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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1208	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Plecotus sardus	Plecotus sardus	Plecotus sardus	Plecotus sardus	Plecotus sardus	Plecotus sardus	Plecotus sardus	Plecotus sardus	Plecotus sardus	Plecotus sardus		[MSW3] Closely related to auritus and alpinus, but clearly distinct.; [HMW] Plecotus sardus Mucedda et al., 2002 , “interior of a cave at Lanaitto’s Valley, Oliena District, Nuoro Province, middle-east Sardinia , Italy ( 40°15'29"N , 9°29'13"E , 150 mas.l.).” Previously treated within either P. austriacus or P. auritus , but recently elevated to species level based on morphological and molecular data. Monotypic.; [batnames2022]  auritus species group. Closely related to auritus and alpinus , but clearly distinct.; [IUCN] Identified as a new species by genetic analyses, this species combines morphological features of both P. austriacus and P. auritus .; [batnames2023]  auritus species group. Closely related to auritus and alpinus , but clearly distinct.; [batnames2025_1.7] auritus species group. Closely related to auritus and alpinus, but clearly distinct.														sardus	Identified as a new species by genetic analyses, this species combines morphological features of both P. austriacus and P. auritus .			sardus 	sardus 			sardus Mucedda, Kiefer, Pidinchedda, & Veith, 2002						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Plecotini	Plecotus sardus	Plecotus		sardus	Mucedda, Kiefer, Pidinchedda, and Veith		2002		Acta Chiropterol.	4		123		Sardinian Long-eared Bat	Italy, Sardinia, Nuoro Province, Oliena District, Lanaitto’s Valley, in a cave, 40°15'29”N, 09°29'13”E, 150 m.	Sardinia (Italy).	IUCN 2003 – Not evaluated (new species).		Closely related to auritus and alpinus, but clearly distinct.	4C3D87E8FF966A29FF8D9CC0189DB854	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	869	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF946A2BFA87930C1665B009.xml	Plecotus sardus	Vespertilionidae	Plecotus	sardus	Mucedda	2002	Oreillard de Sardaigne @fr | Sardinien-Langohr @de | Orejudode Cerdena @es	Plecotus sardus Mucedda et al., 2002 , “interior of a cave at Lanaitto’s Valley, Oliena District, Nuoro Province, middle-east Sardinia , Italy ( 40°15'29"N , 9°29'13"E , 150 mas.l.).” Previously treated within either P. austriacus or P. auritus , but recently elevated to species level based on morphological and molecular data. Monotypic.	Sardinia .	Head-body c. 45 mm , tail ¢.b 1 mm , ear 37- 5-39 mm , hindfoot 6-7-7- 7 mm , forearm 41-1-42- 2 mm . Dorsal fur woolly (c. 10 mm long), brownish, with hairs tricolored (dark brown bases, then grayish, with dark brown tips); ventral fur whitish or pale brown, with bicolored hairs (dark brown and whitish). Ears pale brown; tragus distinctive compared to congeners, being large (over 18 mm ) and broad (over 6 mm ), dark brown at base and yellowish at tip. Muzzle narrow, with round gland under chin. Wings brown, inserting at base of toe. Thumbs large; claws and hindfoot have sticking hair. Calcar long (c. 18 mm ) and slender, with small lobe at tip. Penis is almost cylindrical, unlike in the Brown Longeared Bat ( P. auritus ) and the Gray Long-eared Bat ( P. austriacus ); differs from the Alpine Long-eared Bat ( P. macrobullaris ) in baculum shape. P* is minute.	Very little information. Of three reports, two were from karstic areas with forests nearby at medium elevations, the third being at the coastline. It is suspected that the species is a forest-dweller, foraging within woodlands, and roosting in caves.	No information.	Only four maternity colonies are known with a total estimate of under 1000 breeding bats, but reproductive patterns remain poorly studied.	No information.	No movements have been reported.	Classified as Vulnerable on The IUCN Red List. Limited forest remains in Sardinia , while deforestation is increasing; the species may be at real risk of extinction in the near future. Also, roost visiting and disturbance may threaten potential roost sites.	