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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1251	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens	Pteronotus quadridens		[MSW2] Subgenus Chilonycteris. Includes torrei; For use of quadridens in place of fuliginosus; see Silva-Taboada (1976:7). See Rodriguez-Dur4n and Kunz (1992, Mammalian Species, 395).; [MSW3] Subgenus Chilonycteris. Includes torrei; For use of quadridens in place of fuliginosus, see Silva-Taboada (1976). See Rodríguez-Durán and Kunz (1992) and Timm and Genoways (2003).; [HMW] Lobostoma quadridens Gundlach, 1840 , “Cafetal St. Antonio el Fundador,” Canimar, Cuba . G. Silva-Taboada in 1976 concluded that the description of Lobostoma quadridens by J. Gundlach in 1840 referred to the same species described by J. E. Gray in 1843 as Chilonycteris fuliginosa and consequently should be considered the senior synonym of P. fuliginosus, which was formerly recognized byJ. D. Smith in 1972 in his revision of Mormoopidae . Fossil records of P. quadridens are reported for many islands in the Bahamas , suggesting it had a more extensive distribution in the late Quaternary than today. Two subspecies recognized.; [batnames2022] Subgenus Chilonycteris . Includes torrei ; see Smith (1972). For use of quadridens in place of fuliginosus , see Silva-Taboada (1976). See Rodr&iacute;guez-Dur&aacute;n and Kunz (1992) and Timm and Genoways (2003).; [IUCN] Subgenus  Chilonycteris . Includes torrei . See Rodríguez-Durán and Kunz (1992) and Timm and Genoways (2003).; [batnames2023] Subgenus Chilonycteris . Includes torrei ; see Smith (1972). For use of quadridens in place of fuliginosus , see Silva-Taboada (1976). See Rodr&iacute;guez-Dur&aacute;n and Kunz (1992) and Timm and Genoways (2003).; [batnames2025_1.7] Subgenus Chilonycteris. Includes torrei; see Smith (1972). For use of quadridens in place of fuliginosus, see Silva-Taboada (1976). See Rodr&iacute;guez-Dur&aacute;n and Kunz (1992) and Timm and Genoways (2003).					(fuliginosus)	fuliginosus, inflata, torrei.	fuliginosus, quadridens	quadridens, fuliginosus	torrei; fuliginosus - inflata	quadridens, fuliginosus		quadridens, fuliginosus	quadridens - torrei; fuliginosus - inflata	quadridens, fuliginosus, inflata, torrei	Subgenus  Chilonycteris . Includes torrei . See Rodríguez-Durán and Kunz (1992) and Timm and Genoways (2003).	quadridens, fuliginosus	quadridens - torrei; fuliginosus - inflata	quadridens, fuliginosus, inflata, torrei 	quadridens, fuliginosus, inflatus, torrei	fuliginosus, quadridens	fuliginosus - inflata; torrei - fuliginosus	quadridens (J. Gundlach, 1840)|fuliginosus (J. E. Gray, 1843)|inflatus (Rehn, 1904)|torrei (G. M. Allen, 1916)						N/A					Distribution: Confined to the Greater Antilles.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Sooty moustached bat	Greater Antilles; ref. 4.39	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Gundlach	1840	Arch. Naturgesch., 6:357.	Subgenus Chilonycteris. Includes torrei; For use of quadridens in place of fuliginosus; see Silva-Taboada (1976:7). See Rodriguez-Dur4n and Kunz (1992, Mammalian Species, 395).	Cuba, Jamaica, Hispaniola, Puerto Rico.	Cuba, Matanzas, Canimar.		GUNDLACH	1840	Nasolabial plate with fairly well-developed lateral processes. Plagiopatagium and uropatagium attached to tar sus by a long ligament. Size small (forearm length, 35-40 mm; condylobasal length, 12-14 mm).	Distribution: Confined to the Greater Antilles.	Two subspecies are currently recognized:	P. q. fuliginosus (= inflata) (Jamaica, Hispaniola, Puerto Rico), P. q. quadridens (= torrei) (Cuba).	71	species	P. quadridens	GUNDLACH	1840	Chilonycteris	subgenus	Pteronotus quadridens				Nasolabial plate with fairly well-developed lateral processes. Plagiopatagium and uropatagium attached to tar sus by a long ligament. Size small (forearm length, 35-40 mm; condylobasal length, 12-14 mm).	Two subspecies are currently recognized:		3. P. quadridens (GUNDLACH 1840).	