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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1213	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Plecotus auritus [synonym of]	Plecotus austriacus wardi	Plecotus austriacus wardi	Plecotus wardi	Plecotus wardi	Plecotus wardi	Plecotus wardi	Plecotus wardi	Plecotus wardi	Plecotus wardi	Plecotus wardi	Plecotus wardi		[HMW] Plecotus wardi Thomas, 1911 , Jammu and Kashmir , India . This species is monotypic.; [batnames2022]  auritus species group. Distinct from austriacus; see Spitzenberger et al. (2006). Does not include mordax , which is apparently a synonym of kozlovi ; see Spitzenberger et al. (2006). Possibly conspecific with ariel and P . sp. 2 (Spitzenberger et al. (2006); see Fukui et al. (2020). Pending further study we retain both  ariel and wardi as separate species.; [MDD2022] split from P. austriacus; includes ariel, which has been considered a distinct species; [IUCN] Previously considered as a synonym of ;P. austriacus . Here considered as a distinct species based on Spitzenberger et al. (2006).; [batnames2023]  auritus species group. Distinct from austriacus; see Spitzenberger et al. (2006). Includes ariel and P. sp. 2 (Spitzenberger et al., 2006); see Fukui et al. (2020). Does not include kozlovi or mordax , which is apparently a synonym of kozlovi ; see Spitzenberger et al. (2006).; [MDD2023] split from P. austriacus; includes ariel, which has been considered a distinct species; [MDD2025_2.0] split from P. austriacus; includes ariel, which has been considered a distinct species; [batnames2025_1.7] auritus species group. Distinct from austriacus; see Spitzenberger et al. (2006). Includes ariel and P. sp. 2 (Spitzenberger et al., 2006); see Fukui et al. (2020). Does not include kozlovi or mordax, which is apparently a synonym of kozlovi; see Spitzenberger et al. (2006).; [MDD2025_2.2] split from P. austriacus; includes ariel, which has been considered a distinct species														wardi, ariel	Previously considered as a synonym of ;P. austriacus . Here considered as a distinct species based on Spitzenberger et al. (2006).	wardi	wardi - ariel	ariel, wardi	ariel, wardi	wardi 	wardi - ariel	ariel O. Thomas, 1911|wardi O. Thomas, 1911						N/A																																								NA																											03AD87FAFF81F66C8C753E69FB55F62F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	869	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF946A2BFF8394301DF7BCBA.xml	Plecotus wardi	Vespertilionidae	Plecotus	wardi	Thomas	1911	Oreillard de Ward @fr | Ward-Langohr @de | Orejudo de Ward @es	Plecotus wardi Thomas, 1911 , Jammu and Kashmir , India . This species is monotypic.	N Afghanistan , N Pakistan , N India , and Nepal .	Head—-body 47-53 mm , tail 49-54 mm , car 37-42 mm , hindfoot 7-9- 5 mm , forearm 42-46 mm . Fur is dense (not woolly, 10-13 mm long), brownish, sometimes with golden tips, on dorsum; sparser and pale gray on venter, with white tips ( 6-8 mm long); dorsal hairs tricolored, with dark brown bases, narrow lighter middle zone, and brown tips; ventral hairs bicolored with dark bases and white tips. Face and chin meagerly furred with short hairs. Patagium can be naked or furred; ears and patagium generally brownish. Toes long (c. 8 mm ), slender, and partially covered by long hair. The species seems to be closely related to Strelkov’s Long-eared Bat (P. strelkovr) in skull and dental characteristics, with medium-sized braincase and short, slender, flat rostrum, but it has larger I and canines than its closest relatives, with broader anterior part of zygomatic arch. Condylo-basal length c.16- 5 mm ; zygomatic breadth c.9- 2 mm ; and upper tooth-row length c.6- 5 mm .	Tunnels in flanks of barren mountains surrounded by coniferous forests. Someareas ofits range (e.g. Nepal ) have a very contrasting dry climate. Usually found at elevations of 1700-3600 m .	No information.	No information.	No information.	No information.	Not assessed on The IUCN Red Lust.	