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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L118	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa	Carollia subrufa		[MSW2] Not a subspecies of castanea; see Pine (1972).; [MSW3] Not a subspecies of castanea; see Pine (1972).; [HMW] Hemiderma subrufum Hahn, 1905 , “Santa Ifigenia [= Santa Efigenia], Oaxaca , Mexico .” This species is monotypic. On following pages: 106. Manu Short-tailed Bat ( Carollia manu ); 107. Sowell's Short-tailed Bat ( Carollia sowell); 108. Silky Short-tailed Bat ( Carollia perspicillata ); 111. Behn's Big-eared Bat ( Glyphonycteris behnii ); 112. Davies's Big-eared Bat ( Glyphonycteris ( Trinycteris nicefori ); 115. Least Big-eared Bat ( Neonycteris pusilla ); 116. Hairy Little Fruit Bat ( Rhinophylla alethina ); 117 Short-tailed Bat ( Carollia brevicaudum); 109. Hernandez's Short-tailed Bat ( Carollia monohernandezi ); 110. Seba’s davies); 113. Tricolored Big-eared Bat ( Glyphonycteris sylvestris ); 114. Niceforo’s Big-eared Bat. Fischer's Little Fruit Bat ( Rhinophylla fischerae ); 118. Dwarf Little Fruit Bat ( Rhinophylla pumilio ).; [batnames2022] Not a subspecies of castanea ; see Pine (1972).; [IUCN] This is not a subspecies of castanea .; [batnames2023] Not a subspecies of castanea ; see Pine (1972).; [batnames2025_1.7] Not a subspecies of castanea; see Pine (1972).														subrufa	This is not a subspecies of castanea .			subrufa	subrufa			subrufa (W. L. Hahn, 1905)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		W Mexico – Nicaragua	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Carollia subrufa	Mexico, Oaxaca, N.W. Tapanatepec, Sta. Efigenia.	Hahn	1905	Proc. Biol. Soc. Wash., 18:247.	Distribution: Ranging from southwestern Mexico south (chiefly along the Pacific slope) to Nicaragua; an isolated record from Guyana.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Hahn's short-tailed bat	W Mexico – Nicaragua	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hahn	1905	Proc. Biol. Soc. Washington, 18:247.	Not a subspecies of castanea; see Pine (1972).	Jalisco (Mexico) to NW Nicaragua; Guyana.	Mexico, Oaxaca, NW Tapanatepec, Sta. Efigenia.		HAHN	1905	Size medium (forearm length, 37-43 mm; condylobasal length, 18-21 mm; maxillary toothrow length, 6.5-7.0 mm). Labial margin of maxillary toothrow without a distinct notch but definetely concave. Crown of anterior lower molar not extremely low. Outer lower incisors not reduced, not concealed dorsally by cingula of canines. Mandible somewhat V-shaped. Pelage short and sparse.	Distribution: Ranging from southwestern Mexico south (chiefly along the Pacific slope) to Nicaragua; an isolated record from Guyana.	No subspecies.		84	species	C. subrufa	HAHN	1905	Carollia	genus	Carollia subrufa				Size medium (forearm length, 37-43 mm; condylobasal length, 18-21 mm; maxillary toothrow length, 6.5-7.0 mm). Labial margin of maxillary toothrow without a distinct notch but definetely concave. Crown of anterior lower molar not extremely low. Outer lower incisors not reduced, not concealed dorsally by cingula of canines. Mandible somewhat V-shaped. Pelage short and sparse.	No subspecies.		3. C. subrufa (HAHN 1905).	3	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Carolliinae		Carollia subrufa	Carollia		subrufa	Hahn	y	1905		Proc. Biol. Soc. Wash.	18		247		Gray Short-tailed Bat	Mexico, NW coast of Oaxaca, 8 mi (12 km) NW Tapanatepec, Sta. Efigenia.	Jalisco (Mexico) to NW Nicaragua. A report of this species from Guyana (Koopman, 1993) appears to be in error (B. Lim, pers. comm.).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Not a subspecies of castanea; see Pine (1972).	