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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1264	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus	Pteropus chrysoproctus		[MSW3] chrysoproctus species group. See Flannery (1995b). May not include argentatus, a taxon based on a badly damaged immature specimen thought to be from Ambon (K. Helgen, pers. comm.). Sulawesi specimens previously referred to argentatus were allocated to Acerodon celebensis by Musser et al. (1982a). This complex includes several undescribed species (K. Helgen, pers. comm.).; [HMW] Pteropus chrysoproctus Temminck, 1837 , Ambon , Maluku Islands , Indonesia . Pteropus chrysoproctus is in the melanopogon species group. It might not include synonymized P. argentatus , Sulawesi specimens previously referred to as P. argentatus were identified as Acerodon celebensis . It might contain undescribed subspecies. Monotypic.; [batnames2022]  samoensis species group; see Almeida et al. (2014). See Flannery (1995b). May not include argentatus, a taxon based on a badly damaged immaturespecimen thought to be from Ambon (K. Helgen, pers. comm.). Sulawesi specimens previously referred to argentatus were allocated to Acerodon  celebensis by Musser et al. (1982a). This complex includes several undescribed species (K. Helgen, pers. comm.).; [IUCN] Simmons (2005) includes P. argentatus , as a synonym of Pteropus chrysoproctus .; [batnames2023]  samoensis species group; see Almeida et al. (2014). See Flannery (1995b). May not include argentatus, a taxon based on a badly damaged immaturespecimen thought to be from Ambon (K. Helgen, pers. comm.). Sulawesi specimens previously referred to argentatus were allocated to Acerodon  celebensis by Musser et al. (1982a). This complex includes several undescribed species (K. Helgen, pers. comm.).; [batnames2025_1.7] samoensis species group; see Almeida et al. (2014). See Flannery (1995b). May not include argentatus, a taxon based on a badly damaged immaturespecimen thought to be from Ambon (K. Helgen, pers. comm.). Sulawesi specimens previously referred to argentatus were allocated to Acerodon celebensis by Musser et al. (1982a). This complex includes several undescribed species (K. Helgen, pers. comm.).									argentatus			chrysoproctus 	chrysoproctus - argentatus	chrysoproctus, argentatus, argentatus	Simmons (2005) includes P. argentatus , as a synonym of Pteropus chrysoproctus .	chrysoproctus 	chrysoproctus - argentatus	chrysoproctus, argentatus, argentatus	chrysoproctus, argentatus, argentatus	chrysoproctus 	chrysoproctus - argentatus	chrysoproctus Temminck, 1837|argentatus J. E. Gray, 1843 [nomen nudum]|argentatus J. E. Gray, 1844		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Amboina flying fox	Sanghir Is, S Moluccas	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pteropus chrysoproctus	Indonesia, Molucca Isis., Amboina.	Temminck	1837	Monogr. Mamm., 2:67.	Distribu tion: Occurring in the central Moluccas (Ceram, Buru, and surrounding small islands) and San ghirs.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Amboina flying fox	S Moluccas; ref. 4.110	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1837	Monogr. Mamm., 2:67.		Amboina, Buru, Seram, and small islands east of Seram (Indonesia). A Sangihe Isl record is erroneous; see Bergmans and Rozendaal (1988:65).	Indonesia, Molucca Isis, Amboina.		TEMMINCK	1837	Tibia thinly haired dorsally. Size large (forearm length, 163-177 mm).	Distribu tion: Occurring in the central Moluccas (Ceram, Buru, and surrounding small islands) and San ghirs.	No subspecies.		24	species	P. chrysoproctus	TEMMINCK	1837	Pteropus	genus	Pteropus chrysoproctus				Tibia thinly haired dorsally. Size large (forearm length, 163-177 mm).	No subspecies.		27. P. chrysoproctus TEMMINCK 1837 [chrysop roctus group],	27	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Pteropus chrysoproctus	Pteropus		chrysoproctus	Temminck		1837		Monogr. Mamm.	2		67		Moluccan Flying Fox	Indonesia, Maluku, Ambon.	Ambon, Buru, Seram, and small islands east of Seram (Indonesia). A Sangihe Isl record is erroneous; see Bergmans and Rozendaal (1988).	