Ellerman & Morrison-Scott (1951) | Etemad (1967) | Spitzenberger, Strelkov & Haring (2003) | Spitzenberger, Strelkov, Winkler & Haring (2006) | Srinivasulu & Srinivasulu (2012)	https://zenodo.org/record/6398316/files/figure.png	239. Ward’s Long-eared Bat Plecotus wardi French: Oreillard de Ward / German: Ward-Langohr / Spanish: Orejudo de Ward Taxonomy. Plecotus wardi Thomas, 1911 , Jammu and Kashmir , India . This species is monotypic. Distribution. N Afghanistan , N Pakistan , N India , and Nepal . Descriptive notes. Head—-body 47-53 mm , tail 49-54 mm , car 37-42 mm , hindfoot 7-9- 5 mm , forearm 42-46 mm . Fur is dense (not woolly, 10-13 mm long), brownish, sometimes with golden tips, on dorsum; sparser and pale gray on venter, with white tips ( 6-8 mm long); dorsal hairs tricolored, with dark brown bases, narrow lighter middle zone, and brown tips; ventral hairs bicolored with dark bases and white tips. Face and chin meagerly furred with short hairs. Patagium can be naked or furred; ears and patagium generally brownish. Toes long (c. 8 mm ), slender, and partially covered by long hair. The species seems to be closely related to Strelkov’s Long-eared Bat (P. strelkovr) in skull and dental characteristics, with medium-sized braincase and short, slender, flat rostrum, but it has larger I and canines than its closest relatives, with broader anterior part of zygomatic arch. Condylo-basal length c.16- 5 mm ; zygomatic breadth c.9- 2 mm ; and upper tooth-row length c.6- 5 mm . Habitat. Tunnels in flanks of barren mountains surrounded by coniferous forests. Someareas ofits range (e.g. Nepal ) have a very contrasting dry climate. Usually found at elevations of 1700-3600 m . Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The IUCN Red Lust. Bibliography. Ellerman & Morrison-Scott (1951), Etemad (1967), Spitzenberger, Strelkov & Haring (2003), Spitzenberger, Strelkov, Winkler & Haring (2006), Srinivasulu & Srinivasulu (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Plecotus wardi	Plecotus		wardi	Thomas	1911	0	Ann. Mag. Nat. Hist.	ser.8, 7:209	Ward&apos;s Long-eared Bat	None.	Leh, Ladakh, Jammu and Kashmir, India	N Pakistan, W India through Nepal.	Not listed.	Least Concern	 auritus species group. Distinct from austriacus; see Spitzenberger et al. (2006). Does not include mordax , which is apparently a synonym of kozlovi ; see Spitzenberger et al. (2006). Possibly conspecific with ariel and P . sp. 2 (Spitzenberger et al. (2006); see Fukui et al. (2020). Pending further study we retain both  ariel and wardi as separate species.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Plecotus wardi	23	Ward's Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PLECOTINI	Plecotus	NA	wardi	O. Thomas	1911	0	Plecotus_wardi	Thomas, O. (1911). New Asiatic Muridae. Annals and Magazine of Natural History, ser. 8, 7, 209.	https://www.biodiversitylibrary.org/item/53774#page/229/mode/1up	BM 1906.10.3.2		Jammu and Kashmir, India.			wardi O. Thomas, 1911|ariel O. Thomas, 1911	split from P. austriacus; includes ariel, which has been considered a distinct species	Spitzenberger, F., Strelkov, P. P., Winkler, H., & Haring, E. (2006). A preliminary revision of the genus Plecotus (Chiroptera, Vespertilionidae) based on genetic and morphological results. Zoologica Scripta, 35(3), 187-230.|Fukui, D., Tan Tu, V., Thanh, H. T., Arai, S., Harada, M., Csorba, G., & Son, N. T. (2020). First record of the genus Plecotus from Southeast Asia with notes on the taxonomy, karyology and echolocation call of P. homochrous from Vietnam. Acta Chiropterologica, 22(1), 57-74.	Afghanistan|Pakistan|India|Nepal|China	Asia	Indomalaya	LC	0	0	0	Plecotus_wardi	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Plecotus wardi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Plecotus	wardi	Thomas, 1911	Previously considered as a synonym of ;P. austriacus . Here considered as a distinct species based on Spitzenberger et al. (2006).	90000000	Plecotus wardi	Least Concern		2020	2020-07-05 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution. However there is continuing decline in habitat quality in its range and presumed threat to this species due to tourism-related development; more accurate information in future may lead to more precise assessment.	Occurs in caves and caverns surrounded by coniferous forests, and occasionally in human habitation (T.A. Shah, pers. comm.).	