3	_P. q. fuliginosus_ (Gray, 1843) (synonyms: _inflatus_ (Rehn, 1904)); _P. q. quadridens_ (Gundlach, 1840) (synonyms: _torrei_ (Allen, 1916))			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Mormoopidae			Pteronotus quadridens	Pteronotus	Chilonycteris	quadridens	Gundlach	y	1840		Arch. Naturgesch.	6		357		Sooty Mustached Bat	Cuba, Matanzas, Canimar.	Cuba, Jamaica, Hispaniola, Puerto Rico.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	torrei G. M. Allen, 1916; fuliginosus Gray, 1843; inflata Rehn, 1904.	Subgenus Chilonycteris. Includes torrei; For use of quadridens in place of fuliginosus, see Silva-Taboada (1976). See Rodríguez-Durán and Kunz (1992) and Timm and Genoways (2003).	03A63743915EFFE2E4E4CDC6284BA4B6	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Mormoopidae_424.pdf.imf	hash://md5/ff9f4f3b9157ffebe472c9232f51a072	439	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/37/03A637439152FFEEE1E1CA2B280AADFC.xml	Pteronotus quadridens	Mormoopidae	Pleronotus	quadridens	Gundlach	1840	Ptéronote fuligineux @fr | Rul Rfarbene Schnurrbartfledermaus @de | Pteronotus tiznado @es	Lobostoma quadridens Gundlach, 1840 , “Cafetal St. Antonio el Fundador,” Canimar, Cuba . G. Silva-Taboada in 1976 concluded that the description of Lobostoma quadridens by J. Gundlach in 1840 referred to the same species described by J. E. Gray in 1843 as Chilonycteris fuliginosa and consequently should be considered the senior synonym of P. fuliginosus, which was formerly recognized byJ. D. Smith in 1972 in his revision of Mormoopidae . Fossil records of P. quadridens are reported for many islands in the Bahamas , suggesting it had a more extensive distribution in the late Quaternary than today. Two subspecies recognized.	P.q.quadridensGundlach,1840—Cuba. P. q. fuliginosusJ. E. Gray, 1843 — Jamaica , Hispaniola, and Puerto Rico .	Head-body c.40—-44 mm, tail 16-22 mm, ear 14-16 mm, hindfoot 8-10 mm, forearm 35-40 mm; weight 3-6 g. The Sooty Mustached Bat is the smallest species of mormoopid. It might be confused with individuals ofits sister species, Macleay’s Mustached Bat (P. macleayit), given similarity in external morphology and sympatric distribution, but it is slightly smaller. Pelage is distinctly tricolored dorsally and bicolored ventrally. Labionasal plate has small lateral spikes and 3-4 tubercles on margin above each nostril. Skull rostrum is delicate, slightly elevated, and longer than one-half total length of skull. Braincase is globular. Condylo-basal lengths are 12-14 mm. There is a geographical trend of size increase in overall size from west to east in Cuba . Subspecies fuliginosus is larger than quadridens externally and cranially.	Forest patches, semi-open areas, and near streams to forage and caves to roost. The Sooty Mustached Bat is commonly found foraging in the same habitats as Macleay’s Mustached Bat.	Sooty Mustached Bats eat a wide diversity of insects, including moths, beetles, flies, hymenopterans, homopterans, hemipterans, and orthopterans. It has been suggested that they are opportunistic feeders. Consumption of spiders was reported by different sources and has been associated with hovering ability.	The Sooty Mustached Bat is monoestrous and uniparous; there is one case of twins. Mating season starts in January. Pregnant females are most common in May but have been reported in February—June. Lactation occurs in June-September and peaks in July. Infants form dense clusters of 50-200 individuals in caves.	There is evidence of temporal separation in activity of the Sooty Mustached Bat from other species occupying the same cave. It is the first to leave the cave at dusk or a few minutes before sunset. In Cuba , it has initial activity peak lasting four hours after sunset and becomes active again before sunrise. It usually forages at lower heights in the forest understory than Macleay’s Mustached Bat and the Antillean Ghost-faced Bat ( Mormoops blainviller). Echolocation calls during search phase consist of short CF-FM pulses of c.4 milliseconds, with 2-3 harmonics. Second harmonic is usually most intense and starts with short CF component at ¢.83 kHz and ends at c.62 kHz. Recorded calls of enclosed Sooty Mustached Bats had shorter duration and greater bandwidth than echolocation calls normally found from freeflying bats.	