03A687BCFF82FF8516BFF412FC03F4A3	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	535	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF82FF8516BFF412FC03F4A3.xml	Carollia subrufa	Phyllostomidae	Carollia	subrufa		1905	Carollia de Hahn @fr | Hahn-Kurzschwanzblattnase @de | Carolia de Hahn @es | Hahn's Short-tailed Bat @en	Hemiderma subrufum Hahn, 1905 , “Santa Ifigenia [= Santa Efigenia], Oaxaca , Mexico .” This species is monotypic. On following pages: 106. Manu Short-tailed Bat ( Carollia manu ); 107. Sowell's Short-tailed Bat ( Carollia sowell); 108. Silky Short-tailed Bat ( Carollia perspicillata ); 111. Behn's Big-eared Bat ( Glyphonycteris behnii ); 112. Davies's Big-eared Bat ( Glyphonycteris ( Trinycteris nicefori ); 115. Least Big-eared Bat ( Neonycteris pusilla ); 116. Hairy Little Fruit Bat ( Rhinophylla alethina ); 117 Short-tailed Bat ( Carollia brevicaudum); 109. Hernandez's Short-tailed Bat ( Carollia monohernandezi ); 110. Seba’s davies); 113. Tricolored Big-eared Bat ( Glyphonycteris sylvestris ); 114. Niceforo’s Big-eared Bat. Fischer's Little Fruit Bat ( Rhinophylla fischerae ); 118. Dwarf Little Fruit Bat ( Rhinophylla pumilio ).	Pacific slope from Jalisco (W Mexico ) to NW Costa Rica .	Head-body 56-78 mm, tail 5—15 mm , ear 12-19 mm, hindfoot 10-15 mm, forearm 34-40 mm; weight 18 g . The Gray Short-tailed Bat is intermediate in size between the Chestnut Short-tailed Bat ( C. castanea ) and Seba’s Short-tailed Bat ( C. perspicillata ). It has three dorsal bands, with short gray hair, but basal and intermediate bands are not clearly distinguished. Fur between shoulders is red-brown to dark brown. Forearms are generally hairless. Rostrum is short, and lower lip has central papillae surrounded by smaller warts in a V-shape. Skull is robust, and zygomatic arches are incomplete. Over the palate, half of the tooth row is divergent, and I, are clearly visible in the jaw.	Dry, evergreen, pine-oak primary forests and also cultivated areas and secondary woodlands from sea level to elevations of ¢. 1200 m . In Costa Rica , the Gray Shorttailed Bat has been found in lowland dry forests. In El Salvador , it has been found in hollow Ceiba pentandra ( Malvaceae ).	The Gray Short-tailed Bat is mainly frugivorous and nectarivorous, and it occasionally eats insects. It is reported to prefer fruits of Brosimum alicastrum ( Moraceae ), Cecropia peltate ( Urticaceae ), Ficus morazaniana and FE ovalis ( Moraceae ), Muntingia calabura ( Muntingiaceae ), Piper amalago and P. pseudofuligineum ( Piperaceae ), Mastichodendron sp. ( Sapotaceae ), and Vismia baccifera ( Hypericaceae ).	The Gray Short-tailed Bat seems to have a bimodal polyestrous reproductive cycle. Pregnant females were found in December—March andJuly—October. It is monotocous (one young at a time). In El Salvador , juveniles have been reported in April, September, and October, and males with scrotal testes in January, March, August, and December. In Mexico, pregnant females have been reported in May.	The Gray Short-tailed Bat becomes active at the beginning of the night. It roosts in caves, hollow trees, and human structures.	The Gray Short-tailed Bat is considered gregarious and shares roosts with Pallas’s Long-tongued Bat ( Glossophaga soricina ) and the Long-legged Bat ( Macrophyllum macrophyllum ). Gray Short-tailed Bats are not present in caves with Seba’s Short-tailed Bats.	Classified as Least Concern on The IUCN Red Lust.	Bonaccorso & Gush (1987) | Hahn (1905) | LaVal & Rodriguez-Herrera (2002) | Miller et al. (2015a) | Ortega et al. (2008) | Pine (1972) | Starrett & de la Torre (1964) | Tellez (2014b)	https://zenodo.org/record/6458821/files/figure.png	105. Gray Short-tailed Bat Carollia subrufa French: Carollia de Hahn / German: Hahn-Kurzschwanzblattnase / Spanish: Carolia de Hahn Other common names: Hahn's Short-tailed Bat Taxonomy. Hemiderma subrufum Hahn, 1905 , “Santa Ifigenia [= Santa Efigenia], Oaxaca , Mexico .” This species is monotypic. On following pages: 106. Manu Short-tailed Bat ( Carollia manu ); 107. Sowell's Short-tailed Bat ( Carollia sowell); 108. Silky Short-tailed Bat ( Carollia perspicillata ); 111. Behn's Big-eared Bat ( Glyphonycteris behnii ); 112. Davies's Big-eared Bat ( Glyphonycteris ( Trinycteris nicefori ); 115. Least Big-eared Bat ( Neonycteris pusilla ); 116. Hairy Little Fruit Bat ( Rhinophylla alethina ); 117 Short-tailed Bat ( Carollia brevicaudum); 109. Hernandez's Short-tailed Bat ( Carollia monohernandezi ); 110. Seba’s davies); 