CITES – Appendix II. IUCN/SSC Action Plan (1992) – No Data. IUCN 2003 – Lower Risk (nt).	argentatus Gray, 1843 [nomen nudum]; argentatus Gray, 1844.	chrysoproctus species group. See Flannery (1995b). May not include argentatus, a taxon based on a badly damaged immature specimen thought to be from Ambon (K. Helgen, pers. comm.). Sulawesi specimens previously referred to argentatus were allocated to Acerodon celebensis by Musser et al. (1982a). This complex includes several undescribed species (K. Helgen, pers. comm.).	03AD87FAFF9AF67489B63F12F5EAFB5B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	162	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFACF64289B1368BF8B4FDD7.xml	Pteropus chrysoproctus	Pteropodidae	Pteropus	chrysoproctus	Temminck	1837	Roussette d'Ambon @fr | Sidmolukken-Flughund @de | Zorro volador de Ambon @es | Ambon Flying Fox @en	Pteropus chrysoproctus Temminck, 1837 , Ambon , Maluku Islands , Indonesia . Pteropus chrysoproctus is in the melanopogon species group. It might not include synonymized P. argentatus , Sulawesi specimens previously referred to as P. argentatus were identified as Acerodon celebensis . It might contain undescribed subspecies. Monotypic.	C Moluccas Is (Buru, Ambon , and Seram) and Watubela Archipelago in Indonesia .	Head-body 190-265 mm (tailless), ear 27-30 mm , hindfoot 48-55 mm , forearm 163-177 mm ; weight 300-420 g . Greatest lengths of skulls are 67-77 mm and tibias 75-80 mm . Ears of the South Moluccan Flying Fox are of moderate length and curved into pointed tips. Bases of most hairs are tawny to rufous. On Seram, body is dark chestnut-brown, with more gradation to red-chestnut on sides of upper chest and flanks. Mantle is rufous, with sides occasionally grading to golden rufous to golden toward nape. Crown and sides of head are dark rufous, with back grading from lighter mantle into chestnut. A Buru specimen has dark burnt umber on back, with more golden tawny-tinged hairs toward rump, dark chestnut chest and belly, nearly ocherous buffy thighs, and tawny head, mantle, and sides of neck. Forearm is naked, and upperside oftibia is sparsely furred or naked. Wing membranes are black from uppersides of body. Index claw is present. Canines are slender at bases, I, and P| are smaller, and cheekteeth are narrower compared with Black-bearded Flying Fox ( P. melanopogon ). 1, is 1-5-2 times larger than I. Canines of males are noticeably longer and slightly heavier than females.	Old growth forest and recently mangrove forest.	The South Moluccan Flying Fox is probably frugivorous, but anecdotal evidence from Seram suggests that it feeds on durian ( Durio sp. , Malvaceae ) flowers. Hunters have seen it foraging in stands of Ceiba ( Malvaceae ) and orchards of cultivated fruits ( Syzygium , Myrtaceae ).	No information.	The South Moluccan Flying Fox is crepuscular and nocturnal. It roosts on mangrove islands during the day and leaves around dusk to forage in forested areas with low levels of disturbance.	The South Moluccan Flying Fox is generally gregarious and roosts in large colonies in forested areas. A colony of ¢.300 individuals was found in an unprotected mangrove forest. Single individuals and groups of less than ten individuals have been seen in temporary roosts in fragmented forests near villages. It roosts with the Black-bearded Flying Fox in separate trees.	CITES Appendix II. Classified as Vulnerable on The IUCN Red List. Population decline of the South Moluccan Flying Fox is suspected to have been more than 30% over three generations due to hunting and habitat loss. It is usually hunted at foraging sites as a snack for drinking parties. Deforestation, mining, oil exploration, and agroforestry expansion has resulted in loss of almost one-half of total forest coverage. Some foraging sites occur in Manusela National Park on Seram, but day roosts occur in unprotected mangrove forests on offshore islands nearby. There is no effective protection, and no protected area on Buru. It is likely extinct on Ambon .	