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(1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name 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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1304	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pteropus pohlei	Pteropus pohlei	Pteropus macrotis [synonym of]	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei	Pteropus pohlei		[MSW3] poliocephalus species group. See Flannery (1995b). Includes at least one undescribed species (K. Helgen, pers. comm.).; [HMW] Pteropus pohlei Stein, 1933 , Yapen Island, Province of Papua , Indonesia . Pteropus pohlei is in the griseus species group, and it might contain unnamed taxa (specifically, specimens from Biak-Supiori and Waigeo islands). Monotypic.; [batnames2022]  griseus species group; see Almeida et al. (2014). See Flannery (1995b). Includes at least one undescribed species (K. Helgen, pers. comm.).; [IUCN] Populations from Biak-Supiori and Waigeo that are usually ascribed to this species together represent a related, undescribed species. Pteropus pohlei apparently does not occur on the New Guinea mainland. (K. Helgen pers. comm.).; [batnames2023]  griseus species group; see Almeida et al. (2014). See Flannery (1995b). Includes at least one undescribed species (K. Helgen, pers. comm.).; [batnames2025_1.7] griseusspecies group; see Almeida et al. (2014). See Flannery (1995b). Includes at least one undescribed species (K. Helgen, pers. comm.).														pohlei	Populations from Biak-Supiori and Waigeo that are usually ascribed to this species together represent a related, undescribed species. Pteropus pohlei apparently does not occur on the New Guinea mainland. (K. Helgen pers. comm.).			pohlei 	pohlei 			pohlei G. Stein, 1933		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Geelvink Bay flying fox	W New Guinea	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pteropus pohlei	Indonesia, Irian Jaya, Tjenderawasih Div., Japen Isl.	Stein	1933	Z. Saugetierk., 8:93.	Distribution: Known only from Yapen island off the northern coast of west ern New Guinea.					Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Stein	1933	Z. Saugetierk., 8:93.		Japen Isl (off NW New Guinea).	Indonesia, Irian Jaya, Tjenderawasih Div., Japen Isl.		STEIN	1933	Tibia naked dorsally. Ears relatively short (26-28 mm). Size small for group (forearm length, 126-136 m).	Distribution: Known only from Yapen island off the northern coast of west ern New Guinea.	No subspecies.		26	species	P. pohlei	STEIN	1933	Pteropus	genus	Pteropus pohlei				Tibia naked dorsally. Ears relatively short (26-28 mm). Size small for group (forearm length, 126-136 m).	No subspecies.		52. P. pohlei STEIN 1933 [poliocephalus group].	52	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Pteropus pohlei	Pteropus		pohlei	Stein		1933		Z. Säugetierk.	8		93		Geelvink Bay Flying Fox	Indonesia, Prov. of Papua, Tjenderawasih Div., Yapen Isl.	Yapen, Biak-Supiori, Numfoor, and Rani Isls (off NW New Guinea).	CITES – Appendix II. IUCN/SSC Action Plan (1992) – No Data: Limited Distribution. IUCN 2003 – Vulnerable.		poliocephalus species group. See Flannery (1995b). Includes at least one undescribed species (K. Helgen, pers. comm.).	03AD87FAFF9BF675896833E4F691F217	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	145	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFF93F67D8C6E319DF739F48F.xml	Pteropus pohlei	Pteropodidae	Pteropus	pohlei	Stein	1933	Roussette de Pohle @fr | Pohle-Flughund @de | Zorro volador de Yapen @es | Geelvink Bay Fruit Bat @en | Yapen Flying Fox @en	Pteropus pohlei Stein, 1933 , Yapen Island, Province of Papua , Indonesia . Pteropus pohlei is in the griseus species group, and it might contain unnamed taxa (specifically, specimens from Biak-Supiori and Waigeo islands). Monotypic.	Schouten Is (Rani, Numfor, and Yapen) in Cenderawasih Bay (= Geelvink Bay), off NW New Guinea .	Head-body 176- 210 mm (tailless), ear 25-28 mm , forearm 126-135 mm ; weight 315-360 g (mean 337 g ). Muzzle is narrow, rhinarium is black, and nostrils are short. Eyes are large, with dark brown irises. Ears are relatively long and broad, with narrowly rounded off tips. Head is yellowish, chin and throat are brown to straw yellow, and mantle is golden to orange. Back hairs are silky, adpressed, and brown, with scattered silver hairs. Venter is brownish yellow to fuscous, with scattered yellowish hair tips, and belly often has large dark central patch. Wing membranes are dark brown, and tibia is naked dorsally. Skull has tapering rostrum, large orbits, and very weak sagittal crest; frontal region between orbital processes is concave. All premolars and molars are very weak, high, and widely separated, with rounded edges, and usually tinged in shades of reddish brown.	