113. Tricolored Big-eared Bat ( Glyphonycteris sylvestris ); 114. Niceforo’s Big-eared Bat. Fischer's Little Fruit Bat ( Rhinophylla fischerae ); 118. Dwarf Little Fruit Bat ( Rhinophylla pumilio ). Distribution. Pacific slope from Jalisco (W Mexico ) to NW Costa Rica . Descriptive notes. Head-body 56-78 mm, tail 5—15 mm , ear 12-19 mm, hindfoot 10-15 mm, forearm 34-40 mm; weight 18 g . The Gray Short-tailed Bat is intermediate in size between the Chestnut Short-tailed Bat ( C. castanea ) and Seba’s Short-tailed Bat ( C. perspicillata ). It has three dorsal bands, with short gray hair, but basal and intermediate bands are not clearly distinguished. Fur between shoulders is red-brown to dark brown. Forearms are generally hairless. Rostrum is short, and lower lip has central papillae surrounded by smaller warts in a V-shape. Skull is robust, and zygomatic arches are incomplete. Over the palate, half of the tooth row is divergent, and I, are clearly visible in the jaw. Habitat. Dry, evergreen, pine-oak primary forests and also cultivated areas and secondary woodlands from sea level to elevations of ¢. 1200 m . In Costa Rica , the Gray Shorttailed Bat has been found in lowland dry forests. In El Salvador , it has been found in hollow Ceiba pentandra ( Malvaceae ). Food and Feeding. The Gray Short-tailed Bat is mainly frugivorous and nectarivorous, and it occasionally eats insects. It is reported to prefer fruits of Brosimum alicastrum ( Moraceae ), Cecropia peltate ( Urticaceae ), Ficus morazaniana and FE ovalis ( Moraceae ), Muntingia calabura ( Muntingiaceae ), Piper amalago and P. pseudofuligineum ( Piperaceae ), Mastichodendron sp. ( Sapotaceae ), and Vismia baccifera ( Hypericaceae ). Breeding. The Gray Short-tailed Bat seems to have a bimodal polyestrous reproductive cycle. Pregnant females were found in December—March andJuly—October. It is monotocous (one young at a time). In El Salvador , juveniles have been reported in April, September, and October, and males with scrotal testes in January, March, August, and December. In Mexico, pregnant females have been reported in May. Activity patterns. The Gray Short-tailed Bat becomes active at the beginning of the night. It roosts in caves, hollow trees, and human structures. Movements, Home range and Social organization. The Gray Short-tailed Bat is considered gregarious and shares roosts with Pallas’s Long-tongued Bat ( Glossophaga soricina ) and the Long-legged Bat ( Macrophyllum macrophyllum ). Gray Short-tailed Bats are not present in caves with Seba’s Short-tailed Bats. Status and Conservation. Classified as Least Concern on The IUCN Red Lust. Bibliography. Bonaccorso & Gush (1987), Hahn (1905), LaVal & Rodriguez-Herrera (2002), Miller et al. (2015a), Ortega et al. (2008), Pine (1972), Starrett & de la Torre (1964), Tellez (2014b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Carollia subrufa	Carollia		subrufa	Hahn	1905	1	Proc. Biol. Soc. Wash.	0.9215	Gray Short-tailed Bat	None.	Mexico, NW coast of Oaxaca, 8 mi (12 km) NW Tapanatepec, Sta. Efigenia.	Jalisco (Mexico) to NW Nicaragua. A report of this species from Guyana (Koopman, 1993) appears to be in error (B. Lim, pers. comm.).	Not listed.	Least Concern	Not a subspecies of castanea ; see Pine (1972).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Carollia subrufa	23	Gray Short-tailed Bat	Hahn's Short-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	CAROLLIINAE	NA	Carollia	NA	subrufa	Hahn	1905	1	Hemiderma_subrufum	Hahn, W. L. (1905). A New Bat from Mexico. Proceedings of the Biological Society of Washington, 18, 247.	https://repository.si.edu/bitstream/handle/10088/68345/1905%20Proc%20Bio%20V%2018%20Hahn%20Bat%20pp%20247-248%20pdf.pdf?sequence=1&isAllowed=y	USNM 75127		"Santa Ifigenia [= Santa Efigenia], Oaxaca, Mexico."			subrufa (Hahn, 1905)	NA	NA	Mexico|Guatemala|El Salvador|Honduras|Nicaragua|Costa Rica	North America	Nearctic|Neotropic	LC	0	0	0	Carollia_subrufa	0	sciname match	Carollia_subrufa	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	3906	Carollia subrufa	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Carollia	subrufa	(Hahn, 1905)	This is not a subspecies of castanea .	