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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L131	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus	Chalinolobus nigrogriseus		[MSW2] Subgenus Chalinolobus. Includes rogersi; see Van Deusen and Koopman (1971), who revised the species.; [MSW3] Includes rogersi; see Van Deusen and Koopman (1971), who revised the species. Also see Flannery (1995a, b) and Bonaccorso (1998).; [HMW] Scotophilus nigrogriseus Gould, 1852 , vicinity of Moreton Bay, Queensland , Australia . Two subspecies are recognized.; [batnames2022] Includes rogersi ; see Van Deusen and Koopman (1971), who revised the species. Also see Flannery (1995 a , b ) andBonaccorso (1998).; [batnames2023] Includes rogersi ; see Van Deusen and Koopman (1971), who revised the species. Also see Flannery (1995 a , b ) andBonaccorso (1998).; [batnames2025_1.7] Includes rogersi; see Van Deusen and Koopman (1971), who revised the species. Also see Flannery (1995, 1995) and Bonaccorso (1998). May not be present on New Guinea; see Parnaby et al. (2024).		(Frosted bat)		rogersi		rogersi.	rogersi, nigrogriseus	nigrogriseus , rogersi		nigrogriseus, rogersi		nigrogriseus, rogersi		nigrogriseus, rogersi		nigrogriseus, rogersi		nigrogriseus, rogersi 	nigrogriseus, rogersi 	nigrogriseus, rogersi		nigrogriseus (J. Gould, 1856)|rogersi O. Thomas, 1909		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Hoary Bat	SE New Guinea, Fergusson I, N Australia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Chalinolobus nigrogriseus	Australia, Queensland, vic. of Moreton Bay.	Gould	1852	Mamm. Aust., Part 4, Vol. 3, pl. 43.	Distribution: Confined to coastal areas of northern and eastern Australia; also southeastern New Guinea including Fergusson island in the D'Entrecasteaux group.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Hoary bat (Frosted bat)	SE New Guinea, Fergusson I, N Australia	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Gould	1852	Mamm. Aust, pt. 4, vol. 3, pl. 43.	Subgenus Chalinolobus. Includes rogersi; see Van Deusen and Koopman (1971), who revised the species.	N and E Australia; SE New Guinea and adjacent small islands.	Australia, Queensland, vie. of Moreton Bay.		GOULD	1856	Size relatively small (forearm length, 31-39 mm). Supraorbital swellings well developed. Post-orbital constriction well developed. Low median crest on braincase. Inner upper incisor unicuspid. Outer upper incisor relatively short. Antero-internal cusp on posterior upper premolar generally poorly developed. Fur dark gray with varying amounts of whitish frosting.	Distribution: Confined to coastal areas of northern and eastern Australia; also southeastern New Guinea including Fergusson island in the D'Entrecasteaux group.	Two subspecies are currently recognized:	C. n. rogersi (northeastern Western Australia to northwestern Queensland), C. n. nigrogriseus (New Guinea to northeastern New South Wales).	124	species	C. nigrogriseus	GOULD	1856	Chalinolobus	subgenus	Chalinolobus nigrogriseus				Size relatively small (forearm length, 31-39 mm). Supraorbital swellings well developed. Post-orbital constriction well developed. Low median crest on braincase. Inner upper incisor unicuspid. Outer upper incisor relatively short. Antero-internal cusp on posterior upper premolar generally poorly developed. Fur dark gray with varying amounts of whitish frosting.	Two subspecies are currently recognized:		2. C. nigrogriseus (GOULD 1856).	2	_C. n. nigrogriseus_ (Gould, 1856); _C. n. rogersi_ Thomas, 1909			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Chalinolobus nigrogriseus	Chalinolobus		nigrogriseus	Gould	y	1852		Mamm. Aust.	pt. 4	vol. 3	pl. 43		Hoary Wattled Bat	Australia, Queensland, vic. of Moreton Bay.	N and E Australia; SE New Guinea and adjacent small islands.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	rogersi Thomas, 1909.	