Primary and disturbed forests, village gardens, and offshore islands.	No information.	Pregnant and lactating Geelvink Bay Flying Foxes have been recorded in September; births might occur in December—January. Breeding might be aseasonal.	The Geelvink Bay Flying Fox is nocturnal.	The Geelvink Bay Flying Fox roosts alone or in small groups.	CITES Appendix II. Classified as Vulnerable on The IUCN Red List. The Geelvink Bay Flying Fox is found in few locations. Hunting is a threat, and extent and quality of habitat continues to decline.	Almeida et al. (2014) | Flannery (1995a) | Mildenstein ( 2016g)		154. Geelvink Bay Flying Fox Pteropus pohlei French: Roussette de Pohle / German: Pohle-Flughund / Spanish: Zorro volador de Yapen Other common names: Geelvink Bay Fruit Bat , Yapen Flying Fox Taxonomy. Pteropus pohlei Stein, 1933 , Yapen Island, Province of Papua , Indonesia . Pteropus pohlei is in the griseus species group, and it might contain unnamed taxa (specifically, specimens from Biak-Supiori and Waigeo islands). Monotypic. Distribution. Schouten Is (Rani, Numfor, and Yapen) in Cenderawasih Bay (= Geelvink Bay), off NW New Guinea . Descriptive notes. Head-body 176- 210 mm (tailless), ear 25-28 mm , forearm 126-135 mm ; weight 315-360 g (mean 337 g ). Muzzle is narrow, rhinarium is black, and nostrils are short. Eyes are large, with dark brown irises. Ears are relatively long and broad, with narrowly rounded off tips. Head is yellowish, chin and throat are brown to straw yellow, and mantle is golden to orange. Back hairs are silky, adpressed, and brown, with scattered silver hairs. Venter is brownish yellow to fuscous, with scattered yellowish hair tips, and belly often has large dark central patch. Wing membranes are dark brown, and tibia is naked dorsally. Skull has tapering rostrum, large orbits, and very weak sagittal crest; frontal region between orbital processes is concave. All premolars and molars are very weak, high, and widely separated, with rounded edges, and usually tinged in shades of reddish brown. Habitat. Primary and disturbed forests, village gardens, and offshore islands. Food and Feeding. No information. Breeding. Pregnant and lactating Geelvink Bay Flying Foxes have been recorded in September; births might occur in December—January. Breeding might be aseasonal. Activity patterns. The Geelvink Bay Flying Fox is nocturnal. Movements, Home range and Social organization. The Geelvink Bay Flying Fox roosts alone or in small groups. Status and Conservation. CITES Appendix II. Classified as Vulnerable on The IUCN Red List. The Geelvink Bay Flying Fox is found in few locations. Hunting is a threat, and extent and quality of habitat continues to decline. Bibliography. Almeida et al. (2014), Flannery (1995a), Mildenstein ( 2016g ).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Pteropus pohlei	Pteropus		pohlei	Stein	1933	0	Z. S&auml;ugetierk.	0.3979	Geelvink Bay Flying Fox	None.	Indonesia, Prov. of Papua, Tjenderawasih Div., Yapen Isl.	Yapen, Biak-Supiori, Numfoor, and Rani Isls (off NW New Guinea).	Appendix II	Vulnerable	 griseus species group; see Almeida et al. (2014). See Flannery (1995b). Includes at least one undescribed species (K. Helgen, pers. comm.).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pteropus pohlei	23	Geelvink Bay Flying Fox	Geelvink Bay Fruit Bat|Yapen Flying Fox	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTEROPODINI	Pteropus	NA	pohlei	Stein	1933	0	Pteropus_pohlei	Stein, G. (1933). Weitere Mitteilungen zur Systematik papuanischer SÃ¤uger. Zeitschrift fÃ¼r SÃ¤ugetierkunde 8, 93.	https://www.biodiversitylibrary.org/item/161846#page/107/mode/1up	ZMB 44688		Yapen Island, Province of Papua, Indonesia.			pohlei Stein, 1933	NA	NA	Indonesia	Oceania	Australasia/Oceania	VU	0	0	0	Pteropus_pohlei	0	sciname match	Pteropus_pohlei	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	18750	Pteropus pohlei	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Pteropus	pohlei	Stein, 1933	Populations from Biak-Supiori and Waigeo that are usually ascribed to this species together represent a related, undescribed species. Pteropus pohlei apparently does not occur on the New Guinea mainland. (K. Helgen pers. comm.).	