Large colonies of Sooty Mustached Bats are commonly reported in hot and humid caves in association with other mormoopids and phyllostomids but in spatially separated clusters. They seem to be more restricted by roost conditions than larger congeners, occupying caves with elevated temperature (over 35°C). They can fly up to 9 km between roosts and foraging sites, and some individuals were able to return to their roost from distances up to 30 km . Males and females can disappear from the roost sites during mating season. There is evidence of sexual segregation during maternity period at least in some areas.	Classified as Least Concern on The IUCN Red List. The Sooty Mustached Bat is one of the most abundant bat species in Cuba and Puerto Rico and the least common mormoopid in Jamaica .	Genoways et al. (2005) | Gray (1843a) | Gundlach (1840) | Macias & Mora (2003) | Macias et al. (2006) | Mancina et al. (2012) | Morgan (2001) | Rodriguez-Duran & Kunz (1992) | Silva-Taboada (1976a, 1979) | Simmons & Conway (2001) | Smith (1972)	https://zenodo.org/record/6606802/files/figure.png	7. Sooty Mustached Bat Pleronotus quadridens French: Ptéronote fuligineux / German: RulRfarbene Schnurrbartfledermaus / Spanish: Pteronotus tiznado Taxonomy. Lobostoma quadridens Gundlach, 1840 , “Cafetal St. Antonio el Fundador,” Canimar, Cuba . G. Silva-Taboada in 1976 concluded that the description of Lobostoma quadridens by J. Gundlach in 1840 referred to the same species described by J. E. Gray in 1843 as Chilonycteris fuliginosa and consequently should be considered the senior synonym of P. fuliginosus, which was formerly recognized byJ. D. Smith in 1972 in his revision of Mormoopidae . Fossil records of P. quadridens are reported for many islands in the Bahamas , suggesting it had a more extensive distribution in the late Quaternary than today. Two subspecies recognized. Subspecies and Distribution. P.q.quadridensGundlach,1840—Cuba. P. q. fuliginosusJ. E. Gray, 1843 — Jamaica , Hispaniola, and Puerto Rico . Descriptive notes. Head-body c.40—-44 mm, tail 16-22 mm, ear 14-16 mm, hindfoot 8-10 mm, forearm 35-40 mm; weight 3-6 g. The Sooty Mustached Bat is the smallest species of mormoopid. It might be confused with individuals ofits sister species, Macleay’s Mustached Bat (P. macleayit), given similarity in external morphology and sympatric distribution, but it is slightly smaller. Pelage is distinctly tricolored dorsally and bicolored ventrally. Labionasal plate has small lateral spikes and 3-4 tubercles on margin above each nostril. Skull rostrum is delicate, slightly elevated, and longer than one-half total length of skull. Braincase is globular. Condylo-basal lengths are 12-14 mm. There is a geographical trend of size increase in overall size from west to east in Cuba . Subspecies fuliginosus is larger than quadridens externally and cranially. Habitat. Forest patches, semi-open areas, and near streams to forage and caves to roost. The Sooty Mustached Bat is commonly found foraging in the same habitats as Macleay’s Mustached Bat. Food and Feeding. Sooty Mustached Bats eat a wide diversity of insects, including moths, beetles, flies, hymenopterans, homopterans, hemipterans, and orthopterans. It has been suggested that they are opportunistic feeders. Consumption of spiders was reported by different sources and has been associated with hovering ability. Breeding. The Sooty Mustached Bat is monoestrous and uniparous; there is one case of twins. Mating season starts in January. Pregnant females are most common in May but have been reported in February—June. Lactation occurs in June-September and peaks in July. Infants form dense clusters of 50-200 individuals in caves. Activity patterns. There is evidence of temporal separation in activity of the Sooty Mustached Bat from other species occupying the same cave. It is the first to leave the cave at dusk or a few minutes before sunset. In Cuba , it has initial activity peak lasting four hours after sunset and becomes active again before sunrise. It usually forages at