Includes rogersi; see Van Deusen and Koopman (1971), who revised the species. Also see Flannery (1995a, b) and Bonaccorso (1998).	4C3D87E8FFDD6A62FF56950A168CBA4D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	796	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFDD6A62FF56950A168CBA4D.xml	Chalinolobus nigrogriseus	Vespertilionidae	Chalinolobus	nigrogriseus		1856	Chalinolobe cendré @fr | Schwarzgraue Lappenfledermaus @de | Calinolobocenizo @es | Eastern Wattled Bat @en | Hoary Bat @en | Pied Bat; Blackish-gray Bat (nigrogriseus) @en | Frosted Bat (rogersi) @en	Scotophilus nigrogriseus Gould, 1852 , vicinity of Moreton Bay, Queensland , Australia . Two subspecies are recognized.	C.n.wnigrogriseusGould,1352-SENewGuinea(includingFergussoninD’EntrecasteauxIs)andNEAustralia,fromCapeYorktotheClarenceRiverinNENewSouthWales(includingFraserandStradbrokeIs). C. n. rogersi Thomas, 1909 — N Australia in NE Western Australia , N Northern Territory , and NW Queensland , including Tiwi and Groote Eylandt Is.	Head-body 39-6-58-2 mm,tail 26-43 mm, ear 7-5-13 mm, hindfoot 6-9-8-9 mm, forearm 31-6-39-4 mm; weight 4-2-10 g. Dorsal pelage is dark gray to black, and ventral pelage is grayish brown; fur is frosted with white tips both dorsally and ventrally, giving a hoary appearance, although some individuals lack white frosting, and Western Australian populations have more frosting than Queensland populations; there is some lighter grayish, light brown, or white hair on the ventral margins of the membranes, and light hairs that extend onto the venter of the wing and uropatagium, rarely having a distinctly white Vsshaped band similar to the Little Pied Bat ( C. picatus ). Membranes and ears are black, and some parts of the face and the arms are dark pinkish brown to black. Ears are short and broad with ribbing on inner surface and small fleshy lobes on lower margin close to lips extending under eyes; tragus is pointed inward, rounded, and stubby. There is a fleshy lobe at the corner of the mouth thatis very poorly developed compared to other congeners. Uropatagium extends to end oftail. Skull has well-developed supraorbital swellings; braincase is large and has a low sagittal crest; much contrast between interorbital and intertemporal widths; rostrum is low and swollen. No posterior cusp on I% I? is about one-third height of I* above cingulum; I’ has a notch (unlike the Little Pied Bat); there is a weakly to well-developed anterointernal cusp on P* P? is minute, or even missing in some cases; lower molars are myotodont.	A variety of habitats, including monsoon forests, tall open forests, open woodlands, dry sclerophyll forests ( Eucalyptus camaldulensis , Myrtaceae ), riparian woodlands, littoral rainforest, deciduous vine thickets, coastal scrub, sand dunes, spinifex-covered hills, grasslands, and floodplains. Commonest in lowlands, from sea level up to 300 m .	Hoary Wattled Bats forage close to water and in swampy areas, and have been seen foraging along the forest edge, above the canopy, and close to rivers and creeks in forest, generally avoiding dense vegetation away from water. They catch prey both aerially and by gleaning off the ground and foliage. They are moderately fast fliers, reaching speeds up to 34 km /h; they frequently change direction quickly to catch evasive prey, which they can detect 3-5 m away. They have a varied insectivorous diet, but prefer beetles, ants, and moths; also recorded are spiders, mantises, earwigs, crickets, grasshoppers, cockroaches, cicadas, bugs, flies, and lacewings.	Pregnant females have been recorded in July, August, October, and late November. Twins (rarely single young) are born during October and November and lactating females have been observed from October through January.	