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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1170	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Pipistrellus raceyi	Pipistrellus raceyi	Pipistrellus raceyi	Pipistrellus raceyi	Pipistrellus raceyi	Pipistrellus raceyi	Pipistrellus raceyi	Pipistrellus raceyi	Pipistrellus raceyi		[HMW] Pipistrellus raceyi Bates et al., 2006 , “Kianjavato, Province de Fianarantsoa , Madagascar , 21°22.84°S , 47°51.96’E , 75 m asl.” Relationship of P. raceyi to other species of Pipisitrellus is uncertain because it seems to be highly divergent from other species. As currently defined, western and eastern populations of P. racey: might represent completely distinct species or subspecies based on limited genetic and morphological data. Monotypic.; [MDD2022] recently described; [IUCN] A recently described endemic species (Bates et al. 2006).; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														raceyi	A recently described endemic species (Bates et al. 2006).			raceyi 	raceyi 			raceyi P. J. J. Bates, F. H. Ratrimomanarivo, D. L. Harrison, & S. M. Goodman, 2006						N/A																																								NA																											4C3D87E8FFEC6A53FF909D9D192BB236	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	777	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFE86A57FF869F081428BEDD.xml	Pipistrellus raceyi	Vespertilionidae	Pipistrellus	raceyi	Bates et al.	2006	Pipistrelle de Racey @fr | Racey-Zwergfledermaus @de | Pipistrela de Racey @es | Racey's Pipistrelle Bat @en	Pipistrellus raceyi Bates et al., 2006 , “Kianjavato, Province de Fianarantsoa , Madagascar , 21°22.84°S , 47°51.96’E , 75 m asl.” Relationship of P. raceyi to other species of Pipisitrellus is uncertain because it seems to be highly divergent from other species. As currently defined, western and eastern populations of P. racey: might represent completely distinct species or subspecies based on limited genetic and morphological data. Monotypic.	Lowlands of W & E coasts of Madagascar .	Head-body 46-52 mm,tail 229-34 mm, ear 7-5-11 mm, hindfoot 5-3-7-5 mm, forearm 27-33 mm; weight 3-8-5-8 g. Eastern population has generally larger individuals than western population. Pelage is relatively long. Dorsum of Racey’s Pipistrelle is light rufous, with distinctly darker head; venter is buffy brown. Wing membranes are dark brownish black, and uropatagium is a similar color and stretches to about end oftail. Muzzle is relatively short and inflated, with large glandular swellings that are essentially naked; face is pinkish and darker blackish brown closer to front of muzzle. Ears are uniformly dark brownish black and short, with rounded tips and 3-5 well-developed ridges; tragus is ¢.50% the height of ear, crescent-shaped, and has small notch near base of posterior border and rounded tip. Penis is very long (9-6-11-8 mm; much longer than the Dusky Pipistrelle, P. hesperidus , in Madagascar ). Baculum is long and thin, curving upward near base and downward near tip; tip is always bifurcated, and base is bifurcated in western population and not in eastern population (only with very slight medial notch); base is either relatively skinny in western population but inflated dorsally in eastern population. Rostrum is rounded and flattened, with slightly upturned tip; supraorbital ridges are inflated and angular, forming an oblique welldefined hollow in front of each ridge; I® is about the height of second cusp of I*; I? is bicuspid; I” is unicuspid; P* is comparatively well developed and slightly displaced from tooth row; C! and P* never touch; and lower molars are nyctalodont.	Degraded and relatively intact deciduous forests in western Madagascar and near disturbed lowland and humid forests and near agriculture in eastern Madagascar , recorded at elevations of 10-300 m.	Racey’s Pipistrelle is insectivorous. According to stomach samples, diet consisted of Isoptera (30% volume; 37-8% frequency), Coleoptera (27-8%; 789%), Lepidoptera (26-1%; 65-6%), Hymenoptera (125%; 47-8%), Trichoptera (1-9%; 10%), Homoptera (1-5%; 12-2%), and Arachnida (0-1%; 17%). Fecal samples of three individuals included Coleoptera (32:5% volume), Diptera (28-9%), Lepidoptera (16-:3%), Homoptera (117%), Araneae (6:8%), and Hymenoptera (3-9%).	Births occur at beginning of rainy season (November-December).	