lower heights in the forest understory than Macleay’s Mustached Bat and the Antillean Ghost-faced Bat ( Mormoops blainviller). Echolocation calls during search phase consist of short CF-FM pulses of c.4 milliseconds, with 2-3 harmonics. Second harmonic is usually most intense and starts with short CF component at ¢.83 kHz and ends at c.62 kHz. Recorded calls of enclosed Sooty Mustached Bats had shorter duration and greater bandwidth than echolocation calls normally found from freeflying bats. Movements, Home range and Social organization. Large colonies of Sooty Mustached Bats are commonly reported in hot and humid caves in association with other mormoopids and phyllostomids but in spatially separated clusters. They seem to be more restricted by roost conditions than larger congeners, occupying caves with elevated temperature (over 35°C). They can fly up to 9 km between roosts and foraging sites, and some individuals were able to return to their roost from distances up to 30 km . Males and females can disappear from the roost sites during mating season. There is evidence of sexual segregation during maternity period at least in some areas. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Sooty Mustached Bat is one of the most abundant bat species in Cuba and Puerto Rico and the least common mormoopid in Jamaica . Bibliography. Genoways et al. (2005), Gray (1843a), Gundlach (1840), Macias & Mora (2003), Macias et al. (2006), Mancina et al. (2012), Morgan (2001), Rodriguez-Duran & Kunz (1992), Silva-Taboada (1976a, 1979), Simmons & Conway (2001), Smith (1972).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Mormoopidae	Pteronotus quadridens	Pteronotus	Chilonycteris	quadridens	Gundlach	1840	1	Arch. Naturgesch.	0.4979	Sooty Mustached Bat	 torrei G. M. Allen, 1916; <b>fuliginosus</b> Gray, 1843; inflata Rehn, 1904	Cuba, Matanzas, Canimar	Greater Antilles (Cuba, Jamaica, Hispaniola, Puerto Rico)	Not listed.	Least Concern	Subgenus Chilonycteris . Includes torrei ; see Smith (1972). For use of quadridens in place of fuliginosus , see Silva-Taboada (1976). See Rodr&iacute;guez-Dur&aacute;n and Kunz (1992) and Timm and Genoways (2003).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pteronotus quadridens	23	Sooty Mustached Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	MORMOOPIDAE	NA	NA	Pteronotus	NA	quadridens	Gundlach	1840	1	Lobostoma_quadridens	Gundlach, J. (1840). Beschreibung von vier auf Cuba gefangenen Fledermäusen. Wiegmanns Archiv für Naturgeschichte, 1840, 6, 357.	https://www.biodiversitylibrary.org/item/31587#page/367/mode/1up	ZMB 2458, ZMB 2807, ZMB 2869, ZMB 4210, ZMB 6075, ZMB 67414-67416, ZMB 67418, ZMB A2311a, ZMB A2259b [syntypes]		"Cafetal St. Antonio el Fundador," Canimar, Cuba.			quadridens (Gundlach, 1840)|fuliginosus (J. E. Gray, 1843)|inflata (Rehn, 1904)|torrei (G. M. Allen, 1916)	NA	NA	Cuba|Jamaica|Haiti|Dominican Republic|Puerto Rico	North America	Neotropic	LC	0	0	0	Pteronotus_quadridens	0	sciname match	Pteronotus_quadridens	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	18710	Pteronotus quadridens	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MORMOOPIDAE	Pteronotus	quadridens	(Gundlach, 1840)	Subgenus  Chilonycteris . Includes torrei . See Rodríguez-Durán and Kunz (1992) and Timm and Genoways (2003).	20000000	Pteronotus quadridens	Least Concern		2016	2016-07-01 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its abundance within its restricted distribution, its presumed large population, and because its habitat is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species roosts during the day in deep recesses of hot caves, where it form roosting aggregations containing thousands of individuals (Gannon et al. ;2005, Genoways et al . 2005). An estimate of 140,000 bats of this species roosts in Cucaracha Cave, Puerto Rico (Gannon et al . 