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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1164	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii	Pipistrellus nathusii		[MSW2] Subgenus Pipistrellus.; [MSW3] Subgenus Pipistrellus.; [HMW] Vesperugo nathusii Keyserling & Blasius, 1839 , Berlin , Germany . Pipastrellus nathusii appears to be close to the P. kuhlii complex, which includes paraphyletic P. kuhliiand P. maderensis . Monotypic.; [batnames2022] Subgenus Pipistrellus .; [batnames2023] Subgenus Pipistrellus .; [batnames2025_1.7] Subgenus Pipistrellus.						unicolor.			unicolor			nathusii 	nathusii - unicolor	nathusii, unicolor		nathusii 	nathusii - unicolor	nathusii, unicolor 	nathusii, unicolor, nathusiusi	 nathusii	nathusii - unicolor	nathusii (von Keyserling & J. H. Blasius, 1839)|unicolor (Fatio, 1905)|nathusiusi Bianchi, 1918 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Nathusius' pipistrelle	Spain – Urals, Caucasus	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pipistrellus nathusii	Germany, Berlin.	Keyserling and Blasius	1839	Arch. Naturgesch., 5(1 ):320.	Distribution: Confined to Europe, Asia Minor, and Trans-Caucasia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Nathusius' pipistrelle	Spain – Urals, Caucasus	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Keyserling and Blasius	1839	Arch. Naturgesch., 5(1):320.	Subgenus Pipistrellus.	W Europe to Urals and Caucasus, and W Asia Minor; S England.	Germany, Berlin.		KEYSERLING & BLASIUS	1839	Size fairly small (forearm length, 30-37 mm). Outer upper incisor relatively long. Anterior upper premolar in toothrow. Rostrum relatively long.	Distribution: Confined to Europe, Asia Minor, and Trans-Caucasia.	No subspecies.		112	species	P. nathusii	KEYSERLING & BLASIUS	1839	Pipistrellus	subgenus	Pipistrellus nathusii				Size fairly small (forearm length, 30-37 mm). Outer upper incisor relatively long. Anterior upper premolar in toothrow. Rostrum relatively long.	No subspecies.		3. P. nathusii (KEYSERLING & BLASIUS 1839) [pipistrellus group].	3	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus nathusii	Pipistrellus	Pipistrellus	nathusii	Keyserling and Blasius	y	1839		Arch. Naturgesch.	5	1	320		Nathusius's Pipistrelle	Germany, Berlin.	W Europe to Urals and Caucasus, and W Asia Minor; S England.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	unicolor Fatio, 1905.	Subgenus Pipistrellus.	4C3D87E8FFE86A57FF8494391FE9B23A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	773	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFF46A4BFF8093C617E2BCE3.xml	Pipistrellus nathusii	Vespertilionidae	Pipistrellus	nathusii	Keyserling & Blasius	1839	Pipistrelle de Nathusius @fr | Rauhautfledermaus @de | Pipistrela de Nathusius @es	Vesperugo nathusii Keyserling & Blasius, 1839 , Berlin , Germany . Pipastrellus nathusii appears to be close to the P. kuhlii complex, which includes paraphyletic P. kuhliiand P. maderensis . Monotypic.	Most of Europe (from Ireland and Iberian Peninsula E to Russia and W Kazakhstan ), Turkey , and Caucasus; also on Mediterranean Is such as Crete or Mallorca.	Head-body 44-54 mm, tail 30-40 mm, ear 10-5-16 mm, forearm 32-2-37-1 mm; weight 6-15-5 g. Fur color of Nathusius’s Pipistrelle changes seasonally, being dark brown or gray brown in winter and reddish in summer. Fur is long and uniformly colored, but venteris slightly paler or more yellowish than dorsum. Proximal one-half of uropatagium is covered by fur. Bare skin is dark brown, snout is short and rounded, and ears are triangular and short, with slender and rounded tragus. Compared with other pipistrelle species, wings of Nathusius’s Pipistrelle are quite long (wingspan 230-250 mm), perfectly adapted for long flights and its migratory behavior. Although not always reliable, a particular wing venation pattern is used as a diagnostic character (cell from elbow to fifth finger clear ly divided). I* is bicuspid, I’ is longer than second cusp of I, canines are considerably large, P? is small but visible above gum, and lower molars are nyctalodont. Chromosomal complement has 2n = 44, FNa = 50, and FN = 54.	[Lowland to alpine habitats from sea level up to elevations of ¢. 2200 m in the Alps (probably single vagrant individuals). Habitats selected by Nathusius’s Pipistrelle include woodlands, forest edges, wetlands, open areas, and grasslands. In terms offorests, it occurs in deciduous, lowland, riparian, and coniferous forests and urban gardens with isolated trees. It tends to select those habitats surrounding ponds, lakes, or water bodies; thus,it is common in wetlands, swamps, and floodplains, especially during migration.	Nathusius’s Pipistrelle reportedly eats Diptera and Lepidoptera , especially non-biting midges, mosquitoes, and black flies. Despite being slower than its congeners, it hunts insects (aerial hawker) at 3-20 m aboveground, commonly on forest edges, linear structures, and forest paths and over water bodies (from small ponds to large lakes), and occasionally around streetlamps. It has also been recorded foraging over sea up to 1-3 km from the coast. Dietary composition varies between maternity, swarming, and migration periods. In Latvia , during migration, Nathusius’s Pipistrelles seem to use a “fly-and-forage” feeding strategy, which consists in stopping several times during migration to feed and then continue the flight.	