Hoary Wattled Bats leave their roosts to forage early in the evening (often before other bats have emerged), and have been observed foraging at twilight, often emerging in large numbers at the same time. They roost in hollows in trees and in roofs of buildings, occasionally also in rock crevices. Call shape is FM/QCF with a characteristic frequency greater than 36 kHz (36-1-40-8 kHz).	The Hoary Wattled Bat roosts in colonies of up to several hundred individuals. In New Guinea , a maternity colony was reported with several hundred females and their young in the roof of a building in November. In this maternity colony, bats clustered in groups of 10-40, some solitary, but no males were present.	Classified as Least Concern on The IUCN Red List. The Hoary Wattled Bat is widespread and common throughout its range, and does not seem to face any major threats. Roost disturbance and loss may be locally significant.	Bonaccorso (1998) | Churchill (2008) | Flannery (1995a, 1995b) | Hutson, Schlitter, Csorba, Thomson & McKenzie (2008) | Kutt et al. (2008) | McKenzie, Fontanini et al. (1995) | Milledge et al. (1992) | Milne (2002) | Ryan (1966) | Van Deusen & Koopman (1971) | Vestjens & Hall (1977)	https://zenodo.org/record/6397924/files/figure.png	73. Hoary Wattled Bat Chalinolobus nigrogriseus French: Chalinolobe cendré / German: Schwarzgraue Lappenfledermaus / Spanish: Calinolobo cenizo Other common names: Eastern Wattled Bat , Hoary Bat , Pied Bat; Blackish-gray Bat (nigrogriseus) , Frosted Bat (rogersi) Taxonomy. Scotophilus nigrogriseus Gould, 1852 , vicinity of Moreton Bay, Queensland , Australia . Two subspecies are recognized. Subspecies and Distribution. C.n.wnigrogriseusGould,1352-SENewGuinea(includingFergussoninD’EntrecasteauxIs)andNEAustralia,fromCapeYorktotheClarenceRiverinNENewSouthWales(includingFraserandStradbrokeIs). C. n. rogersi Thomas, 1909 — N Australia in NE Western Australia , N Northern Territory , and NW Queensland , including Tiwi and Groote Eylandt Is. Descriptive notes. Head-body 39-6-58-2 mm,tail 26-43 mm, ear 7-5-13 mm, hindfoot 6-9-8-9 mm, forearm 31-6-39-4 mm; weight 4-2-10 g. Dorsal pelage is dark gray to black, and ventral pelage is grayish brown; fur is frosted with white tips both dorsally and ventrally, giving a hoary appearance, although some individuals lack white frosting, and Western Australian populations have more frosting than Queensland populations; there is some lighter grayish, light brown, or white hair on the ventral margins of the membranes, and light hairs that extend onto the venter of the wing and uropatagium, rarely having a distinctly white Vsshaped band similar to the Little Pied Bat ( C. picatus ). Membranes and ears are black, and some parts of the face and the arms are dark pinkish brown to black. Ears are short and broad with ribbing on inner surface and small fleshy lobes on lower margin close to lips extending under eyes; tragus is pointed inward, rounded, and stubby. There is a fleshy lobe at the corner of the mouth thatis very poorly developed compared to other congeners. Uropatagium extends to end oftail. Skull has well-developed supraorbital swellings; braincase is large and has a low sagittal crest; much contrast between interorbital and intertemporal widths; rostrum is low and swollen. No posterior cusp on I% I? is about one-third height of I* above cingulum; I’ has a notch (unlike the Little Pied Bat); there is a weakly to well-developed anterointernal cusp on P* P? is minute, or even missing in some cases; lower molars are myotodont. Habitat. A variety of habitats, including monsoon forests, tall open forests, open woodlands, dry sclerophyll forests ( Eucalyptus camaldulensis , Myrtaceae ), riparian woodlands, littoral rainforest, deciduous vine thickets, coastal scrub, sand dunes, spinifex-covered hills, grasslands, and floodplains. Commonest in lowlands, from sea level up to 300 m . Food and Feeding. Hoary Wattled Bats forage close to water and in swampy areas, and have been seen foraging