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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1281	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei	Pteropus lylei		[MSW3] vampyrus species group.; [HMW] Pteropus lylei K. Andersen, 1908 , “ Bangkok , Siam [= Thailand ].” Pteropus lylei 1s in the vampyrus species group. Monotypic.; [batnames2022]  vampyrus species group; see Almeida et al. (2014).; [batnames2023]  vampyrus species group; see Almeida et al. (2014).; [batnames2025_1.7] vampyrusspecies group; see Almeida et al. (2014).										lombocensis, heudei, salottii				lylei				lylei 	lylei 			lylei Andersen, 1908		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Thailand, Indochina	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pteropus lylei	Thailand, Bangkok.	K. Andersen	1908	Ann. Mag. Nat. Hist., ser. 8, 2:367.	Dis tribution: Known only from Thailand, Cambodia, and Vietnam.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Lyle's flying fox	Thailand, Indochina	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	K. Andersen	1908	Ann. Mag. Nat. Hist., ser. 8, 2:367.		Thailand, Vietnam.	Thailand, Bangkok.		ANDERSEN	1908	Posterior basal ledges of premolars practically ob literated. Tibia naked dorsally. Premolars and molars not extremely narrow (breadth of last up per premolar at least one fourth width of palate between them). Last two upper premolars clearly elongate. Inner edges of plagiopatagia well sep arated. Ears long and sharply pointed. Size small for group (forearm length, 145-160 mm).	Dis tribution: Known only from Thailand, Cambodia, and Vietnam.	No subspecies.		25	species	P. lylei	ANDERSEN	1908	Pteropus	genus	Pteropus lylei				Posterior basal ledges of premolars practically ob literated. Tibia naked dorsally. Premolars and molars not extremely narrow (breadth of last up per premolar at least one fourth width of palate between them). Last two upper premolars clearly elongate. Inner edges of plagiopatagia well sep arated. Ears long and sharply pointed. Size small for group (forearm length, 145-160 mm).	No subspecies.		44. P. lylei ANDERSEN 1908 [vampyrus group],	44	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Pteropus lylei	Pteropus		lylei	K. Andersen		1908		Ann. Mag. Nat. Hist., ser. 8	2		367		Lyle's Flying Fox	Thailand, Bangkok.	Thailand, Vietnam, Cambodia.	CITES – Appendix II. IUCN/SSC Action Plan (1992) – Not Threatened. IUCN 2003 – Lower Risk (lc).		vampyrus species group.		Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0			157	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFA7F649896B3279F689FAC0.xml	Pteropus lylei	Pteropodidae	Pteropus	lylei	K. Andersen	1908	Roussette de Lyle @fr | Lyle-Flughund @de | Zorro volador de Lyle @es	Pteropus lylei K. Andersen, 1908 , “ Bangkok , Siam [= Thailand ].” Pteropus lylei 1s in the vampyrus species group. Monotypic.	S Thailand , S Cambodia , and S Vietnam around Gulf of Thailand , entering inland C Cambodia . Single specimen recorded in Yunnan ( China ) considered alien.	Head-body 200 mm (tailless), ear 34-36 mm , hindfoot 50-54 mm , forearm 145-160 mm ; weight 390-480 g . Lyle’s Flying Fox is similar to the Indian Flying Fox ( P. medius ) but smaller. Muzzle is long, tapering, dark brown, and sparsely haired; nostrils are shortly tubular. Eyes are large, with brown irises. Ears are long and pointed. Head pelage is dark brown around eyes; cap and sides of head are reddish to yellowish brown. Nape and mantle are distinctly yellowish to buff or ocherous buff, orange in some individuals, with longer and hirsute hairs, extending to sides of neck; mantle ends abruptly on back, which is dark brown to rump, with closely adpressed, distinctly grizzled hairs. Throat is dark; collar is reddish brown to russet, separate from seal-brown, grizzled belly, often washed in reddish tinge; and in some individuals, belly is russet or orange-brown down to black genitals. Wing membranes are dark brown