20000000	Carollia subrufa	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern because of its wide distribution, presumed large population, occurrence in a number of protected areas, tolerance to some degree of habitat modification, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	This species can be found in dry, deciduous forest and second growth woodland (Reid 1997). It roosts in caves, empty wells, culverts, hollow trees and buildings (Reid 1997). This bat feeds on fruits of Piper , Cecropia , Muntingia ;and Solanum spp. (Fleming 1988). Reproductive cycle appears to be bimodal, with pregnant females from December to May and July to October (LaVal and Rodriguez-H 2002).	In Mexico, there is no major habitat loss (Arroyo-Cabrales pers. comm.). There are no widespread threats to this species.	This bat is common to abundant in dry forest, and absent in rainforest (Reid 1997). ;This species is really difficult to identify and may be confused with other species of ;Carollia .	Stable	This species is known from Jalisco (Mexico) to northwestern Nicaragua, and in Costa Rica. A report of this species from Guyana (Koopman 1993) appears to be in error (Lim pers. comm., Simmons 2005). It occurs from lowlands to 1,200 m asl (Reid 1997).	This species is not used.	Terrestrial	It is found in protected areas. Research actions are recommended.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Carollia		subrufa	Hahn	1905	1	Proc. Biol. Soc. Wash.	0.921528	Gray Short-tailed Bat	None.	Mexico, NW coast of Oaxaca, 8 mi (12 km) NW Tapanatepec, Sta. Efigenia.	Jalisco (Mexico) to NW Nicaragua. A report of this species from Guyana (Koopman, 1993) appears to be in error (B. Lim, pers. comm.).	Not listed.	Least Concern	Not a subspecies of castanea ; see Pine (1972).	Carollia subrufa	1004879	23	Gray Short-tailed Bat	Hahn's Short-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	CAROLLIINAE	NA	Carollia	NA	subrufa	Hahn	1905	1	Hemiderma_subrufum	Hahn, W. L. (1905). A New Bat from Mexico. Proceedings of the Biological Society of Washington, 18, 247.	https://repository.si.edu/bitstream/handle/10088/68345/1905%20Proc%20Bio%20V%2018%20Hahn%20Bat%20pp%20247-248%20pdf.pdf?sequence=1&isAllowed=y	USNM 75127		"Santa Ifigenia [= Santa Efigenia], Oaxaca, Mexico."			subrufa (Hahn, 1905)	NA	NA				Mexico|Guatemala|El Salvador|Honduras|Nicaragua|Costa Rica	North America	Nearctic|Neotropic	LC	0	0	0	Carollia_subrufa	0	sciname match	Carollia_subrufa	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Carollia_subrufa	1004879	23	Gray Short-tailed Bat	Hahn's Short-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Carolliinae	NA	Carollia	NA	subrufa	W. L. Hahn	1	Hemiderma subrufum	Hahn, W.L. 1905-12-09. A new bat from Mexico. Proceedings of the Biological Society of Washington 18:247-248.	https://www.biodiversitylibrary.org/page/3336845	USNM:MAMM:75127	holotype	http://n2t.net/ark:/65665/3ffd1c186-627a-4cfa-bd59-73d2bdfca3d3	"Santa Ifigenia [= Santa Efigenia], Oaxaca, Mexico."			NA	NA				Mexico|Guatemala|El Salvador|Honduras|Nicaragua|Costa Rica	North America	Nearctic|Neotropic	LC	0	0	0	Carollia_subrufa	0	sciname match	Carollia_subrufa	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Carollia		subrufa	Hahn	1905	1	Proc. Biol. Soc. Wash.	0.921528	Gray Short-tailed Bat	None.	Mexico, NW coast of Oaxaca, 8 mi (12 km) NW Tapanatepec, Sta. Efigenia.	Jalisco (Mexico) to NW Nicaragua. A report of this species from Guyana (Koopman, 1993) appears to be in error (B. Lim, pers. comm.).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/3906/22133926/' target='_blank'>Least Concern</a>	Not a subspecies of castanea; see Pine (1972).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Carollia subrufa; Carollia subrufa; Carollia subrufa; Carollia subrufa; Carollia subrufa; Carollia subrufa; subrufa; Carollia de Hahn; Hahn-Kurzschwanzblattnase; Carolia de Hahn; Hahn's Short-tailed Bat; Gray Short-tailed Bat; Hahn's Short-tailed Bat; Gray Short-tailed Bat; Gray Short-tailed Bat; Hemiderma subrufum; C. subrufa