Andersen (1912b) | Corbet & Hill (1992) | Flannery (1995a) | Musseretal. (1982) | Simmons (2005) | Tsang (2016j) | Tsang et al. (2015)		190. South Moluccan Flying Fox Pteropus chrysoproctus French: Roussette d'Ambon / German: Sidmolukken-Flughund / Spanish: Zorro volador de Ambon Other common names: Ambon Flying Fox Taxonomy. Pteropus chrysoproctus Temminck, 1837 , Ambon , Maluku Islands , Indonesia . Pteropus chrysoproctus is in the melanopogon species group. It might not include synonymized P. argentatus , Sulawesi specimens previously referred to as P. argentatus were identified as Acerodon celebensis . It might contain undescribed subspecies. Monotypic. Distribution. C Moluccas Is (Buru, Ambon , and Seram) and Watubela Archipelago in Indonesia . Descriptive notes. Head-body 190-265 mm (tailless), ear 27-30 mm , hindfoot 48-55 mm , forearm 163-177 mm ; weight 300-420 g . Greatest lengths of skulls are 67-77 mm and tibias 75-80 mm . Ears of the South Moluccan Flying Fox are of moderate length and curved into pointed tips. Bases of most hairs are tawny to rufous. On Seram, body is dark chestnut-brown, with more gradation to red-chestnut on sides of upper chest and flanks. Mantle is rufous, with sides occasionally grading to golden rufous to golden toward nape. Crown and sides of head are dark rufous, with back grading from lighter mantle into chestnut. A Buru specimen has dark burnt umber on back, with more golden tawny-tinged hairs toward rump, dark chestnut chest and belly, nearly ocherous buffy thighs, and tawny head, mantle, and sides of neck. Forearm is naked, and upperside oftibia is sparsely furred or naked. Wing membranes are black from uppersides of body. Index claw is present. Canines are slender at bases, I, and P| are smaller, and cheekteeth are narrower compared with Black-bearded Flying Fox ( P. melanopogon ). 1, is 1-5-2 times larger than I. Canines of males are noticeably longer and slightly heavier than females. Habitat. Old growth forest and recently mangrove forest. Food and Feeding. The South Moluccan Flying Fox is probably frugivorous, but anecdotal evidence from Seram suggests that it feeds on durian ( Durio sp. , Malvaceae ) flowers. Hunters have seen it foraging in stands of Ceiba ( Malvaceae ) and orchards of cultivated fruits ( Syzygium , Myrtaceae ). Breeding. No information. Activity patterns. The South Moluccan Flying Fox is crepuscular and nocturnal. It roosts on mangrove islands during the day and leaves around dusk to forage in forested areas with low levels of disturbance. Movements, Home range and Social organization. The South Moluccan Flying Fox is generally gregarious and roosts in large colonies in forested areas. A colony of ¢.300 individuals was found in an unprotected mangrove forest. Single individuals and groups of less than ten individuals have been seen in temporary roosts in fragmented forests near villages. It roosts with the Black-bearded Flying Fox in separate trees. Status and Conservation. CITES Appendix II. Classified as Vulnerable on The IUCN Red List. Population decline of the South Moluccan Flying Fox is suspected to have been more than 30% over three generations due to hunting and habitat loss. It is usually hunted at foraging sites as a snack for drinking parties. Deforestation, mining, oil exploration, and agroforestry expansion has resulted in loss of almost one-half of total forest coverage. Some foraging sites occur in Manusela National Park on Seram, but day roosts occur in unprotected mangrove forests on offshore islands nearby. There is no effective protection, and no protected area on Buru. It is likely extinct on Ambon . Bibliography. Andersen (1912b), Corbet & Hill (1992), Flannery (1995a), Musseretal. (1982), Simmons (2005), Tsang (2016j), Tsang et al. (2015).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Pteropus chrysoproctus	Pteropus		chrysoproctus	Temminck	1837	0	Monogr. Mamm.	0.1299	Moluccan Flying Fox	 argentatus Gray, 1843 [ nomen nudum ], argentatus Gray, 1844.	Indonesia, Maluku, Ambon.	Ambon, Buru, Seram, and small islands east of Seram (Indonesia). A Sangihe Isl record is erroneous; see Bergmans and Rozendaal (1988).	