2005). A cave occupied by this bat usually shelters two to five other species. It is insectivorous, and apparently is an opportunistic forager to some degree, like many insectivorous bats. Insects from one to seven different orders have been found in stomach or faecal samples of a single bat; moths, flies, and true bugs are taken consistently as well, and wasps and flying ants, which occur in large but unpredictable swarms, are eaten when available. Females generally give birth to a single young; twinning is extremely rare. Pregnant females are found from February through June, with the largest percentage occurring in May, when births begin (Silva-Taboada 1979, Gannon et al . 2005).	Major threats are on hot caves. Caves are not protected in Jamaica (Davalos pers. comm.). It is extinct from Abaco, Andros and New Providence (Bahamas) (Turvey pers. comm.).	It is abundant in Puerto Rico (Gannon et al. ;2005).	Stable	This species is known from Cuba, Jamaica, Hispaniola, and Puerto Rico (Simmons 2005). Extinct from the Bahamas (Morgan 2001).		Terrestrial	The species is found in protected areas.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Mormoopidae	Pteronotus	Chilonycteris	quadridens	Gundlach	1840	1	Arch. Naturgesch.	0.497917	Sooty Mustached Bat	 torrei G. M. Allen, 1916; <b>fuliginosus</b> Gray, 1843; inflata Rehn, 1904	Cuba, Matanzas, Canimar	Greater Antilles (Cuba, Jamaica, Hispaniola, Puerto Rico)	Not listed.	Least Concern	Subgenus Chilonycteris . Includes torrei ; see Smith (1972). For use of quadridens in place of fuliginosus , see Silva-Taboada (1976). See Rodr&iacute;guez-Dur&aacute;n and Kunz (1992) and Timm and Genoways (2003).	Pteronotus quadridens	1004866	23	Sooty Mustached Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Mormoopidae	NA	NA	Pteronotus	NA	quadridens	Gundlach	1840	1	Lobostoma_quadridens	Gundlach, J. (1840). Beschreibung von vier auf Cuba gefangenen Fledermäusen. Wiegmanns Archiv für Naturgeschichte, 1840, 6, 357.	https://www.biodiversitylibrary.org/item/31587#page/367/mode/1up	ZMB 2458, ZMB 2807, ZMB 2869, ZMB 4210, ZMB 6075, ZMB 67414-67416, ZMB 67418, ZMB A2311a, ZMB A2259b [syntypes]		"Cafetal St. Antonio el Fundador," Canimar, Cuba.			quadridens (Gundlach, 1840)|fuliginosus (J. E. Gray, 1843)|inflata (Rehn, 1904)|torrei (G. M. Allen, 1916)	NA	NA				Cuba|Jamaica|Haiti|Dominican Republic|Puerto Rico	North America	Neotropic	LC	0	0	0	Pteronotus_quadridens	0	sciname match	Pteronotus_quadridens	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pteronotus_quadridens	1004866	23	Sooty Mustached Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Mormoopidae	NA	NA	Pteronotus	Chilonycteris	quadridens	J. Gundlach	1	Lobostoma quadridens	Gundlach, J. 1840. Beschreibung von vier auf Cuba gefangenen Fledermäusen. Archiv für Naturgeschichte 6(1):356-358.	https://www.biodiversitylibrary.org/page/7203178	ZMB 2458, ZMB 2484, ZMB 2485, ZMB 2807, ZMB 2869, ZMB 4210, ZMB 6075, ZMB 67413, ZMB 67414, ZMB 67415, ZMB 67416, ZMB 67418, ZMB 67419, ZMB (unnumbered)	syntypes		"Cafetal St. Antonio el Fundador," Canimar, Cuba.			NA	NA				Cuba|Jamaica|Haiti|Dominican Republic|Puerto Rico	North America	Neotropic	LC	0	0	0	Pteronotus_quadridens	0	sciname match	Pteronotus_quadridens	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Mormoopidae	Pteronotus	Chilonycteris	quadridens	Gundlach	1840	1	Arch. Naturgesch.	0.497917	Sooty Mustached Bat	torrei G. M. Allen, 1916; fuliginosus Gray, 1843; inflata Rehn, 1904	Cuba, Matanzas, Canimar	Greater Antilles (Cuba, Jamaica, Hispaniola, Puerto Rico)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/18710/22076753/' target='_blank'>Least Concern</a>	Subgenus Chilonycteris. Includes torrei; see Smith (1972). For use of quadridens in place of fuliginosus, see Silva-Taboada (1976). See Rodr&iacute;guez-Dur&aacute;n and Kunz (1992) and Timm and Genoways (2003).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pteronotus quadridens; Pteronotus quadridens; Pteronotus quadridens; Pteronotus quadridens; Pteronotus quadridens; Pteronotus quadridens; quadridens; fuliginosus; torrei; fuliginosus - inflata; quadridens; fuliginosus; fuliginosus; torrei; fuliginosus - inflata; quadridens; fuliginosus; inflata; torrei; Ptéronote fuligineux; Rul Rfarbene Schnurrbartfledermaus; Pteronotus tiznado; Sooty Mustached Bat; Sooty Mustached Bat; Sooty Mustached Bat; P. quadridens