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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L120	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Casinycteris campomaanensis	Casinycteris campomaanensis	Casinycteris campomaanensis	Casinycteris campomaanensis	Casinycteris campomaanensis	Casinycteris campomaanensis	Casinycteris campomaanensis	Casinycteris campomaanensis	Casinycteris campomaanensis		[HMW] Casinycteris campomaanensis Hassanin, 2014 , Cameroon , South region, “Village of Nkoélon-Mvini, Campo-Ma’an area, South Region, Cameroon , 02°23.831’N , 10°02.691 °F,, 117 m above sea level.” Casinycteris campomaanensis is larger than its sister species,; C. argynnis ,; from which it also differs in nDNA and mtDNA sequences. These species split c.1:6 million years ago. Monotypic.; [MDD2022] recently described; [IUCN] Casinycteris campomaanis was described from a new specimen collected in southwest Cameroon and replaced the Cameroon distribution of the previously recognized Scotonycteris ophidon (Hassanin 2014). It is recognized as a species by Simmons and Cirranello (2020).; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														campomaanensis	Casinycteris campomaanis was described from a new specimen collected in southwest Cameroon and replaced the Cameroon distribution of the previously recognized Scotonycteris ophidon (Hassanin 2014). It is recognized as a species by Simmons and Cirranello (2020).			campomaanensis	campomaanensis			campomaanensis Hassanin, 2014						N/A																																								NA																											03AD87FAFFDDF63389693C5BF7A2FCA2	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	83	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFDDF63389693C5BF7A2FCA2.xml	Casinycteris campomaanensis	Pteropodidae	Casinycteris	campomaanensis	Hassanin	2014	Casinyctére de Campo-Ma’an @fr | Campo-Ma'an-Kurzschnauzenflughund @de | Casinicterio de Campo-Ma @es	Casinycteris campomaanensis Hassanin, 2014 , Cameroon , South region, “Village of Nkoélon-Mvini, Campo-Ma’an area, South Region, Cameroon , 02°23.831’N , 10°02.691 °F,, 117 m above sea level.” Casinycteris campomaanensis is larger than its sister species,; C. argynnis ,; from which it also differs in nDNA and mtDNA sequences. These species split c.1:6 million years ago. Monotypic.	SW Cameroon ; possibly in Nigeria and Republic of the Congo .	Ear 22- 3 mm , hindfoot 15 mm , forearm 73- 2 mm ; weight 49 g ; no tail. The Campo-Ma’an Fruit Bat has conspicuous face markings consisting of white patch on forehead in between eyes, white posterior eyespots, white band around lips, and white cheeks. Rostrum skin and eyelids are greenish yellow, and nostrils are reddish. Head is dog-like, eyes are large and brown, and ears have anterior basal ear patches. There are no epaulettes on adult males. Dorsum is rusty brown; hairs have dark brown bases, pale middles, and rusty brown tips; and pelage is dense, soft, and woolly. Ventral side is paler, with breast, belly, and throat whitish; pelage of underparts is shorter; and flanks are brown. Claw is present on second digit. Wing membranes are brown, attaching to first toes; finger joints are yellow. Skull is comparatively robust; zygomatic width is relatively wide, and arches are abruptly expanded; braincase is rounded; rostrum is short and broad; bony palate is arched medially, with deep groove along median palatine suture; there are five thick, undivided, anteriorly arched interdental palatal ridges and seven irregular, thin serrated post-dental ridges of which last two are medially divided; and post-dental palate tapers backward, ending with large posterior spine. In occlusal view, P is smaller than incisors; C' is longer than other teeth, slightly curving backward and having indistinct vertical groove on anteromedial surface; obvious diastema is present between C' and P* upper premolars and molars are subcircular, with secondary inner cusp; C, is less developed than C!, with small secondary inner cusps; and lower cheekteeth appear bifurcated, with two distinct cusps on inner and outer surfaces.	Atlantic Equatorial coastal forest biome, with low undulating hills and plains, from sea level up to elevations of ¢. 500 m .	The Campo-Ma’an Fruit Bat is presumably frugivorous.	No information.	No information.	No information.	Classified as Data Deficient on The IUCN Red List. The Campo-Ma’an Fruit Bat is only known from a very limited number of specimens, and its conservation threats are unknown. The Campo-Ma’an Fruit Bat occurs in Campo Ma’an National Park ( Cameroon ).	