Appendix II	Vulnerable	 samoensis species group; see Almeida et al. (2014). See Flannery (1995b). May not include argentatus, a taxon based on a badly damaged immaturespecimen thought to be from Ambon (K. Helgen, pers. comm.). Sulawesi specimens previously referred to argentatus were allocated to Acerodon  celebensis by Musser et al. (1982a). This complex includes several undescribed species (K. Helgen, pers. comm.).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pteropus chrysoproctus	23	South Moluccan Flying Fox	Ambon Flying Fox	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTEROPODINI	Pteropus	NA	chrysoproctus	Temminck	1837	0	Pteropus_chrysoproctus	Temminck, C. J. (1837). Monographies de mammalogie, ou Description de quelques genres de mammifères, dont les espèces ont été observées dans les différens musées de l'Europe, Vol. 2. Leiden, 67.	https://www.google.com/books/edition/Monographies_de_mammalogie/yHQQAQAAMAAJ?hl=en&gbpv=1&dq=monographies+de+mammalogie+temminck+volume+2&printsec=frontcover	ZMB 2626 [syntype]		Ambon, Maluku Islands, Indonesia.			chrysoproctus Temminck, 1837|argentatus J. E. Gray, 1843 [nomen nudum]|argentatus J. E. Gray, 1844	NA	NA	Indonesia	Oceania	Australasia/Oceania	VU	0	0	0	Pteropus_chrysoproctus	0	sciname match	Pteropus_chrysoproctus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	100000000	Pteropus chrysoproctus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Pteropus	chrysoproctus	Temminck, 1837	Simmons (2005) includes P. argentatus , as a synonym of Pteropus chrysoproctus .	20000000	Pteropus chrysoproctus	Vulnerable	A3cd	2016	2016-01-18 00:00:00 UTC	3.1	English	The Moluccan Flying Fox is assessed as Vulnerable under A3cd because of a population decline of more than 30% is suspected to occur over the next three generations (24 years based on Pacifici et al.  2013). It is rarely encountered today despite being commonly hunted in previous decades, although it is difficult to determine how precipitous the decline in the species has been. As a distinctive, large species, hunting is a major threat and ongoing decline of its habitat is predicted to continue due to logging and agricultural expansion.	It is largely dependent on intact forest, but on Obi it was also collected while feeding in a garden, and several others when roosting in a Sago palm. It has also been collected from disturbed forest. It occurs in small groups in lightly disturbed areas. A large colony was recorded from an isolated mangrove forest in Seram (Tsang et al.  2015).	Deforestation due to clear-cut logging and agricultural expansion is a major threat. Hunting is also likely to be a problem as it is a large and distinctive bat.	This species has been rarely collected, but probably it is not uncommon. It was definitely detected more often in the 19th century than in recent surveys. Local people have noted that there are now fewer bats than there were in past decades. A large colony was found in a mangrove forest in Seram in a recent survey, but it is presently the only known persistent colony. However, the colony is located near Manusela National Park, but not within it, and only afforded protection by a local resort owner due to its ecotourism value (Tsang et al. 2015). Small groups have been seen in fragmentary secondary forest near villages as well (S.M. Tsang, pers. comm).	Decreasing	The species is found in the Maluku Islands, Ambon, Buru, Seram, and small islands east of Seram in Indonesia. A Sangihe Island record is erroneous (Bergmans and Rozendaal 1988). It is found below 250 m asl. It is now likely absent from Ambon.		Terrestrial	This species occurs in Manusela National Park. However, the only recorded roosting colony currently resides outside of the national park lands, and is afforded minor protection by a local resort owner due to its value for ecotourism. Taxonomic research is needed to resolve this species complex. Further surveys are needed to determine the current population status of this species, its roosting preferences, and its feeding ecology. It is listed on CITES Appendix II.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Pteropus		chrysoproctus	Temminck	1837	0	Monogr. Mamm.	0.129861	Moluccan Flying Fox	 argentatus Gray, 1843 [ nomen nudum ], argentatus Gray, 1844.	