Racey’s Pipistrelle is nocturnal. It roosts in houses, other man-made structures, and vegetation. Call shape is FM/QCEF, and recordings of four individuals in Madagascar (three western and one eastern) had peak frequencies of 53-4-58-2 kHz, maximum frequencies of 91-:7-128-3 kHz, minimum frequencies of 41-7-51-4 kHz, and durations of 2-3-3-1 milliseconds.	Racey’s Pipistrelle roosted in the same building with the Malagasy Serotine ( Neoromicia matroka ) and the Malagasy Myotis ( Myotis goudotii ) in Ambalahosy, eastern Madagascar , but each was found in different positions between ceilingjoints and roofing.	Classified as Data Deficient on The IUCN Red List. Preliminary evidence suggests that Racey’s Pipistrelle is common in lowland regions in Madagascar , but information is very limited on its ecology and threats.	ACR (2018) | Bates et al. (2006) | Goodman, Rakotondramanana etal. (2015) | Goodman, Rasoanoro et al. (2014) | Jenkins et al. (2008) | Rakotondramanana et al. (2015) | Rasoanoro et al. (2015) | Wilkinson et al. (2012)	https://zenodo.org/record/6397836/files/figure.png	31. Racey’s Pipistrelle Pipistrellus raceyi French: Pipistrelle de Racey / German: Racey-Zwergfledermaus / Spanish: Pipistrela de Racey Other common names: Racey's Pipistrelle Bat Taxonomy. Pipistrellus raceyi Bates et al., 2006 , “Kianjavato, Province de Fianarantsoa , Madagascar , 21°22.84°S , 47°51.96’E , 75 m asl.” Relationship of P. raceyi to other species of Pipisitrellus is uncertain because it seems to be highly divergent from other species. As currently defined, western and eastern populations of P. racey: might represent completely distinct species or subspecies based on limited genetic and morphological data. Monotypic. Distribution. Lowlands of W & E coasts of Madagascar . Descriptive notes. Head-body 46-52 mm,tail 229-34 mm, ear 7-5-11 mm, hindfoot 5-3-7-5 mm, forearm 27-33 mm; weight 3-8-5-8 g. Eastern population has generally larger individuals than western population. Pelage is relatively long. Dorsum of Racey’s Pipistrelle is light rufous, with distinctly darker head; venter is buffy brown. Wing membranes are dark brownish black, and uropatagium is a similar color and stretches to about end oftail. Muzzle is relatively short and inflated, with large glandular swellings that are essentially naked; face is pinkish and darker blackish brown closer to front of muzzle. Ears are uniformly dark brownish black and short, with rounded tips and 3-5 well-developed ridges; tragus is ¢.50% the height of ear, crescent-shaped, and has small notch near base of posterior border and rounded tip. Penis is very long (9-6-11-8 mm; much longer than the Dusky Pipistrelle, P. hesperidus , in Madagascar ). Baculum is long and thin, curving upward near base and downward near tip; tip is always bifurcated, and base is bifurcated in western population and not in eastern population (only with very slight medial notch); base is either relatively skinny in western population but inflated dorsally in eastern population. Rostrum is rounded and flattened, with slightly upturned tip; supraorbital ridges are inflated and angular, forming an oblique welldefined hollow in front of each ridge; I® is about the height of second cusp of I*; I? is bicuspid; I” is unicuspid; P* is comparatively well developed and slightly displaced from tooth row; C! and P* never touch; and lower molars are nyctalodont. Habitat. Degraded and relatively intact deciduous forests in western Madagascar and near disturbed lowland and humid forests and near agriculture in eastern Madagascar , recorded at elevations of 10-300 m. Food and Feeding. Racey’s Pipistrelle is insectivorous. According to stomach samples, diet consisted of Isoptera (30% volume; 37-8% frequency), Coleoptera (27-8%; 789%), Lepidoptera (26-1%; 65-6%), Hymenoptera (125%; 47-8%), Trichoptera (1-9%; 10%), Homoptera (1-5%; 12-2%), and Arachnida (0-1%; 17%). Fecal samples of three individuals included Coleoptera (32:5% volume), Diptera (28-9%), Lepidoptera (16-:3%), Homoptera (117%), Araneae (6:8%), and Hymenoptera (3-9%). Breeding. Births occur at beginning of rainy season (November-December). Activity patterns. Racey’s Pipistrelle is nocturnal. It roosts in houses, other man-made structures, and vegetation. Call shape is FM/QCEF, and recordings of four individuals in Madagascar (three western and one eastern) had peak frequencies of 53-4-58-2 kHz, maximum frequencies of 91-:7-128-3 kHz, minimum frequencies of 41-7-51-4 kHz, and durations of 2-3-3-1 milliseconds. Movements, Home range and Social organization. Racey’s Pipistrelle roosted in the same building with the Malagasy Serotine ( Neoromicia matroka ) and the Malagasy Myotis ( Myotis goudotii ) in Ambalahosy, eastern Madagascar , but each was found in different positions between ceilingjoints and roofing. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Preliminary evidence suggests that Racey’s Pipistrelle is common in lowland regions in Madagascar , but information is very limited on its ecology and threats. Bibliography. ACR (2018), Bates et al. (2006), Goodman, Rakotondramanana etal. (2015), Goodman, Rasoanoro et al. (2014), Jenkins et al. (2008), Rakotondramanana et al. (2015), Rasoanoro et al. (2015), Wilkinson et al. (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pipistrellus raceyi	Pipistrellus	Pipistrellus	raceyi	Bates, Ratrimomanarivo, Harrison, & Goodman	2006	0	Acta Chiropterol.	8(2): 301	Racey&apos;s Pipistelle	None.	Madagascar, Province de Fianarantsoa, Kianjavato	Madagascar.	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pipistrellus raceyi	23	Racey's Pipistrelle	Racey's Pipistrelle Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	raceyi	P. J. J. Bates, Ratrimomanarivo, D. L. Harrison, & Goodman	2006	0	Pipistrellus_raceyi	Bates, P. J., Ratrimomanarivo, F. H., Harrison, D. L., & Goodman, S. M. (2006). A description of a new species of Pipistrellus (Chiroptera: Vespertilionidae) from Madagascar with a review of related Vespertilioninae from the island. Acta Chiropterologica, 8(2), 301.	https://bioone.org/journals/acta-chiropterologica/volume-8/issue-2/1733-5329(2006)8[299:ADOANS]2.0.CO;2/A-description-of-a-new-species-of-Pipistrellus-Chiroptera/10.3161/1733-5329(2006)8[299:ADOANS]2.0.CO;2.full	FMNH 185567		"Kianjavato, Province de Fianarantsoa, Madagascar, 21°22.84'S, 47°51.96'E, 75 m a.s.l."	-21.38	47.866	raceyi P. J. J. Bates, Ratrimomanarivo, D. L. Harrison, & Goodman, 2006	recently described	Bates, P. J., Ratrimomanarivo, F. H., Harrison, D. L., & Goodman, S. M. (2006). A description of a new species of Pipistrellus (Chiroptera: Vespertilionidae) from Madagascar with a review of related Vespertilioninae from the island. Acta Chiropterologica, 8(2), 299-324.	Madagascar	Africa	Afrotropic	DD	0	0	0	Pipistrellus_raceyi	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136646	Pipistrellus raceyi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pipistrellus	raceyi	Bates, Ratrimomanarivo, Harrison &; Goodman, 2006	A recently described endemic species (Bates et al. 2006).	20000000	Pipistrellus raceyi	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	This species is listed as Data Deficient because although preliminary evidence suggests that it is widely distributed in lowland areas of parts of eastern and western Madagascar and it exists in a range of vegetation types, there are insufficient data on its habitat requirements and major threats.	Individuals from the east have been netted near to disturbed lowland and humid forest but also in proximity to agriculture and there is no evidence that this species requires humid forest vegetation to survive (Bates et al. 2006). In the west, P. raceyi was netted inside degraded and relatively intact dry deciduous forest (Bates et al . 2006). In the east it is known to roost inside buildings (Bates et al . 2006). Nevertheless, all four localities are noteworthy for the presence of lowland forest formations and additional surveys are now needed to determine whether this is a genuine reflection of habitat requirements or an artifact of how biologists select their survey sites. If it is the latter case, then P. raceyi may turn out to be a common species in the low-lying areas across much of Madagascar.	No major threats are documented, but forest degradation may conceivably reduce availability of suitable foraging and roosting habitat in the west where this species presumably roosts in trees and appears to be associated with forest habitats.	This recently described species is currently known only from 15 specimens (Bates et al. 2006); however, with further survey work, it may be shown to more common.	