along the forest edge, above the canopy, and close to rivers and creeks in forest, generally avoiding dense vegetation away from water. They catch prey both aerially and by gleaning off the ground and foliage. They are moderately fast fliers, reaching speeds up to 34 km /h; they frequently change direction quickly to catch evasive prey, which they can detect 3-5 m away. They have a varied insectivorous diet, but prefer beetles, ants, and moths; also recorded are spiders, mantises, earwigs, crickets, grasshoppers, cockroaches, cicadas, bugs, flies, and lacewings. Breeding. Pregnant females have been recorded in July, August, October, and late November. Twins (rarely single young) are born during October and November and lactating females have been observed from October through January. Activity patterns. Hoary Wattled Bats leave their roosts to forage early in the evening (often before other bats have emerged), and have been observed foraging at twilight, often emerging in large numbers at the same time. They roost in hollows in trees and in roofs of buildings, occasionally also in rock crevices. Call shape is FM/QCF with a characteristic frequency greater than 36 kHz (36-1-40-8 kHz). Movements, Home range and Social organization. The Hoary Wattled Bat roosts in colonies of up to several hundred individuals. In New Guinea , a maternity colony was reported with several hundred females and their young in the roof of a building in November. In this maternity colony, bats clustered in groups of 10-40, some solitary, but no males were present. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Hoary Wattled Bat is widespread and common throughout its range, and does not seem to face any major threats. Roost disturbance and loss may be locally significant. Bibliography. Bonaccorso (1998), Churchill (2008), Flannery (1995a, 1995b), Hutson, Schlitter, Csorba, Thomson & McKenzie (2008), Kutt et al. (2008), McKenzie, Fontanini et al. (1995), Milledge et al. (1992), Milne (2002), Ryan (1966), Van Deusen & Koopman (1971), Vestjens & Hall (1977).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Chalinolobus nigrogriseus	Chalinolobus		nigrogriseus	Gould	1852	1	Mamm. Aust.	pt. 4, vol. 3, pl. 43	Hoary Wattled Bat	<b> rogersi </b>Thomas, 1909.	Australia, Queensland, vic. of Moreton Bay.	N and E Australia; SE New Guinea and adjacent small islands.	Not listed.	Least Concern	Includes rogersi ; see Van Deusen and Koopman (1971), who revised the species. Also see Flannery (1995 a , b ) andBonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Chalinolobus nigrogriseus	23	Hoary Wattled Bat	Eastern Wattled Bat|Hoary Bat|Pied Bat|Blackish-gray Bat|Frosted Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Chalinolobus	NA	nigrogriseus	Gould	1852	1						vicinity of Moreton Bay, Queensland, Australia.			nigrogriseus (Gould, 1856)|rogersi O. Thomas, 1909	NA	NA	Papua New Guinea|Australia	Oceania	Australasia/Oceania	LC	0	0	0	Chalinolobus_nigrogriseus	0	sciname match	Chalinolobus_nigrogriseus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	4421	Chalinolobus nigrogriseus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Chalinolobus	nigrogriseus	(Gould, 1856)		20000000	Chalinolobus nigrogriseus	Least Concern		2020	2019-08-23 00:00:00 UTC	3.1	English	This species is listed as Least Concern because it has a wide distribution and populations are thought to be secure.	This species is present in lowland habitats where it has been recorded from wet sclerophyll forest, open woodlands, scrub, and over sand dunes. It roosts in tree holes, rock crevices, and in buildings. It is an aerial insectivore that forages over wetlands, including waterholes, marshes, creeks, and sewage ponds, and within forested areas. Females give birth to one or two young (Flannery 1995a, Bonaccorso 1998).	