Hassanin (2014, 2017) | Hassanin et al. (2015)		37. Campo-Ma’an Fruit Bat Casinycteris campomaanensis French: Casinyctére de Campo-Ma’an / German: Campo-Ma'an-Kurzschnauzenflughund / Spanish: Casinicterio de Campo-Ma Taxonomy. . Casinycteris campomaanensis Hassanin, 2014 , Cameroon , South region, “Village of Nkoélon-Mvini, Campo-Ma’an area, South Region, Cameroon , 02°23.831’N , 10°02.691 °F,, 117 m above sea level.” Casinycteris campomaanensis is larger than its sister species,; C. argynnis ,; from which it also differs in nDNA and mtDNA sequences. These species split c.1:6 million years ago. Monotypic. Distribution. SW Cameroon ; possibly in Nigeria and Republic of the Congo . Descriptive notes. Ear 22- 3 mm , hindfoot 15 mm , forearm 73- 2 mm ; weight 49 g ; no tail. The Campo-Ma’an Fruit Bat has conspicuous face markings consisting of white patch on forehead in between eyes, white posterior eyespots, white band around lips, and white cheeks. Rostrum skin and eyelids are greenish yellow, and nostrils are reddish. Head is dog-like, eyes are large and brown, and ears have anterior basal ear patches. There are no epaulettes on adult males. Dorsum is rusty brown; hairs have dark brown bases, pale middles, and rusty brown tips; and pelage is dense, soft, and woolly. Ventral side is paler, with breast, belly, and throat whitish; pelage of underparts is shorter; and flanks are brown. Claw is present on second digit. Wing membranes are brown, attaching to first toes; finger joints are yellow. Skull is comparatively robust; zygomatic width is relatively wide, and arches are abruptly expanded; braincase is rounded; rostrum is short and broad; bony palate is arched medially, with deep groove along median palatine suture; there are five thick, undivided, anteriorly arched interdental palatal ridges and seven irregular, thin serrated post-dental ridges of which last two are medially divided; and post-dental palate tapers backward, ending with large posterior spine. In occlusal view, P is smaller than incisors; C' is longer than other teeth, slightly curving backward and having indistinct vertical groove on anteromedial surface; obvious diastema is present between C' and P* upper premolars and molars are subcircular, with secondary inner cusp; C, is less developed than C!, with small secondary inner cusps; and lower cheekteeth appear bifurcated, with two distinct cusps on inner and outer surfaces. Habitat. Atlantic Equatorial coastal forest biome, with low undulating hills and plains, from sea level up to elevations of ¢. 500 m . Food and Feeding. The Campo-Ma’an Fruit Bat is presumably frugivorous. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. The Campo-Ma’an Fruit Bat is only known from a very limited number of specimens, and its conservation threats are unknown. The Campo-Ma’an Fruit Bat occurs in Campo Ma’an National Park ( Cameroon ). Bibliography. Hassanin (2014, 2017), Hassanin et al. (2015).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Casinycteris campomaanensis	Casinycteris		campomaanensis	Hassanin	2014	0	C. R. Biologie	337(2014): 137	Campo-Ma'an Fruit Bat	None.	Cameroon, South Region, Campo-Ma'an Area, Nkoelon-Mvini Village	Known only from the type locality.	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Casinycteris campomaanensis	23	Campo-Ma'an Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	ROUSETTINAE	SCOTONYCTERINI	Casinycteris	NA	campomaanensis	Hassanin	2014	0	Casinycteris_campomaanensis	Hassanin, A. (2014). Description of a new bat species of the tribe Scotonycterini (Chiroptera, Pteropodidae) from southwestern Cameroon. Comptes Rendus Biologies, 337(2), 137.	https://www.sciencedirect.com/science/article/pii/S1631069113003119	MNHN 2011-637		Cameroon, South region, "Village of Nkoélon-Mvini, Campo-Ma'an area, South Region, Cameroon, 02°23.831'N, 10°02.691'E, 117 m above sea level."	2.3972	10.045	campomaanensis Hassanin, 2014	recently described	Hassanin, A. (2014). Description of a new bat species of the tribe Scotonycterini (Chiroptera, Pteropodidae) from southwestern Cameroon. Comptes Rendus Biologies, 337(2), 134-142.	Cameroon|Nigeria?|Republic of the Congo?	Africa	Afrotropic	DD	0	0	0	Casinycteris_campomaanensis	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Casinycteris campomaanensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Casinycteris	campomaanensis	Hassanin, 2014	Casinycteris campomaanis was described from a new specimen collected in southwest Cameroon and replaced the Cameroon distribution of the previously recognized Scotonycteris ophidon (Hassanin 2014). It is recognized as a species by Simmons and Cirranello (2020).	