Bosso et al. (2016) | Dietz & von Helversen (2004) | Dietz & Kiefer (2016) | GIRC (2004) | Mucedda et al. (2002) | Pacifici et al. (2013) | Piraccini (2016e) | Simmons (2005)	https://zenodo.org/record/6398320/files/figure.png	241. Sardinian Long-eared Bat Plecotus sardus French: Oreillard de Sardaigne / German: Sardinien-Langohr / Spanish: Orejudo de Cerdena Taxonomy. Plecotus sardus Mucedda et al., 2002 , “interior of a cave at Lanaitto’s Valley, Oliena District, Nuoro Province, middle-east Sardinia , Italy ( 40°15'29"N , 9°29'13"E , 150 mas.l.).” Previously treated within either P. austriacus or P. auritus , but recently elevated to species level based on morphological and molecular data. Monotypic. Distribution. Sardinia . Descriptive notes. Head-body c. 45 mm , tail ¢.b 1 mm , ear 37- 5-39 mm , hindfoot 6-7-7- 7 mm , forearm 41-1-42- 2 mm . Dorsal fur woolly (c. 10 mm long), brownish, with hairs tricolored (dark brown bases, then grayish, with dark brown tips); ventral fur whitish or pale brown, with bicolored hairs (dark brown and whitish). Ears pale brown; tragus distinctive compared to congeners, being large (over 18 mm ) and broad (over 6 mm ), dark brown at base and yellowish at tip. Muzzle narrow, with round gland under chin. Wings brown, inserting at base of toe. Thumbs large; claws and hindfoot have sticking hair. Calcar long (c. 18 mm ) and slender, with small lobe at tip. Penis is almost cylindrical, unlike in the Brown Longeared Bat ( P. auritus ) and the Gray Long-eared Bat ( P. austriacus ); differs from the Alpine Long-eared Bat ( P. macrobullaris ) in baculum shape. P* is minute. Habitat. Very little information. Of three reports, two were from karstic areas with forests nearby at medium elevations, the third being at the coastline. It is suspected that the species is a forest-dweller, foraging within woodlands, and roosting in caves. Food and Feeding. No information. Breeding. Only four maternity colonies are known with a total estimate of under 1000 breeding bats, but reproductive patterns remain poorly studied. Activity patterns. No information. Movements, Home range and Social organization. No movements have been reported. Status and Conservation. Classified as Vulnerable on The IUCN Red List. Limited forest remains in Sardinia , while deforestation is increasing; the species may be at real risk of extinction in the near future. Also, roost visiting and disturbance may threaten potential roost sites. Bibliography. Bosso et al. (2016), Dietz & von Helversen (2004), Dietz & Kiefer (2016), GIRC (2004), Mucedda et al. (2002), Pacifici et al. (2013), Piraccini (2016e), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Plecotus sardus	Plecotus		sardus	Mucedda, Kiefer, Pidinchedda & Veith	2002	0	Acta Chiropterol.	0.2521	Sardinian Long-eared Bat	None.	Italy, Sardinia, Nuoro Province, Oliena District, Lanaittoâs Valley, in a cave, 40Â°15'29âN, 09Â°29'13âE, 150 m.	Sardinia (Italy).	Not listed.	Vulnerable	 auritus species group. Closely related to auritus and alpinus , but clearly distinct.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Plecotus sardus	23	Sardinian Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PLECOTINI	Plecotus	NA	sardus	Mucedda, Kiefer, Pidinchedda, & Veith	2002	0	Plecotus_sardus	Mucedda, M., Kiefer, A., Pidinchedda, E. & Veith, M. (2002). A new species of long-eared bat (Chiroptera, Vespertilionidae) from Sardinia (Italy). Acta Chiropterologica, 4(2), 123.	https://bioone.org/journals/acta-chiropterologica/volume-4/issue-2/001.004.0202/A-New-Species-of-Long-Eared-Bat-Chiroptera-Vespertilionidae-from/10.3161/001.004.0202.full	DZAB 0023		"interior of a cave at Lanaitto's Valley, Oliena District, Nuoro Province, middle-east Sardinia, Italy (40Â°15'29"N, 9Â°29'13"E, 150 m a.s.l.)."	40.258	9.4869	sardus Mucedda, Kiefer, Pidinchedda, & Veith, 2002	NA	NA	Italy	Europe	Palearctic	VU	0	0	0	Plecotus_sardus	0	sciname match	Plecotus_sardus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136503	Plecotus sardus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Plecotus	sardus	Mucedda, Kiefer, Pidinchedda, &; Veith, 2002	Identified as a new species by genetic analyses, this species combines morphological features of both P. austriacus and P. auritus .	518549	Plecotus sardus	Vulnerable	B2ab(iii)	2016	2016-05-25 00:00:00 UTC	3.1	English	This species is considered to be Vulnerable at the Global, European and EU 28 levels. It is restricted to forest fragments in Sardinia (Italy). Since this species was first identified in 2002, it has been captured in only three localities, two in the same National Park. The population is probably very small, although more localities will probably be discovered in the future. Forests in Sardinia sum up to less than 2,000 km<sup>2</sup>, and their quality is declining.	