from slightly above medial plane, index claw is present, and all claws are black. Forearm and tibia are naked dorsally. Uropatagium is narrow at center; calcaris short. Skull is typical pteropine, with strong basicranial deflection; rostrum is long and tapering; orbit is large; zygoma is thin, weak, and gently arched; braincase is domed; postorbital foramina are large; postorbital constriction is poorly marked; temporal lines do notjoin in raised sagittal crest; nuchal crest is obvious but low; palate is flat and long; tooth rows are slightly divergent posteriorly; and post-dental palate is relatively short. Mandible is rather delicate; symphysis is long and sloping. Dentition is typical pteropine but generally weak, with rather rounded cusps; M,is small and peglike. Chromosomal complement has 2n = 40 and FN = 76.	Mangrove forests, freshwater and peat swamp forests, lowland deciduous dry forests, mountain rainforests, and rainforests. Roosting habitat of Lyle’s Flying Fox can be in close proximity to human settlements, with marked preference (53% of roosts) for trees growing in Buddhist temples, and is usually near watercourses.	Liyle’s Flying Fox is primarily frugivorous. In Thailand ,it eats fruits of 34 plant genera, predominantly figs ( Ficus , Moraceae ) and cultivated plants in orchards. Plants visited for flowers and leaves include Acacia (Fabaceae) , Eucalyptus ( Myrtaceae ), and Markhamia ( Bignoniaceae ). Lyle’s Flying Foxes ingest ¢.50% their body weight/day. It is an important seed disperser and a central-place forager that returns to the same roosts after foraging.	Lyle’s Flying Fox is seasonally monoestrous. Mating and courtship occur in morning and late afternoon at roosts. Litter size is one, occasionally two. Young are more common in colonies in April-May (hot inter-monsoonal period). Young remains attached to, and flies with, its mother for most of the lactation period.	Lyle’s Flying Foxes are crepuscular and nocturnal. They leave day roosts in the evening to forage and return in early morning (03:00-05:00 h). Lunar phase had no influence on timing of colony-wide emergence; during breeding season, females were the last to emerge. During daytime, sleeping, grooming, and wing flapping are the most frequent behaviors of roosting bats. Wing spreading, movement, and courtship are more frequent in males than females; the opposite is true for grooming and antagonistic behaviors.	Lyle’s Flying Fox is highly colonial and roosts in tall trees; in Thailand , roosts contain 100-1000 individuals of both sexes, with one colony of more than 11,000 individuals. It is highly mobile and can travel up to 24 km /night from day roosts to foraging areas; cumulative distances traveled per night are up to 50 km . Fidelity for roosting and feeding sites is high. Lyle’s Flying Fox spent 5—-8% of time commuting to foraging sites, 58-78% foraging, and 12-37% roosting depending on season (more foraging in November, more roosting in April-May); 62-65% of roosting time was spent resting.	CITES Appendix II. Classified as Vulnerable on The IUCN Red List. Lyle’s Flying Fox is perceived to have decline of more than 30% over the past 15 years due to increase hunting and trade, decline in habitat quality and extent due to deforestation, and loss of tall roosting trees. It occurs in protected areas. It is tolerant of human presence; roosting sites are in close proximity to settlements, particularly Buddhist temples where it is protected by monks (58% of 30 known roosts in Thailand ). The Wildlife Preservation and Protection Act protects it in Thailand , and overall population is estimated to be 75,000 individuals.	