200000000	Casinycteris campomaanensis	Data Deficient		2017	2016-06-16 00:00:00 UTC	3.1	English	This species is listed as Data Deficient because it is only known from three specimens and no other informations are available regarding its demographic trend, threats and ecology.	The two identified specimens were recorded from lowland tropical moist forest in southern Cameroon and southern Congo. Currently, there is little information currently available on the natural history of this species.	Threats to this species are currently unknown.	Nothing is known about population trend and size for Casinycteris campomaanensis .	Unknown	The species has only been recorded from the type locality: Village of Nkoélon-Mvini, Campo-Ma’an area, South Region, Cameroon. A second specimen collected at Goumina (Republic of the Congo) was originally identified as Casinycteris ophiodon (Dowsett et al . 1991), but its measurements fit well with Casinycteris campomaanensis . In addition, a third specimen collected in Nigeria may belong to this species (J. Fahr, personal communication).		Terrestrial	This species is undoubtedly present in the Campo Ma'an National Park, a protected area located near the type locality. Further studies are however needed to better understand the distribution, natural history and threats of this species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Casinycteris		campomaanensis	Hassanin	2014	0	C. R. Biologie	337(2014): 137	Campo-Ma'an Fruit Bat	None.	Cameroon, South Region, Campo-Ma'an Area, Nkoelon-Mvini Village	Known only from the type locality.	Not listed.	Data Deficient		Casinycteris campomaanensis	1004548	23	Campo-Ma'an Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	ROUSETTINAE	SCOTONYCTERINI	Casinycteris	NA	campomaanensis	Hassanin	2014	0	Casinycteris_campomaanensis	Hassanin, A. (2014). Description of a new bat species of the tribe Scotonycterini (Chiroptera, Pteropodidae) from southwestern Cameroon. Comptes Rendus Biologies, 337(2), 137.	https://www.sciencedirect.com/science/article/pii/S1631069113003119	MNHN 2011-637		Cameroon, South region, "Village of Nkoélon-Mvini, Campo-Ma'an area, South Region, Cameroon, 02°23.831'N, 10°02.691'E, 117 m above sea level."	2.397183	10.04485	campomaanensis Hassanin, 2014	recently described	Hassanin, A. (2014). Description of a new bat species of the tribe Scotonycterini (Chiroptera, Pteropodidae) from southwestern Cameroon. Comptes Rendus Biologies, 337(2), 134-142.				Cameroon|Nigeria?|Republic of the Congo?	Africa	Afrotropic	DD	0	0	0	Casinycteris_campomaanensis	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Casinycteris_campomaanensis	1004548	23	Campo-Ma'an Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Epomophorinae	Scotonycterini	Casinycteris	NA	campomaanensis	Hassanin	0	Casinycteris campomaanensis	Hassanin, A. 2014-02. Description of a new bat species of the tribe Scotonycterini (Chiroptera, Pteropodidae) from southwestern Cameroon. Comptes Rendus Biologies 337(2):134-142.	https://doi.org/10.1016/j.crvi.2013.12.006	MNHN-ZM-2011-637	holotype	http://coldb.mnhn.fr/catalognumber/mnhn/zm/2011-637	Cameroon, South region, "Village of Nkoélon-Mvini, Campo-Ma'an area, South Region, Cameroon, 02°23.831'N, 10°02.691'E, 117 m above sea level."	2.397183	10.04485	recently described	Hassanin, A. (2014). Description of a new bat species of the tribe Scotonycterini (Chiroptera, Pteropodidae) from southwestern Cameroon. Comptes Rendus Biologies, 337(2), 134-142.				Cameroon|Nigeria?|Republic of the Congo?	Africa	Afrotropic	DD	0	0	0	Casinycteris_campomaanensis	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Casinycteris		campomaanensis	Hassanin	2014	0	C. R. Biologie	337(2014): 137	Campo-Ma'an Fruit Bat	None.	Cameroon, South Region, Campo-Ma'an Area, Nkoelon-Mvini Village	Known only from the type locality.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84455300/214846046/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Casinycteris campomaanensis; Casinycteris campomaanensis; Casinycteris campomaanensis; Casinycteris campomaanensis; Casinycteris campomaanensis; campomaanensis; Casinyctére de Campo-Ma’an; Campo-Ma'an-Kurzschnauzenflughund; Casinicterio de Campo-Ma; Campo-Ma'an Fruit Bat; Campo-Ma'an Fruit Bat; C. campomaanensis