In general, there appear to be no major threats to this species. However, it may be locally threatened in parts of its range by disturbance of roosting sites or loss of tree hollows. Urban expansion, mining and agriculture are emerging threats as urban development is occurring in prime habitat zones within SE Queensland and NE New South Wales.	This species is common and widespread within its Australian range (Kutt et al. 2008). Bonaccorso (1998) suggests that it is not commonly encountered in Papua New Guinea, even though it occurs in association with humans.	Stable	This species has been recorded on the island of New Guinea (present in Papua New Guinea only), on Fergusson Island in the D'Entrecasteaux Islands (Papua New Guinea), and from much of northern Australia (Flannery 1995a,b; Bonaccorso 1998; Kutt et al. 2008). It ranges from sea Level to 300 m asl.		Terrestrial	This species occurs in a number of protected areas. Bonaccorso (1998) suggests that local educational campaigns may assist the conservation of this species in New Guinea.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Chalinolobus		nigrogriseus	Gould	1852	1	Mamm. Aust.	pt. 4, vol. 3, pl. 43	Hoary Wattled Bat	<b> rogersi </b>Thomas, 1909.	Australia, Queensland, vic. of Moreton Bay.	N and E Australia; SE New Guinea and adjacent small islands.	Not listed.	Least Concern	Includes rogersi ; see Van Deusen and Koopman (1971), who revised the species. Also see Flannery (1995 a , b ) andBonaccorso (1998).	Chalinolobus nigrogriseus	1005707	23	Hoary Wattled Bat	Eastern Wattled Bat|Hoary Bat|Pied Bat|Blackish-gray Bat|Frosted Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Chalinolobus	NA	nigrogriseus	Gould	1852	1						vicinity of Moreton Bay, Queensland, Australia.			nigrogriseus (Gould, 1856)|rogersi O. Thomas, 1909	NA	NA				Papua New Guinea|Australia	Oceania	Australasia/Oceania	LC	0	0	0	Chalinolobus_nigrogriseus	0	sciname match	Chalinolobus_nigrogriseus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Chalinolobus_nigrogriseus	1005707	23	Hoary Wattled Bat	Eastern Wattled Bat|Hoary Bat|Pied Bat|Blackish-gray Bat|Frosted Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Chalinolobus	NA	nigrogriseus	J. Gould	1	Scotophilus nigrogriseus	Gould, J. 1856. [Part viii]. Pl. I-12, I-16, I-17, I-18, I-19, I-33, II-16, II-17, II-28, II-48, II-49, II-50, III-22, III-33, III-44 in Gould, J. 1845-1863. Mammals of Australia. In three volumes. John Gould, London.	https://www.biodiversitylibrary.org/page/49706126	BMNH:Mamm:1856.10.28.3	holotype	https://data.nhm.ac.uk/object/dae75288-03d5-40bf-b8da-bd73b62ab6be	vicinity of Moreton Bay, Queensland, Australia.			NA	NA				Papua New Guinea|Australia	Oceania (Continent)	Australasia	LC	0	0	0	Chalinolobus_nigrogriseus	0	sciname match	Chalinolobus_nigrogriseus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Chalinolobus		nigrogriseus	Gould	1852	1	Mamm. Aust.	pt. 4, vol. 3, pl. 43	Hoary Wattled Bat	rogersi Thomas, 1909.	Australia, Queensland, vic. of Moreton Bay	N and E Australia; SE New Guinea and adjacent small islands	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/4421/21984276/' target='_blank'>Least Concern</a>	Includes rogersi; see Van Deusen and Koopman (1971), who revised the species. Also see Flannery (1995, 1995) and Bonaccorso (1998). May not be present on New Guinea; see Parnaby et al. (2024).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Chalinolobus nigrogriseus; Chalinolobus nigrogriseus; Chalinolobus nigrogriseus; Chalinolobus nigrogriseus; Chalinolobus nigrogriseus; Chalinolobus nigrogriseus; nigrogriseus ; rogersi; nigrogriseus; rogersi; rogersi; nigrogriseus; rogersi; Chalinolobe cendré; Schwarzgraue Lappenfledermaus; Calinolobocenizo; Eastern Wattled Bat; Hoary Bat; Pied Bat; Blackish-gray Bat (nigrogriseus); Frosted Bat (rogersi); Hoary Wattled Bat; Eastern Wattled Bat; Hoary Bat; Pied Bat; Blackish-gray Bat; Frosted Bat; Hoary Wattled Bat; Hoary Wattled Bat; C. nigrogriseus
