 2008, Woinarski et al. 2014). In Queensland, it primarily occurs in the Brigalow Belt Bioregion where there has been substantial vegetation loss (Woinarski et al. 2014).</p>		Terrestrial	It has been recorded from a number of protected areas. There is a need to identify and protect important roosting and foraging sites from habitat conversion and degradation, and from invasive species (goats, cats, foxes). Further studies are needed into the distribution, abundance, natural history, and threats to this species.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Chalinolobus		picatus	Gould	1852	1	Mamm. Aust.	pt. 4, vol. 3, pl. 43	Little Pied Bat	None.	Australia, New South Wales, Capt. Sturt's Depot.	NW New South Wales, C and S Queensland, and South Australia (Australia).	Not listed.	Near Threatened	Reviewed by Van Deusen and Koopman (1971).	Chalinolobus picatus	1005708	23	Little Pied Bat	Pied Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Chalinolobus	NA	picatus	Gould	1852	1	Scotophilus_picatus	Gould, J. (1852). The mammals of Australia, Vol. 3. PI. 43 (with text).	https://www.biodiversitylibrary.org/item/197034#page/181/mode/1up	BM 1853.10.22.33		Capt. Sturt's Depot, New South Wales, Australia.			picatus (Gould, 1852)	NA	NA				Australia	Oceania	Australasia/Oceania	NT	0	0	0	Chalinolobus_picatus	0	sciname match	Chalinolobus_picatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Chalinolobus_picatus	1005708	23	Little Pied Bat	Pied Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Chalinolobus	NA	picatus	J. Gould	1	Scotophilus picatus	Gould, J. 1852. [Part iv]. Pl. I-2, I-41, I-42, I-43, I-45, II-12, II-13, II-18, II-19, II-20, II-21, II-61, II-62, III-14, III-43 in Gould, J. 1845-1863. Mammals of Australia. In three volumes. John Gould, London.	https://www.biodiversitylibrary.org/page/49706122	BMNH:Mamm:1853.10.22.33	holotype	https://data.nhm.ac.uk/object/df725c5e-f60a-4eee-80dd-9c352b4779c6	Capt. Sturt's Depot, New South Wales, Australia.			NA	NA				Australia	Oceania (Continent)	Australasia	NT	0	0	0	Chalinolobus_picatus	0	sciname match	Chalinolobus_picatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Chalinolobus		picatus	Gould	1852	1	Mamm. Aust.	pt. 4, vol. 3, pl. 43	Little Pied Bat	None.	Australia, New South Wales, Capt. Sturt's Depot.	NW New South Wales, C and S Queensland, and South Australia (Australia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/4422/21984147/' target='_blank'>Near Threatened</a>	Reviewed by Van Deusen and Koopman (1971).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Chalinolobus picatus; Chalinolobus picatus; Chalinolobus picatus; Chalinolobus picatus; Chalinolobus picatus; Chalinolobus picatus; picatus; Chalinolobe pie; Kleine Lappenfledermaus; Calinolobopequeno; Pied Bat; Little Pied Bat; Pied Bat; Little Pied Bat; Little Pied Bat; C. picatus