The Sardinian Long-eared Bat occurs in the most wooded areas parts of Sardinia, and roosts in natural caves. Two of the three known localities are situated in karstic areas, one locality lies near the sea coast (Mucedda et al. 2002). Artificial habitats are not used. It is a sedentary species.	Roost disturbance (as a result of tourism) and habitat loss (caused by forestry management) are the main threats to the species.	This species was first described in 2002 and is poorly known. It is considered to be rare.	Decreasing	The Sardinian Long-eared Bat (Plecotus sardus ) ;is restricted to the island of Sardinia (Italy). The three known localities are situated near to the coast or at low elevations.		Terrestrial	It is protected by national legislation in most range states. There are also international legal obligations for its protection through the Bonn Convention (Eurobats) and Bern Convention. It is included in Annex IV of EU Habitats &; Species Directive. Two of the three known localities are in the National Park of Gennargentu and the Orosei Gulf. Recommended research actions include surveys and monitoring to determine distribution, status, and population trend. Protection of roosting and foraging habitat is also recommended.	Palearctic		FALSE	FALSE	Global, Europe & Mediterranean	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Plecotus		sardus	Mucedda, Kiefer, Pidinchedda & Veith	2002	0	Acta Chiropterol.	0.252083	Sardinian Long-eared Bat	None.	Italy, Sardinia, Nuoro Province, Oliena District, Lanaittoâ€™s Valley, in a cave, 40Â°15'29â€N, 09Â°29'13â€E, 150 m.	Sardinia (Italy).	Not listed.	Vulnerable	 auritus species group. Closely related to auritus and alpinus , but clearly distinct.	Plecotus sardus	1005673	23	Sardinian Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PLECOTINI	Plecotus	NA	sardus	Mucedda, Kiefer, Pidinchedda, & Veith	2002	0	Plecotus_sardus	Mucedda, M., Kiefer, A., Pidinchedda, E. & Veith, M. (2002). A new species of long-eared bat (Chiroptera, Vespertilionidae) from Sardinia (Italy). Acta Chiropterologica, 4(2), 123.	https://bioone.org/journals/acta-chiropterologica/volume-4/issue-2/001.004.0202/A-New-Species-of-Long-Eared-Bat-Chiroptera-Vespertilionidae-from/10.3161/001.004.0202.full	DZAB 0023		"interior of a cave at Lanaitto's Valley, Oliena District, Nuoro Province, middle-east Sardinia, Italy (40Â°15'29"N, 9Â°29'13"E, 150 m a.s.l.)."	40.25806	9.486944	sardus Mucedda, Kiefer, Pidinchedda, & Veith, 2002	NA	NA				Italy	Europe	Palearctic	VU	0	0	0	Plecotus_sardus	0	sciname match	Plecotus_sardus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Plecotus_sardus	1005673	23	Sardinian Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Plecotini	Plecotus	NA	sardus	Mucedda, Kiefer, Pidinchedda, & Veith	0	Plecotus sardus	Mucedda, M., Kiefer, A., Pidinchedda, E. and Veith, M. 2002-12. A new species of long-eared bat (Chiroptera, Vespertilionidae) from Sardinia (Italy). Acta Chiropterologica 4(2):121-135.	https://doi.org/10.3161/001.004.0202	DZAB 0023	holotype		"interior of a cave at Lanaitto's Valley, Oliena District, Nuoro Province, middle-east Sardinia, Italy (40Â°15'29"N, 9Â°29'13"E, 150 m a.s.l.)."	40.25806	9.486944	NA	NA				Italy	Europe	Palearctic	CR	0	0	0	Plecotus_sardus	0	sciname match	Plecotus_sardus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Plecotus		sardus	Mucedda, Kiefer, Pidinchedda & Veith	2002	0	Acta Chiropterol.	0.252083	Sardinian Long-eared Bat	None.	Italy, Sardinia, Nuoro Province, Oliena District, Lanaittoâ€™s Valley, in a cave, 40Â°15'29â€N, 09Â°29'13â€E, 150 m.	Sardinia (Italy).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136503/211020578/' target='_blank'>Critically Endangered</a>	auritus species group. Closely related to auritus and alpinus, but clearly distinct.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Plecotus sardus; Plecotus sardus; Plecotus sardus; Plecotus sardus; Plecotus sardus; Plecotus sardus; sardus; Oreillard de Sardaigne; Sardinien-Langohr; Orejudode Cerdena; Sardinian Long-eared Bat; Sardinian Long-eared Bat; Sardinian Long-eared Bat; P. sardus