Almeida et al. (2014) | Andersen (1912b) | Boonneung (1977) | Bumrungsri, Suyanto & Francis (2008) | Chaiyes et al. (2017) | Hengjan (2011) | Hondo et al. (2010) | Hood et al. (1988) | Smith, A.T. & Xie Yan (2008) | Smith, J.D.L. & Gregory (2009) | Weber et al. (2015) | Zhang Jinshuo et al. (2010)		179. Lyle’s Flying Fox Pteropus lylei French: Roussette de Lyle / German: Lyle-Flughund / Spanish: Zorro volador de Lyle Taxonomy. Pteropus lylei K. Andersen, 1908 , “ Bangkok , Siam [= Thailand ].” Pteropus lylei 1s in the vampyrus species group. Monotypic. Distribution. S Thailand , S Cambodia , and S Vietnam around Gulf of Thailand , entering inland C Cambodia . Single specimen recorded in Yunnan ( China ) considered alien. Descriptive notes. Head-body 200 mm (tailless), ear 34-36 mm , hindfoot 50-54 mm , forearm 145-160 mm ; weight 390-480 g . Lyle’s Flying Fox is similar to the Indian Flying Fox ( P. medius ) but smaller. Muzzle is long, tapering, dark brown, and sparsely haired; nostrils are shortly tubular. Eyes are large, with brown irises. Ears are long and pointed. Head pelage is dark brown around eyes; cap and sides of head are reddish to yellowish brown. Nape and mantle are distinctly yellowish to buff or ocherous buff, orange in some individuals, with longer and hirsute hairs, extending to sides of neck; mantle ends abruptly on back, which is dark brown to rump, with closely adpressed, distinctly grizzled hairs. Throat is dark; collar is reddish brown to russet, separate from seal-brown, grizzled belly, often washed in reddish tinge; and in some individuals, belly is russet or orange-brown down to black genitals. Wing membranes are dark brown from slightly above medial plane, index claw is present, and all claws are black. Forearm and tibia are naked dorsally. Uropatagium is narrow at center; calcaris short. Skull is typical pteropine, with strong basicranial deflection; rostrum is long and tapering; orbit is large; zygoma is thin, weak, and gently arched; braincase is domed; postorbital foramina are large; postorbital constriction is poorly marked; temporal lines do notjoin in raised sagittal crest; nuchal crest is obvious but low; palate is flat and long; tooth rows are slightly divergent posteriorly; and post-dental palate is relatively short. Mandible is rather delicate; symphysis is long and sloping. Dentition is typical pteropine but generally weak, with rather rounded cusps; M,is small and peglike. Chromosomal complement has 2n = 40 and FN = 76. Habitat. Mangrove forests, freshwater and peat swamp forests, lowland deciduous dry forests, mountain rainforests, and rainforests. Roosting habitat of Lyle’s Flying Fox can be in close proximity to human settlements, with marked preference (53% of roosts) for trees growing in Buddhist temples, and is usually near watercourses. Food and Feeding. Liyle’s Flying Fox is primarily frugivorous. In Thailand ,it eats fruits of 34 plant genera, predominantly figs ( Ficus , Moraceae ) and cultivated plants in orchards. Plants visited for flowers and leaves include Acacia (Fabaceae) , Eucalyptus ( Myrtaceae ), and Markhamia ( Bignoniaceae ). Lyle’s Flying Foxes ingest ¢.50% their body weight/day. It is an important seed disperser and a central-place forager that returns to the same roosts after foraging. Breeding. Lyle’s Flying Fox is seasonally monoestrous. Mating and courtship occur in morning and late afternoon at roosts. Litter size is one, occasionally two. Young are more common in colonies in April-May (hot inter-monsoonal period). Young remains attached to, and flies with, its mother for most of the lactation period. Activity patterns. Lyle’s Flying Foxes are crepuscular and nocturnal. They leave day roosts in the evening to forage and return in early morning (03:00-05:00 h). Lunar phase had no influence on timing of colony-wide emergence; during breeding season, females were the last to emerge. During daytime, sleeping, grooming, and wing flapping are the most frequent behaviors of roosting bats. Wing spreading, movement, and courtship are more frequent in males than females; the opposite is true for grooming and antagonistic behaviors. Movements, Home range and Social organization. Lyle’s Flying Fox is highly colonial