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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1206	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Plecotus auritus [synonym of]	N/A	Plecotus auritus sacrimontis	Plecotus ognevi	Plecotus ognevi	Plecotus ognevi	Plecotus ognevi	Plecotus ognevi	Plecotus ognevi	Plecotus ognevi	Plecotus ognevi	Plecotus ognevi		[HMW] Plecotus auritus ognevi Kishida, 1927 , Sakhalin , Russia . Previously considered a race of P. auritus . Monotypic.; [batnames2022]  sacrimontis species group. Distinct from auritus and  includes uenoi; see Strelkov (1988) and Spitzenberger et al. (2006). This name is considered a nomen nudum by Kaneko and Maeda (2002).; [MDD2022] split from P. auritus; [IUCN] In XX century was treated as a part of P. auritus and was known under the name P. a. sacrimontis G. Allen, 1908 (Strelkov, 1988, Tiunov, 1997). This taxon was elevated to species level and divided from P. sacrimontis by Spitzenberger et al. (2006). Border between P. ognevi and P. sacrimontis is not clear, e.g. population from Kuril Islands is allocated to P. ognevi by Spitzenberger et al. (2006) and to P. sacrimontis by Fukui (2015).; [batnames2023]  sacrimontis species group. Distinct from auritus and  includes uenoi; see Strelkov (1988) and Spitzenberger et al. (2006). This name is considered a nomen nudum by Kaneko and Maeda (2002).; [MDD2023] split from P. auritus; [MDD2025_2.0] split from P. auritus; [batnames2025_1.7] sacrimontis species group. Distinct from auritus and includes uenoi; see Strelkov (1988) and Spitzenberger et al. (2006). This name is considered a nomen nudum by Kaneko and Maeda (2002).; [MDD2025_2.2] split from P. auritus												ognevi	ognevi - uenoi	ognevi, uenoi, noemroegi	In XX century was treated as a part of P. auritus and was known under the name P. a. sacrimontis G. Allen, 1908 (Strelkov, 1988, Tiunov, 1997). This taxon was elevated to species level and divided from P. sacrimontis by Spitzenberger et al. (2006). Border between P. ognevi and P. sacrimontis is not clear, e.g. population from Kuril Islands is allocated to P. ognevi by Spitzenberger et al. (2006) and to P. sacrimontis by Fukui (2015).	ognevi, nomrogi 	ognevi - uenoi	ognevi, uenoi, noemroegi	ognevi, uenoi, noemroegi	nomrogi, ognevi 	ognevi - uenoi	ognevi Kishida, 1927|uenoi Imaizumi & Yoshiyuki, 1969|nomrogi Dolch, M. Stubbe, GÃ¤rtner, K. Thiele, Ariunbold, Batsaikhan, Lkhagvasuren, A. Stubbe, & Steinhauser, 2021 [nomen nudum]						N/A																																								NA																											4C3D87E8FF956A2BFA4A901C1D3DB911	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	867	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF966A29FA8895F51406B2A1.xml	Plecotus ognevi	Vespertilionidae	Plecotus	ognevi	Kishida	1927	Oreillard d'Ognev @fr | Ognev-Langohr @de | Orejudo de Ognev @es	Plecotus auritus ognevi Kishida, 1927 , Sakhalin , Russia . Previously considered a race of P. auritus . Monotypic.	Extreme E Kazakhstan and C Asian mountains E through Mongolia and NE China to Korea and Russian Far East, including Sakhalin I.	Head-body 48-50 mm , tail 44-46 mm , ear 31-33 mm , hindfoot c. 8 mm , forearm 40-42 mm ; weight 7-9 g . Similar to the Brown Long-eared Bat ( P. auritus ) but forearm is slightly larger, whereas it is shorter than in the Japanese Long-eared Bat ( P. sacrimontis ) and Kozlov’s Long-eared Bat ( P kozlovi ). Fur almost woolly, dark brown dorsally and pale brown to white ventrally (hairs bicolored, with dark bases and white tips); some individuals have some pale hairs surrounding neck and throat. Toes are brown and generally covered by long dense hair. Face quite dark, naked from nose to muzzle and eyes. Membranes usually translucent and delicate. Tragus 13-15 mm long. Tibia 18-20 mm long. Thumbs and claws remarkably long. Skull medium-sized, smaller in males than in females, very similar in structure to Strelkov’s Long-eared Bat ( P. strelkovi ); there is a diagnostic pronounced concavity in dorsal profile of anterior part of skull. Dental formula is 12/3, C1/1,P 2/3, M 3/3 (x2) = 36. Chromosomal complement has 2n = 32 and FNa = 50.	