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(1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name 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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1261	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pteropus brunneus	Pteropus hypomelanus [synonym of]	Pteropus brunneus	Pteropus brunneus	Pteropus brunneus	Pteropus brunneus	N/A	Pteropus brunneus	Pteropus brunneus	Pteropus brunneus	Pteropus brunneus	Pteropus brunneus	Pteropus brunneus	Pteropus brunneus	Pteropus brunneus		[MSW2] See Koopman (1984c:2-3) for status.; [MSW3] subniger species group. Known only from the holotype. It is not clear that this taxon represents a valid species, see Koopman (1984c).; [batnames2022]  samoensis species group; see Almeida et al. (2014). Known only from the holotype. It is not clear that this taxon represents a valid species, see Koopman (1984c).; [IUCN] The taxonomic status of this species is unclear (Mickleburgh et al. 1992). Conder (2008) states that the species is distinct from all other Australian Pteropus species, but there has been speculation that the single specimen might in fact be a vagrant of another species. Despite much survey effort, the species has not been found on the Percy Islands recently. Koopman (1984) discusses the uncertain taxonomic status of P. brunneus at length due to how distinct it is from other known Pteropus in the region. More taxonomic work is needed.; [batnames2023]  samoensis species group; see Almeida et al. (2014). Known only from the holotype. It is not clear that this taxon represents a valid species, see Koopman (1984c).; [batnames2025_1.7] samoensisspecies group; see Almeida et al. (2014). Known only from the holotype. It is not clear that this taxon represents a valid species, see Koopman (1984c).														brunneus	The taxonomic status of this species is unclear (Mickleburgh et al. 1992). Conder (2008) states that the species is distinct from all other Australian Pteropus species, but there has been speculation that the single specimen might in fact be a vagrant of another species. Despite much survey effort, the species has not been found on the Percy Islands recently. Koopman (1984) discusses the uncertain taxonomic status of P. brunneus at length due to how distinct it is from other known Pteropus in the region. More taxonomic work is needed.			brunneus 	brunneus 			brunneus Dobson, 1878		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Percy I, off E Queensland; perhaps applied to a vagrant P. hypomelanus		N/A					Distribution: A poorly known form (only one slightly immature specimen collected on the Percy island off the eastern coast of Queensland, Australia and possibly an accidental from some where in the Solomon islands) and conspecific with P. admiralitatum (KOOPMAN 1984 a).		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Percy I, off E Queensland; perhaps applied to a vagrant P. hypomelanus	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Dobson	1878	Cat. Chiroptera Brit. Mus., p. 37.	See Koopman (1984c:2-3) for status.	Known from the type locality only.	Australia, Queensland, Percy Island.		DOBSON	1878	Medium in size (forearm length, 118 mm; maxil lary tooth row length, 24 mm). Tibia hairy dor sally.	Distribution: A poorly known form (only one slightly immature specimen collected on the Percy island off the eastern coast of Queensland, Australia and possibly an accidental from some where in the Solomon islands) and conspecific with P. admiralitatum (KOOPMAN 1984 a).	No subspecies.		22	species	P. brunneus	DOBSON	1878	Pteropus	genus	Pteropus brunneus				Medium in size (forearm length, 118 mm; maxil lary tooth row length, 24 mm). Tibia hairy dor sally.	No subspecies.		8. P. brunneus DOBSON 1878 [subniger group],	8	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Pteropus brunneus	Pteropus		brunneus	Dobson		1878		Cat. Chiroptera Brit. Mus.			37		Dusky Flying Fox	Australia, Queensland, Percy Isl.	Known from the type locality only.	CITES – Appendix II. IUCN/SSC Action Plan (1992) – Extinct? IUCN 2003 – Extinct.		subniger species group. Known only from the holotype. It is not clear that this taxon represents a valid species, see Koopman (1984c).								N/A					1878														Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Pteropus brunneus	Pteropus		brunneus	Dobson	1878	0	Cat. Chiroptera Brit. Mus.	p. 37	Dusky Flying Fox	None.	Australia, Queensland, Percy Isl.	Known from the type locality only.	Not listed.	Extinct	 samoensis species group; see Almeida et al. (2014). Known only from the holotype. It is not clear that this taxon represents a valid species, see Koopman (1984c).