and roosts in tall trees; in Thailand , roosts contain 100-1000 individuals of both sexes, with one colony of more than 11,000 individuals. It is highly mobile and can travel up to 24 km /night from day roosts to foraging areas; cumulative distances traveled per night are up to 50 km . Fidelity for roosting and feeding sites is high. Lyle’s Flying Fox spent 5—-8% of time commuting to foraging sites, 58-78% foraging, and 12-37% roosting depending on season (more foraging in November, more roosting in April-May); 62-65% of roosting time was spent resting. Status and Conservation. CITES Appendix II. Classified as Vulnerable on The IUCN Red List. Lyle’s Flying Fox is perceived to have decline of more than 30% over the past 15 years due to increase hunting and trade, decline in habitat quality and extent due to deforestation, and loss of tall roosting trees. It occurs in protected areas. It is tolerant of human presence; roosting sites are in close proximity to settlements, particularly Buddhist temples where it is protected by monks (58% of 30 known roosts in Thailand ). The Wildlife Preservation and Protection Act protects it in Thailand , and overall population is estimated to be 75,000 individuals. Bibliography. Almeida et al. (2014), Andersen (1912b), Boonneung (1977), Bumrungsri, Suyanto & Francis (2008), Chaiyes et al. (2017), Hengjan (2011), Hondo et al. (2010), Hood et al. (1988), Smith, A.T. & Xie Yan (2008), Smith, J.D.L. & Gregory (2009), Weber et al. (2015), Zhang Jinshuo et al. (2010).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Pteropus lylei	Pteropus		lylei	K. Andersen	1908	0	Ann. Mag. Nat. Hist.	ser. 8, 2: 367	Lyle's Flying Fox	None.	Thailand, Bangkok.	Thailand, Vietnam, Cambodia.	Appendix II	Vulnerable	 vampyrus species group; see Almeida et al. (2014).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pteropus lylei	23	Lyle's Flying Fox		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTEROPODINI	Pteropus	NA	lylei	K. Andersen	1908	0						"Bangkok, Siam [= Thailand]."			lylei K. Andersen, 1908	NA	NA	Thailand|Cambodia|Vietnam	Asia	Indomalaya	VU	0	0	0	Pteropus_lylei	0	sciname match	Pteropus_lylei	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	18734	Pteropus lylei	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Pteropus	lylei	K. Andersen, 1908		20000000	Pteropus lylei	Vulnerable	A4cd	2021	2020-11-27 00:00:00 UTC	3.1	English	Pteropus lylei is assessed as Vulnerable under criterion A4cd as it is estimated that a decline of over 30-35% has occurred a 15-year period (three generations, GL = 5 years; Pacifici et al. ;2013) including the past and the present (2010 to 2025). This decline is expected to continue due to increasing persecution, hunting pressure, and ongoing decline in the extent and quality of its forest habitat used for roosting and foraging.	Pteropus lylei is known to form colonies up to 8,000 individuals (CITE) in trees that can become stripped of leaves by the bats' activity. The species has been documented to co-roost with Pteropus vampyrus in at least three of the four known roosts in Viet Nam where the largest mixed species colony (1,500 to 2,000 individuals) was historically found in the Doi pagoda, P. lylei ;was the more abundant species (Son et al. ;2009). Most colonies for the species are located in close proximity to human populations where the roosts receive some degree of protection and foraging areas are available in the surrounding agricultural landscapes (Weber et al. ;2015, Chaiyes et al. 2017, Choden et al. ; 2019, Duengkae et al. ;2019). In Thailand, 53% of the roost sites were located within Buddhist temples (Chaiyes et al. 2017) where the bats receive some degree of protection during the day (Weber et al. ;2015, Chaiyes et al. 2017, Duengkae et al. ;2019). Similarly, many flying fox roosts in Cambodia are located in pagodas where monks provide protection from hunting (Choden et al. 2019). The roosting sites of P. lylei were