<p>Little is known about this species, however it is potentially threatened by decline in habitat quality due to cave tourism and related development, cave destruction in certain areas, and demolition of abandoned buildings where it may roost.</p>	The species is occasionally reported.	Unknown	A palaearctic species found in the Himalayan regions from Pakistan to Nepal, and the Pamir mountains in Afghanistan. It occurs at elevations between 1,700 m to 3,600 m asl.	Nothing is known about its use and trade.	Terrestrial		Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Plecotus		wardi	Thomas	1911	0	Ann. Mag. Nat. Hist.	ser.8, 7:209	Ward's Long-eared Bat	ariel	Leh, Ladakh, Jammu and Kashmir, India	N Afghanistan, N Pakistan, W India through Nepal; Sichuan (China)	Not listed.	Least Concern	 auritus species group. Distinct from austriacus; see Spitzenberger et al. (2006). Includes ariel and P. sp. 2 (Spitzenberger et al., 2006); see Fukui et al. (2020). Does not include kozlovi or mordax , which is apparently a synonym of kozlovi ; see Spitzenberger et al. (2006).	Plecotus wardi	1005678	23	Ward's Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PLECOTINI	Plecotus	NA	wardi	O. Thomas	1911	0	Plecotus_wardi	Thomas, O. (1911). New Asiatic Muridae. Annals and Magazine of Natural History, ser. 8, 7, 209.	https://www.biodiversitylibrary.org/item/53774#page/229/mode/1up	BM 1906.10.3.2		Jammu and Kashmir, India.			wardi O. Thomas, 1911|ariel O. Thomas, 1911	split from P. austriacus; includes ariel, which has been considered a distinct species	Spitzenberger, F., Strelkov, P. P., Winkler, H., & Haring, E. (2006). A preliminary revision of the genus Plecotus (Chiroptera, Vespertilionidae) based on genetic and morphological results. Zoologica Scripta, 35(3), 187-230.|Fukui, D., Tan Tu, V., Thanh, H. T., Arai, S., Harada, M., Csorba, G., & Son, N. T. (2020). First record of the genus Plecotus from Southeast Asia with notes on the taxonomy, karyology and echolocation call of P. homochrous from Vietnam. Acta Chiropterologica, 22(1), 57-74.				Afghanistan|Pakistan|India|Nepal|China	Asia	Indomalaya	LC	0	0	0	Plecotus_wardi	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Plecotus_wardi	1005678	23	Ward's Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Plecotini	Plecotus	NA	wardi	O. Thomas	0	Plecotus wardi	Thomas, O. 1911-02-01. New Asiatic Muridae. Annals and Magazine of Natural History (8)7(38):205-209.	https://www.biodiversitylibrary.org/page/15814917	BMNH:Mamm:1906.10.3.2	holotype	https://data.nhm.ac.uk/object/09434b68-d610-454a-912a-2f647565c99e	Jammu and Kashmir, India.			split from P. austriacus; includes ariel, which has been considered a distinct species	Spitzenberger, F., Strelkov, P. P., Winkler, H., & Haring, E. (2006). A preliminary revision of the genus Plecotus (Chiroptera, Vespertilionidae) based on genetic and morphological results. Zoologica Scripta, 35(3), 187-230.|Fukui, D., Tan Tu, V., Thanh, H. T., Arai, S., Harada, M., Csorba, G., & Son, N. T. (2020). First record of the genus Plecotus from Southeast Asia with notes on the taxonomy, karyology and echolocation call of P. homochrous from Vietnam. Acta Chiropterologica, 22(1), 57-74.				Afghanistan|Pakistan|India|Nepal|China	Asia	Indomalaya	LC	0	0	0	Plecotus_wardi	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Plecotus		wardi	Thomas	1911	0	Ann. Mag. Nat. Hist.	ser.8, 7:209	Ward's Long-eared Bat	ariel	Leh, Ladakh, Jammu and Kashmir, India	N Afghanistan, N Pakistan, W India through Nepal; Sichuan (China)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85535265/85535306/' target='_blank'>Least Concern</a>	auritus species group. Distinct from austriacus; see Spitzenberger et al. (2006). Includes ariel and P. sp. 2 (Spitzenberger et al., 2006); see Fukui et al. (2020). Does not include kozlovi or mordax, which is apparently a synonym of kozlovi; see Spitzenberger et al. (2006).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Plecotus wardi; Plecotus wardi; Plecotus wardi; Plecotus wardi; Plecotus wardi; wardi; ariel; Oreillard de Ward; Ward-Langohr; Orejudo de Ward; Ward's Long-eared Bat; Ward's Long-eared Bat; P. wardi