Unknown	This is an endemic species to Madagascar that is currently known from four localities (Bates et al. 2006), two in the west (Mikea forest and Kirindy CFPF) and two in the east (Kianjavato and Tampolo forests), all below 80 m elevation (from 16 - 70 m asl.).		Terrestrial	Even though this species roosts synanthropically in the east and may therefore be expected to be relatively common given the abundance of suitable roosting sites, it is only known from four locations across Madagascar despite the large survey effort on Malagasy bats in recent years (Goodman et al. 2005, Bates et al . 2006, Cardiff 2006, Rakotonandrasana and Goodman 2007). Of the four known localities, all receive some form of active forest management and it is expected that deforestation at these sites has been and will be less than for non-protected forest. Further studies of this species are needed to investigate why it appears to be forest-dependent in the west but not in the east of its range.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pipistrellus	Pipistrellus	raceyi	Bates, Ratrimomanarivo, Harrison, & Goodman	2006	0	Acta Chiropterol.	8(2): 301	Racey&apos;s Pipistelle	None.	Madagascar, Province de Fianarantsoa, Kianjavato	Madagascar.	Not listed.	Data Deficient		Pipistrellus raceyi	1005634	23	Racey's Pipistrelle	Racey's Pipistrelle Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	raceyi	P. J. J. Bates, Ratrimomanarivo, D. L. Harrison, & Goodman	2006	0	Pipistrellus_raceyi	Bates, P. J., Ratrimomanarivo, F. H., Harrison, D. L., & Goodman, S. M. (2006). A description of a new species of Pipistrellus (Chiroptera: Vespertilionidae) from Madagascar with a review of related Vespertilioninae from the island. Acta Chiropterologica, 8(2), 301.	https://bioone.org/journals/acta-chiropterologica/volume-8/issue-2/1733-5329(2006)8[299:ADOANS]2.0.CO;2/A-description-of-a-new-species-of-Pipistrellus-Chiroptera/10.3161/1733-5329(2006)8[299:ADOANS]2.0.CO;2.full	FMNH 185567		"Kianjavato, Province de Fianarantsoa, Madagascar, 21°22.84'S, 47°51.96'E, 75 m a.s.l."	-21.3807	47.866	raceyi P. J. J. Bates, Ratrimomanarivo, D. L. Harrison, & Goodman, 2006	recently described	Bates, P. J., Ratrimomanarivo, F. H., Harrison, D. L., & Goodman, S. M. (2006). A description of a new species of Pipistrellus (Chiroptera: Vespertilionidae) from Madagascar with a review of related Vespertilioninae from the island. Acta Chiropterologica, 8(2), 299-324.				Madagascar	Africa	Afrotropic	DD	0	0	0	Pipistrellus_raceyi	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pipistrellus_raceyi	1005634	23	Racey's Pipistrelle	Racey's Pipistrelle Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus	Pipistrellus	raceyi	P. J. J. Bates, F. H. Ratrimomanarivo, D. L. Harrison, & S. M. Goodman	0	Pipistrellus raceyi	Bates, P.J.J., Ratrimomanarivo, F.H., Harrison, D.L. and Goodman, S.M. 2006-12. A description of a new species of _Pipistrellus_ (Chiroptera: Vespertilionidae) from Madagascar with a review of related Vespertilioninae from the island. Acta Chiropterologica 8(2):299-324.	https://doi.org/10.3161/1733-5329(2006)8[299:ADOANS]2.0.CO;2	FMNH:Mamm:185567	holotype	https://collections-zoology.fieldmuseum.org/catalogue/2684335	"Kianjavato, Province de Fianarantsoa, Madagascar, 21°22.84'S, 47°51.96'E, 75 m a.s.l."	-21.3807	47.866	recently described	Bates, P. J., Ratrimomanarivo, F. H., Harrison, D. L., & Goodman, S. M. (2006). A description of a new species of Pipistrellus (Chiroptera: Vespertilionidae) from Madagascar with a review of related Vespertilioninae from the island. Acta Chiropterologica, 8(2), 299-324.				Madagascar	Africa	Afrotropic	DD	0	0	0	Pipistrellus_raceyi	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pipistrellus	Pipistrellus	raceyi	Bates, Ratrimomanarivo, Harrison,  & Goodman	2006	0	Acta Chiropterol.	8(2): 301	Racey&apos;s Pipistelle	None.	Madagascar, Province de Fianarantsoa, Kianjavato	Madagascar.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136646/21989768/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pipistrellus raceyi; Pipistrellus raceyi; Pipistrellus raceyi; Pipistrellus raceyi; Pipistrellus raceyi; raceyi; Pipistrelle de Racey; Racey-Zwergfledermaus; Pipistrela de Racey; Racey's Pipistrelle Bat; Racey's Pipistrelle; Racey's Pipistrelle Bat; Racey&apos;s Pipistelle; P. raceyi