at an average elevation of 23.73 m (0–81 m), and near rivers and water bodies (Chaiyes et al. 2017). The species has been documented to travel from 2.2–23.6 km between day roosts and foraging areas (Weber et al. ; 2015) and in some landscapes, it may travel up to 105 km between colonies (Choden et al. ;2019). In Kandal, Cambodia, radio tagged bats flew ca 20 km from the roost to the foraging area which were often in residential areas (53.7%) and plantations (26.6%), traveling an average of 28.3 km per night (range 6.9–105 km, Choden et al. ;2019). In Cambodia, 6 of 14 tagged P. lylei used a second roost site for at least one night (Choden et al. ;2019) and relatively frequent movements among colonies in Thailand have resulted in high levels of genetic connectedness of the broader species distribution in the region (Sukgosa et al. 2018). Once individuals arrive at their foraging area, they make small-scale movements within their foraging area and appear to exhibit high foraging area fidelity within a season (Weber et al. ; 2015). Although most tracked bats foraged in farmland, plantations, and gardens, research indicates that small remnants of mangroves are important habitats of P. lylei (Weber et al. ;2015). The species has a fairly diverse diet (34 food plant species) of exotic crops and native plants available in a landscape (Weber et al. ;2015). As it feeds extensively in orchards, it is considered a pest in Thailand (S. Bumrungsri pers. comm.).  Pteropus lylei have been found to have unusually high antibody titers to the Nipah Virus, suggesting that it circulates in this species in Thailand and Cambodia (Wacharapluesadee et al. , 2005). Nothing is known of the species ecology in China (Smith et al. 2008), but it is logical to assume similar patterns are observed. Research is needed to fully understand the species distribution, ecology, habitat use, and population connectedness.	The close proximity between P. lylei and people has resulted in direct conflict as farmers view bats as major pests in agricultural lands. Persecution, hunting, and forest degradation and loss are the major threats to P. lylei. In Thailand, 57% of known roost sites that contained ca 62% of the species total population was located in areas where hunting was known to occur (Chaiyes et al. 2017). In Viet Nam, flying foxes are hunted to supply speciality restaurants in some towns in Soc Trang, Ca Mau, and Kien Giang provinces (Choden et al. 2019). It is also threatened by hunting in Cambodia (P. Bates pers. comm.). Native forests are degraded and converted to agriculture or to community development where the bats are hunted and persecuted. The threat of increased persecution of P. lylei out of fear of Nipah (Wacharapluesadee et al. ;2005) and other viruses has escalated with the impacts of the COVID-19 global pandemic (Fenton et al. ; 2020).	A comprehensive range-wide population assessment for the species is lacking, but systematic counts at 30 roosts in Thailand provided an estimate of ca 75,000 bats (Chaiyes et al. 2017). There are at least four (4) known colonies in Viet Nam (Son et al. ;2009), while the number of colonies in Cambodia and Yunnan China are unknown. Most colonies in Thailand are ca 1,000 bats (Sukgosa et al. ;2018), there is a colony of ca 4,000 bats in Cambodia (Choden et al. 2019), and the largest known colony is in Thailand with ca 8,000 bats (Chaiyes et al. 2017). The species total population and the mature individuals is suspected to be declining by 30-35% over a 15-year period including the past and the present (2010 to 2025). This decline is expected to continue due to increasing trade and hunting pressure and ongoing decline in the extent and quality of its forest habitat used for roosting and foraging.	Decreasing	This species is known from Cambodia (Choden et al. 2019), central and eastern Thailand (Weber et al. 2015, Chaiyes et al. 2017, Sukgosa et al. ;2018, Duengkae et al. ;2019), and Viet Nam (Son et al. 2009). There is an unconfirmed record of the species in Yunnan in China (Smith et al. 2008).	