Typical foraging habitat includes forest-steppe,taiga, and mountain and temperate mixed deciduous forest, always in highlands; also river catchment areas and riparian vegetation.	Diet is mostly moths ( Lepidoptera ) and mosquitoes ( Diptera ). Forages both by aerial-hawking and by gleaning.	Breeds in small colonies of a maximum of few tens of individuals, roosting in crevices of old trees. A maternity colony was found in a cave near Beijing , China .	Recorded hibernating in a cave in the Primorsky Krai , Russia .	No information.	Classified as Least Concern on The IUCN Red List.	Datzmann et al. (2012) | Dorjderem (2004) | Kartavtseva et al. (2014) | Kruskop et al. (2012) | Nyambayar et al. (2010) | Orlova et al. (2016) | Scheffler et al. (2012) | Sokolov & Orlov (1980) | Spitzenberger et al. (2006) | Stubbe, Ariunbold, Buuveibaatar, Dorjderem, Monkhzul, Otgonbaatar & Tsogbadrakh (2008a) | Tiunov (2016)	https://zenodo.org/record/6398302/files/figure.png	233. Ognev’s Long-eared Bat Plecotus ognevi French: Oreillard d'Ognev / German: Ognev-Langohr / Spanish: Orejudo de Ognev Taxonomy. Plecotus auritus ognevi Kishida, 1927 , Sakhalin , Russia . Previously considered a race of P. auritus . Monotypic. Distribution. Extreme E Kazakhstan and C Asian mountains E through Mongolia and NE China to Korea and Russian Far East, including Sakhalin I. Descriptive notes. Head-body 48-50 mm , tail 44-46 mm , ear 31-33 mm , hindfoot c. 8 mm , forearm 40-42 mm ; weight 7-9 g . Similar to the Brown Long-eared Bat ( P. auritus ) but forearm is slightly larger, whereas it is shorter than in the Japanese Long-eared Bat ( P. sacrimontis ) and Kozlov’s Long-eared Bat ( P kozlovi ). Fur almost woolly, dark brown dorsally and pale brown to white ventrally (hairs bicolored, with dark bases and white tips); some individuals have some pale hairs surrounding neck and throat. Toes are brown and generally covered by long dense hair. Face quite dark, naked from nose to muzzle and eyes. Membranes usually translucent and delicate. Tragus 13-15 mm long. Tibia 18-20 mm long. Thumbs and claws remarkably long. Skull medium-sized, smaller in males than in females, very similar in structure to Strelkov’s Long-eared Bat ( P. strelkovi ); there is a diagnostic pronounced concavity in dorsal profile of anterior part of skull. Dental formula is 12/3, C1/1,P 2/3, M 3/3 (x2) = 36. Chromosomal complement has 2n = 32 and FNa = 50. Habitat. Typical foraging habitat includes forest-steppe,taiga, and mountain and temperate mixed deciduous forest, always in highlands; also river catchment areas and riparian vegetation. Food and Feeding. Diet is mostly moths ( Lepidoptera ) and mosquitoes ( Diptera ). Forages both by aerial-hawking and by gleaning. Breeding. Breeds in small colonies of a maximum of few tens of individuals, roosting in crevices of old trees. A maternity colony was found in a cave near Beijing , China . Activity patterns. Recorded hibernating in a cave in the Primorsky Krai , Russia . Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. Bibliography. Datzmann et al. (2012), Dorjderem (2004), Kartavtseva et al. (2014), Kruskop et al. (2012), Nyambayar et al. (2010), Orlova et al. (2016), Scheffler et al. (2012), Sokolov & Orlov (1980), Spitzenberger et al. (2006), Stubbe, Ariunbold, Buuveibaatar, Dorjderem, Monkhzul, Otgonbaatar & Tsogbadrakh (2008a), Tiunov (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Plecotus ognevi	Plecotus		ognevi	Kishida	1927	0	Zoological Magazine of Tokyo	45:58:00	Ognev&apos;s Long-eared Bat	 uenoi Imaizumi and Yoshiyuki, 1969.	no type locality as no holotype is designated	NE China, SE Russia and E Mongolia through SC Russia; Sakhalin and Itutrup Isls; extreme W Mongolia.	Not listed.	Least Concern	 sacrimontis species group. Distinct from auritus and  includes uenoi; see Strelkov (1988) and Spitzenberger et al. (2006). This name is considered a nomen nudum by Kaneko and Maeda (2002).