20000000	Pteropus pohlei	Vulnerable	B1ab(i,v)	2016	2015-04-20 00:00:00 UTC	3.1	English	The species is endemic to West Papua and occurs in only three small island. It is known in fewer than six locations; its extent of occurrence is 11,975 kmÂ², the number of mature individuals is declining due to hunting for food and there is a continuing decline in the extent and quality of its habitat, due to forest conversion. The species is listed as Vulnerable.	The species occurs in both primary tropical forest and disturbed forest. It has been recorded from beaches and offshore islands. Flannery (1995) mentions the capture of an individual in a rural garden. This is a foliage rooster, likely in groups and sometimes on offshore islands (K. Helgen pers. comm.).	This species is threatened by hunting for food, and general forest clearance for timber.	It is common over a small area, and the population trend is decreasing.	Decreasing	This species is endemic to West Papua and restricted to the islands of Numfoor, Rani, and Yapen (all in Indonesia).	The species is hunted for food.	Terrestrial	This species is listed on Appendix II of CITES. It has been recorded from some reserves, however, these are not considered to be well protected. <u> Conservation needs/priorities</u> Studies are needed on the speciesâ€™ population sizes, distribution, and extent of occurrence throughout its range. Monitoring of population sizes and locations over time are also important to establish whether these are stable or experiencing trends of decline. The threats to these bats are poorly understood. Studies are needed on the speciesâ€™ habitat requirements and on the effects of forest loss and degradation on the speciesâ€™ population sizes/distribution. Research is also needed on the amount of hunting and the level of bushmeat trade, and the effects of that hunting on population sizes and persistence. Effective roost site protection efforts are needed to minimize hunting mortality and disturbance to non-target individuals. Similar to most threatened flying foxes, local capacity building for conservation managers and education and awareness within local communities are greatly needed to support conservation efforts.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Pteropus		pohlei	Stein	1933	0	Z. S&auml;ugetierk.	0.397917	Geelvink Bay Flying Fox	None.	Indonesia, Prov. of Papua, Tjenderawasih Div., Yapen Isl.	Yapen, Biak-Supiori, Numfoor, and Rani Isls (off NW New Guinea).	Appendix II	Vulnerable	 griseus species group; see Almeida et al. (2014). See Flannery (1995b). Includes at least one undescribed species (K. Helgen, pers. comm.).	Pteropus pohlei	1004489	23	Geelvink Bay Flying Fox	Geelvink Bay Fruit Bat|Yapen Flying Fox	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTEROPODINI	Pteropus	NA	pohlei	Stein	1933	0	Pteropus_pohlei	Stein, G. (1933). Weitere Mitteilungen zur Systematik papuanischer SÃ¤uger. Zeitschrift fÃ¼r SÃ¤ugetierkunde 8, 93.	https://www.biodiversitylibrary.org/item/161846#page/107/mode/1up	ZMB 44688		Yapen Island, Province of Papua, Indonesia.			pohlei Stein, 1933	NA	NA				Indonesia	Oceania	Australasia/Oceania	VU	0	0	0	Pteropus_pohlei	0	sciname match	Pteropus_pohlei	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pteropus_pohlei	1004489	23	Geelvink Bay Flying Fox	Geelvink Bay Fruit Bat|Yapen Flying Fox	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteropodini	Pteropus	NA	pohlei	G. Stein	0	Pteropus pohlei	Stein, G. 1933-04-24. Weitere Mitteilungen zur Systematik papuanischer SÃ¤uger. Zeitschrift fÃ¼r SÃ¤ugetierkunde 8(2):87-95.	https://www.biodiversitylibrary.org/page/45342796	ZMB 44688	holotype		Yapen Island, Province of Papua, Indonesia.			NA	NA				Indonesia	Oceania (Continent)	Australasia	VU	0	0	0	Pteropus_pohlei	0	sciname match	Pteropus_pohlei	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Pteropus		pohlei	Stein	1933	0	Z. S&auml;ugetierk.	0.397917	Geelvink Bay Flying Fox	None.	Indonesia, Prov. of Papua, Tjenderawasih Div., Yapen Isl.	Yapen, Biak-Supiori, Numfoor, and Rani Isls (off NW New Guinea).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Appendix II</a>	<a href='https://www.iucnredlist.org/species/18750/22085786/' target='_blank'>Vulnerable</a>	griseusspecies group; see Almeida et al. (2014). See Flannery (1995b). Includes at least one undescribed species (K. Helgen, pers. comm.).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pteropus pohlei; Pteropus pohlei; Pteropus pohlei; Pteropus pohlei; Pteropus pohlei; Pteropus pohlei; pohlei; Roussette de Pohle; Pohle-Flughund; Zorro volador de Yapen; Geelvink Bay Fruit Bat; Yapen Flying Fox; Geelvink Bay Flying Fox; Geelvink Bay Fruit Bat; Yapen Flying Fox; Geelvink Bay Flying Fox; Geelvink Bay Flying Fox; P. pohlei