Indonesia, Maluku, Ambon.	Ambon, Buru, Seram, and small islands east of Seram (Indonesia). A Sangihe Isl record is erroneous; see Bergmans and Rozendaal (1988).	Appendix II	Vulnerable	 samoensis species group; see Almeida et al. (2014). See Flannery (1995b). May not include argentatus, a taxon based on a badly damaged immaturespecimen thought to be from Ambon (K. Helgen, pers. comm.). Sulawesi specimens previously referred to argentatus were allocated to Acerodon  celebensis by Musser et al. (1982a). This complex includes several undescribed species (K. Helgen, pers. comm.).	Pteropus chrysoproctus	1004457	23	South Moluccan Flying Fox	Ambon Flying Fox	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTEROPODINI	Pteropus	NA	chrysoproctus	Temminck	1837	0	Pteropus_chrysoproctus	Temminck, C. J. (1837). Monographies de mammalogie, ou Description de quelques genres de mammifères, dont les espèces ont été observées dans les différens musées de l'Europe, Vol. 2. Leiden, 67.	https://www.google.com/books/edition/Monographies_de_mammalogie/yHQQAQAAMAAJ?hl=en&gbpv=1&dq=monographies+de+mammalogie+temminck+volume+2&printsec=frontcover	ZMB 2626 [syntype]		Ambon, Maluku Islands, Indonesia.			chrysoproctus Temminck, 1837|argentatus J. E. Gray, 1843 [nomen nudum]|argentatus J. E. Gray, 1844	NA	NA				Indonesia	Oceania	Australasia/Oceania	VU	0	0	0	Pteropus_chrysoproctus	0	sciname match	Pteropus_chrysoproctus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pteropus_chrysoproctus	1004457	23	South Moluccan Flying Fox	Ambon Flying Fox	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteropodini	Pteropus	NA	chrysoproctus	Temminck	0	Pteropus chrysoproctus	Temminck, C.J. 1837. Livraison 2. Pp. 49–140 in Temminck, C.J. 1835-1841. Monographies de Mammalogie. Tome second. C. C. van der Hoek, Leiden, 392 pp.	https://archive.org/details/monographiedema00temmgoog	RMNH.MAM.37984, RMNH.MAM.37985, RMNH.MAM.37986, RMNH.MAM.37987, RMNH.MAM.37988, ZMB 2626	syntypes	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37984.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37984.b | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37985.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37985.b | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37986.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37986.b | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37987.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37987.b | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37988.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37988.b	Ambon, Maluku Islands, Indonesia.			NA	NA				Indonesia	Oceania (Continent)	Australasia	VU	0	0	0	Pteropus_chrysoproctus	0	sciname match	Pteropus_chrysoproctus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Pteropus		chrysoproctus	Temminck	1837	0	Monogr. Mamm.	0.129861	Moluccan Flying Fox	argentatus Gray, 1843 [nomen nudum], argentatus Gray, 1844.	Indonesia, Maluku, Ambon.	Ambon, Buru, Seram, and small islands east of Seram (Indonesia). A Sangihe Isl record is erroneous; see Bergmans and Rozendaal (1988).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Appendix II</a>	<a href='https://www.iucnredlist.org/species/99688187/22078625/' target='_blank'>Vulnerable</a>	samoensis species group; see Almeida et al. (2014). See Flannery (1995b). May not include argentatus, a taxon based on a badly damaged immaturespecimen thought to be from Ambon (K. Helgen, pers. comm.). Sulawesi specimens previously referred to argentatus were allocated to Acerodon celebensis by Musser et al. (1982a). This complex includes several undescribed species (K. Helgen, pers. comm.).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pteropus chrysoproctus; Pteropus chrysoproctus; Pteropus chrysoproctus; Pteropus chrysoproctus; Pteropus chrysoproctus; Pteropus chrysoproctus; argentatus; argentatus; chrysoproctus; argentatus; argentatus; Roussette d'Ambon; Sidmolukken-Flughund; Zorro volador de Ambon; Ambon Flying Fox; South Moluccan Flying Fox; Ambon Flying Fox; Moluccan Flying Fox; Moluccan Flying Fox; P. chrysoproctus