The species is hunted for food and traditional uses. It is not known to be exported.	Terrestrial	<p>Pteropus lylei is listed on Appendix II of CITES and is protected by the Wildlife Preservation and Protection Act, B.E. 2535 (1992) in Thailand which prohibits hunting, possession, and trade or export (Chaiyes et al. 2017). The species is protected within designated areas, including religious establishments (temples and mosques), national parks, wildlife sanctuaries, and non-hunting areas (Chaiyes et al. 2017). The species is known from at least two national parks (Vo Doi National Park, Ca Mau Province and U Minh Thuong National Park, Kien Giang Province) in Viet Nam (Son et al. 2009) but is not known from other protected areas across the species broader distribution. Roosts sites are protected by monks at temples in Thailand (Weber et al. 2015, Chaiyes et al. 2017, Duengkae et al. 2019) and by monks at pagodas in Vietnam (Son et al. 2019). Conservation, restoration, and management across landscapes is warranted to enable protection of remaining native trees patches and associated natural vegetation, which should help reduce bat-human conflict over the bats foraging in orchards, plantations, and gardens (Weber et al. ;2015, Choden et al. 2019). Effective and sustain awareness campaigns with the public and governments are needed to inform proactive management and minimize persecution. Expanded research is needed to improve our understanding of the species population status and trends, ecology, habitat requirements, landscape movements, and threat mitigation. Enhanced protection and management of known roosts and restoration of native forests to reduce foraging pressure in agricultural landscapes are important to the species survival (Weber et al. ;2015). There also needs to be effective work on conflict reduction/resolution with farmers since native forests are not going to recover in the short term. </p>	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Pteropus		lylei	K. Andersen	1908	0	Ann. Mag. Nat. Hist.	ser. 8, 2: 367	Lyle's Flying Fox	None.	Thailand, Bangkok.	Thailand, Vietnam, Cambodia.	Appendix II	Vulnerable	 vampyrus species group; see Almeida et al. (2014).	Pteropus lylei	1004472	23	Lyle's Flying Fox		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTEROPODINI	Pteropus	NA	lylei	K. Andersen	1908	0						"Bangkok, Siam [= Thailand]."			lylei K. Andersen, 1908	NA	NA				Thailand|Cambodia|Vietnam	Asia	Indomalaya	VU	0	0	0	Pteropus_lylei	0	sciname match	Pteropus_lylei	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pteropus_lylei	1004472	23	Lyle's Flying Fox		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteropodini	Pteropus	NA	lylei	Andersen	0	Pteropus lylei	Andersen, K.C. 1908-10-01. Twenty new forms of _Pteropus_. Annals and Magazine of Natural History (8)2(10):361-370.	https://www.biodiversitylibrary.org/page/22071136	BMNH:Mamm:1904.4.7.2	holotype	https://data.nhm.ac.uk/object/0bbab2a6-ae65-4788-83d7-d4bea3947428	"Bangkok, Siam [= Thailand]."			NA	NA				Thailand|Cambodia|Vietnam	Asia	Indomalaya	VU	0	0	0	Pteropus_lylei	0	sciname match	Pteropus_lylei	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Pteropus		lylei	K. Andersen	1908	0	Ann. Mag. Nat. Hist.	ser. 8, 2: 367	Lyle's Flying Fox	None.	Thailand, Bangkok.	Thailand, Vietnam, Cambodia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Appendix II</a>	<a href='https://www.iucnredlist.org/species/18734/22082429/' target='_blank'>Vulnerable</a>	vampyrusspecies group; see Almeida et al. (2014).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pteropus lylei; Pteropus lylei; Pteropus lylei; Pteropus lylei; Pteropus lylei; Pteropus lylei; lombocensis; heudei; salottii; lylei; Roussette de Lyle; Lyle-Flughund; Zorro volador de Lyle; Lyle's Flying Fox; Lyle's Flying Fox; Lyle's Flying Fox; P. lylei