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Plecotus ognevi	23	Ognev's Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PLECOTINI	Plecotus	NA	ognevi	Kishida	1927	0		Kishida, K. (1927). Review of Ognev's â€˜A synopsis of the Russian bats' with notes on some species. Zoological Magazine of Tokyo, 36, 138â€“193 (in Japanese).		NA		Sakhalin, Russia.			ognevi Kishida, 1927|uenoi Imaizumi & Yoshiyuki, 1969|noemroegi Dolch, M. Stubbe, GÃ¤rtner, Thiele, Ariunbold, Batsaikhan, Lkhagvasuren, A. Stubbe, & Steinhauser, 2021	split from P. auritus	Spitzenberger, F., Strelkov, P. P., Winkler, H., & Haring, E. (2006). A preliminary revision of the genus Plecotus (Chiroptera, Vespertilionidae) based on genetic and morphological results. Zoologica Scripta, 35(3), 187-230.	Kazakhstan|Mongolia|Russia|China|North Korea|South Korea	Asia	Palearctic	LC	0	0	0	Plecotus_ognevi	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136598	Plecotus ognevi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Plecotus	ognevi	Kishida, 1927	In XX century was treated as a part of P. auritus and was known under the name P. a. sacrimontis G. Allen, 1908 (Strelkov, 1988, Tiunov, 1997). This taxon was elevated to species level and divided from P. sacrimontis by Spitzenberger et al. (2006). Border between P. ognevi and P. sacrimontis is not clear, e.g. population from Kuril Islands is allocated to P. ognevi by Spitzenberger et al. (2006) and to P. sacrimontis by Fukui (2015).	20000000	Plecotus ognevi	Least Concern		2019	2018-08-08 00:00:00 UTC	3.1	English	Listed as Least Concern, because it is widespread, relatively adaptable and there are no major threats.	Eurytopic species: occurs in mountain and lowland taiga, mixed lowland and riverine deciduous forests and forest-steppe. May inhabits even dry steppes in case of presence of open water sources and suitable day roosts. Sometime occur in human populated places (Sokolov and Orlov 1980, Botvinkin 2002). Summer roosts situated mainly in hollow trees and under tree bark, however, also grottos, stone crevices and human buildings can be used. In Baikal area summer records of P. ognevi were made in railway tunnels (Botvinkin 2002). In steppe regions most probably caves and grottos could be the main roosts around the year. All known winter records were made in caves. P. ognevi feeds mainly on small and middle-size lepidopterans (about 95% of ration both in forest and steppe habitat, Noctuidae and Geometridae are dominate in food remains), but takes also insects of at least 8 other orders (Botvinkin et al. 1998). Presence in food remains of diurnal, poorly flying or non-flying insects suggests that part of its prey P. ognevi takes from tree bark, brunches or from soil surface. No exact information about breeding, but probably this species breeds in small colonies, young are born in June, usually just one offspring is produced.	There are no major threats to the species throughout its range.	There is no information on population sizes at present. Treated as rare species in Khabarovsky Krai, Zabaikalsky krai and Altaisky Kray, however, in some areas of Altai, Baikal area and Transbaikalia this is one of most common local bat species (S. Kruskop, pers. comm., August 2018). It is listed as LC in Mongolia, its population there is treated as quite large and stable (Clark et al. 2006). Large part of the species distribution range covers territory with quite low mankind population, there is high probability that in present the whole species population is stable.	Unknown	This species ranges from the Altai mountains in the Eastern Kazakhstan and middle Enisei basin, eastwards through southern Siberia, Transbaikalia, Mongolia, Russian Far East north to ca. 59 parallel (Tiunov 1997 as P. auritus ), northern China, to the Republic of Korea and the Russian island of Sakhalin. Assigning of the population from Kunashir and Iturup Island to this species requires clarification. In Mongolia it is distributed throughout northern Mongolia, in Mongol Altai, Hangai, HÃ¶vsgÃ¶l and Hentii mountain ranges (Sokolov and Orlov 1980 as P. auritus ), and Eastern Mongolia (Dorjderem 2004 as P. auritus ).	