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pteropus brunneus	23	Percy Island Flying Fox		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTEROPODINI	Pteropus	NA	brunneus	Dobson	1872	0						Percy Is, Queensland, Australia.			brunneus Dobson, 1878	NA	NA	Australia	Oceania	Australasia/Oceania	EX	1	0	0	Pteropus_brunneus	0	sciname match	Pteropus_brunneus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	18718	Pteropus brunneus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Pteropus	brunneus	Dobson, 1878	The taxonomic status of this species is unclear (Mickleburgh et al. 1992). Conder (2008) states that the species is distinct from all other Australian Pteropus species, but there has been speculation that the single specimen might in fact be a vagrant of another species. Despite much survey effort, the species has not been found on the Percy Islands recently. Koopman (1984) discusses the uncertain taxonomic status of P. brunneus at length due to how distinct it is from other known Pteropus in the region. More taxonomic work is needed.	20000000	Pteropus brunneus	Extinct		2020	2019-08-09 00:00:00 UTC	3.1	English	This species is listed as Extinct because it has not been found in its only known range, or nearby, much after the original collection in ;1874 ;(specific date uncertain) despite extensive surveys.	Nothing is known about the habitat requirements of this species.	It is possible that the species declined and eventually became extinct because of its vulnerability to habitat loss (Conder 2008).	This species is known only from the holotype. It was reported as being plentiful at the close of the 19<sup>th</sup> century (Conder 2008).		This species is known only from a single specimen (collected in 1874) from the Percy Islands, Queensland, Australia, and subsequent non-specimen records of a colony (L. Hall pers. comm.). It is not known from which island the specimen originally came. Despite more survey effort recently, the species has not been found on the Percy Islands.		Terrestrial	This species is listed on Appendix II of CITES. Despite extensive surveys so far, further field studies on the Percy Islands and other islands in the region are needed to determine if any remnant populations of this species persist (L. Hall pers. comm.). There is also a need to try to find skeletal remains near old camps, in order to confirm the locality of this specimen (L. Hall pers. comm.). Additional taxonomic work is needed to resolve the status of this species.	Australasian	1874 (with certainty); observations from the 1890s	FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Pteropus		brunneus	Dobson	1878	0	Cat. Chiroptera Brit. Mus.	p. 37	Dusky Flying Fox	None.	Australia, Queensland, Percy Isl.	Known from the type locality only.	Not listed.	Extinct	 samoensis species group; see Almeida et al. (2014). Known only from the holotype. It is not clear that this taxon represents a valid species, see Koopman (1984c).	Pteropus brunneus	1004454	23	Percy Island Flying Fox		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTEROPODINI	Pteropus	NA	brunneus	Dobson	1872	0						Percy Is, Queensland, Australia.			brunneus Dobson, 1878	NA	NA				Australia	Oceania	Australasia/Oceania	EX	1	0	0	Pteropus_brunneus	0	sciname match	Pteropus_brunneus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pteropus_brunneus	1004454	23	Percy Island Flying Fox		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteropodini	Pteropus	NA	brunneus	Dobson	0	Pteropus brunneus	Dobson, G.E. 1878-06-15. Catalogue of the Chiroptera in the Collection of the British Museum. Taylor and Francis, London, 567 pp.	https://www.biodiversitylibrary.org/page/37695103	BMNH:Mamm:1874.3.16.2	holotype	https://data.nhm.ac.uk/object/2d2cd00e-50c4-4e0b-87e8-6a1a191469d5	Percy Is, Queensland, Australia.			NA	NA				Australia	Oceania (Continent)	Australasia	EX	1	0	0	Pteropus_brunneus	0	sciname match	Pteropus_brunneus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Pteropus		brunneus	Dobson	1878	0	Cat. Chiroptera Brit. Mus.	p. 37	Dusky Flying Fox	None.	Australia, Queensland, Percy Isl.	Known from the type locality only.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/18718/22078015/' target='_blank'>Extinct</a>	samoensisspecies group; see Almeida et al. (2014). Known only from the holotype. It is not clear that this taxon represents a valid species, see Koopman (1984c).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pteropus brunneus; Pteropus brunneus; Pteropus brunneus; Pteropus brunneus; Pteropus brunneus; brunneus; Percy Island Flying Fox; Dusky Flying Fox; Dusky Flying Fox; P. brunneus