This species it not involved into trade and have no commercial value. No evidences known for using of this species in local cuisine or local medicine anywhere within its range. It plays certain role as a natural pest controller, but just minor in connection to commercial agriculture.	Terrestrial	This bat requires no special conservations actions besides conservation activities common for all bats of the temperate zone. Its range does include a few protected areas in Russia, Mongolia and Korea. This species is protected as rare in few eastern regions of Russia (as P. auritus , S. Kruskop, pers comm., August 2018). Taxonomic status of the Kunashir population should be studied with advanced molecular approach.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Plecotus		ognevi	Kishida	1927	0	Zoological Magazine of Tokyo	45:58:00	Ognev&apos;s Long-eared Bat	 uenoi Imaizumi and Yoshiyuki, 1969.	no type locality as no holotype is designated	NE China, SE Russia and E Mongolia through SC Russia; Sakhalin and Itutrup Isls; extreme W Mongolia.	Not listed.	Least Concern	 sacrimontis species group. Distinct from auritus and  includes uenoi; see Strelkov (1988) and Spitzenberger et al. (2006). This name is considered a nomen nudum by Kaneko and Maeda (2002).	Plecotus ognevi	1005671	23	Ognev's Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PLECOTINI	Plecotus	NA	ognevi	Kishida	1927	0		Kishida, K. (1927). Review of Ognev's â€˜A synopsis of the Russian bats' with notes on some species. Zoological Magazine of Tokyo, 36, 138â€“193 (in Japanese).		NA		Sakhalin, Russia.			ognevi Kishida, 1927|uenoi Imaizumi & Yoshiyuki, 1969|noemroegi Dolch, M. Stubbe, GÃ¤rtner, Thiele, Ariunbold, Batsaikhan, Lkhagvasuren, A. Stubbe, & Steinhauser, 2021	split from P. auritus	Spitzenberger, F., Strelkov, P. P., Winkler, H., & Haring, E. (2006). A preliminary revision of the genus Plecotus (Chiroptera, Vespertilionidae) based on genetic and morphological results. Zoologica Scripta, 35(3), 187-230.				Kazakhstan|Mongolia|Russia|China|North Korea|South Korea	Asia	Palearctic	LC	0	0	0	Plecotus_ognevi	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Plecotus_ognevi	1005671	23	Ognev's Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Plecotini	Plecotus	NA	ognevi	Kishida	0	Plecotus auritus ognevi	Kishida, K. 1927. Ognev, S. O.â€”May, 1927.â€”A synopsis of the Russian Bats.â€”Journ. Mammal., Vol. 8, No. 2, pp. 140â€“157. Zoological Magazine, Tokyo 39:417-418.	https://hdl.handle.net/2027/uc1.31210014193435?urlappend=%3Bseq=464%3Bownerid=117225338-488				Sakhalin, Russia.			split from P. auritus	Spitzenberger, F., Strelkov, P. P., Winkler, H., & Haring, E. (2006). A preliminary revision of the genus Plecotus (Chiroptera, Vespertilionidae) based on genetic and morphological results. Zoologica Scripta, 35(3), 187-230.				Kazakhstan|Mongolia|Russia|China|North Korea|South Korea	Asia	Palearctic	LC	0	0	0	Plecotus_ognevi	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Plecotus		ognevi	Kishida	1927	0	Zoological Magazine of Tokyo	45:58:00	Ognev&apos;s Long-eared Bat	uenoi Imaizumi and Yoshiyuki, 1969.	no type locality as no holotype is designated	NE China, SE Russia and E Mongolia through SC Russia; Sakhalin and Itutrup Isls; extreme W Mongolia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136598/21996784/' target='_blank'>Least Concern</a>	sacrimontis species group. Distinct from auritus and includes uenoi; see Strelkov (1988) and Spitzenberger et al. (2006). This name is considered a nomen nudum by Kaneko and Maeda (2002).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Plecotus ognevi; Plecotus ognevi; Plecotus ognevi; Plecotus ognevi; Plecotus ognevi; uenoi; ognevi; uenoi; noemroegi; Oreillard d'Ognev; Ognev-Langohr; Orejudo de Ognev; Ognev's Long-eared Bat; Ognev&apos;s Long-eared Bat; P. ognevi