Mating of Nathusius’s Pipistrelles occurs in June-November, including when they roost in nursery colonies (June-August), during migration (August-November), and at their destination at swarming and hibernation sites (November). Males can be resident in swarming areas (which they defend from other males), visitors to specific mating locations, or sporadic visitors that try to mate opportunistically. Maternity colonies are only found in northern and Eastern Europe (Britain, eastern Germany , Baltic States, Belarus , Ukraine , and Russia ). During breeding season, Nathusius’s Pipistrelles congregate in small colonies of some tens to a few hundred individuals, roosting in tree holes, bat boxes, and buildings, usually in lowland forests and wetlands. They tend to give birth to twins in June and leave colonies by the end ofJuly. Young females become sexually active during their first year of life. Nathusius’s Pipistrelles can live up to 13 years.	Nathusius’s Pipistrelle emerges at dusk to hunt flying insects. Unlike other pipistrelles, its flight is not that erratic, being straighter and more predictable. It roosts in hollow trees, bird and bat boxes, rock crevices, expansion joints in bridges, houses, and a wide range of different buildings, timber claddings, or wooden churches. In winter,it tends to roost in rock crevices, cliffs, caves, mines, and buildings in urban areas (most likely benefiting from urban heat islands). It hibernates from mid-winter to early spring. Its echolocation is very similar to that of Kuhl’s Pipistrelle and has high FM sweeps with CF cue that resembles a hockey stick-shaped call when represented on sonograms. Peak frequencies are ¢.38—43 kHz. Echolocation calls of Nathusius’s Pipistrelle can be confused with those of Kuhl’s Pipistrelle (PF. kuhliz) and Savi’s Pipistrelle ( Hypsugo savii ). Calls are variable; in open spaces, FM component can disappear or be lower in frequency, making identification difficult; and in cluttered environments, pulses become highly modulated resembling Myotis echolocation. Pulses last up to 12 milliseconds. Socials calls are very distinct from those of Kuhl’s Pipistrelle and are typically used to reliably identify recorded free-flying bat calls.	Nathusius’s Pipistrelle is one of the few long-distance migratory bat species in Europe, for which migration patterns of more than 1900 km have been reported. In Europe, it tends to breed in north-eastern countries in summer and migrate toward south-western countries to hibernate. In these regions,it positively selects natural wetlands for foraging and avoids agricultural lands and pastures. During migration, it presumably follows coastlines (it has been captured or recorded in offshore platforms and fishing boats), but it has also been captured over high mountains or following waterways. It has also been recorded migrating from continental Europe to Great Britain . Females start migration before males. Migration extends from August to October or November. Nathusius’s Pipistrelles can fly up to 29-48 km/night and only occasionally up to 80 km /night. During the maternity period, foraging areas are relatively close to maternity colonies (less than 6-5 km). Maternity colonies usually share space with the Common Pipistrelle, Brandt's Myotis ( Myotis brandtii ), and the Pond Myotis ( M. dasycneme ).	Classified as Least Concern on The IUCN Red List. Although not very common, Nathusius’s Pipistrelle is quite widespread. Although it has been more common in northern countries,it is now more frequently reported in the south, and during the last few decades, it has been reported or established in the Netherlands , UK , Switzerland , Italy , and Spain . Because it is one of the few long-distance migratory bat species in Europe, modification or fragmentation of habitat and commuting routes might compromise its movements. Roost disturbance and destruction, loss of mature trees in wetlands, and direct persecution are recognized as direct threats.	Arslan & Zima (2014) | Benda, Georgiakakis et al. (2008) | Bogdanowicz (1999a) | Boshamer & Bekker (2008) | Brosset (1990) | Dietz & Kiefer (2016) | Flaguer, Puig-Montserrat et al. (2009) | Flaquer, Ruiz-Jarillo et al. (2005) | Harris &Yalden (2008) | Jahelkova et al. (2008) | Kriiger et al. (2014) | Limpens & Schulte (2000) | Lundy et al. (2010) | Pacifici et al. (2013) | Paunovi¢ & Juste (2016) | Pétersons (2004) | Rodriguez-Munoz et al. (1993) | Russ & Racey (2007) | Russ, Hutson et al. (2001) | Russ, O'Neill & Montgomery (1998) | Sachanowicz et al. (2019) | Schober & Grimmberger (1998) | Speakman et al. (1991) | Suba et al. (2012) | Trujillo & Garcia (2009) | Vierhaus (2004) | Voigt et al. (2012)	https://zenodo.org/record/6397816/files/figure.png	22. Nathusius’s Pipistrelle Pipistrellus nathusii French: Pipistrelle de Nathusius / German: Rauhautfledermaus / Spanish: Pipistrela de Nathusius Taxonomy. Vesperugo nathusii Keyserling & Blasius, 1839 , Berlin , Germany . Pipastrellus nathusii appears to be close to the P. kuhlii complex, which includes paraphyletic P. kuhliiand P. maderensis . Monotypic. Distribution. Most of Europe (from Ireland and Iberian Peninsula E to Russia and W Kazakhstan ), Turkey , and Caucasus; also on Mediterranean Is such as Crete or Mallorca. Descriptive notes. Head-body 44-54 mm, tail 30-40 mm, ear 10-5-16 mm, forearm 32-2-37-1 mm; weight 6-15-5 g. Fur color of Nathusius’s Pipistrelle changes seasonally, being dark brown or gray brown in winter and reddish in summer. Fur is long and uniformly colored, but venteris slightly paler or more yellowish than dorsum. Proximal one-half of uropatagium is covered by fur. Bare skin is dark brown, snout is short and rounded, and ears are triangular and short, with slender and rounded tragus. Compared with other pipistrelle species, wings of Nathusius’s Pipistrelle are quite long (wingspan 230-250 mm), perfectly adapted for long flights and its migratory behavior. Although not always reliable, a particular wing venation pattern is used as a diagnostic character (cell from elbow to fifth finger clear ly divided). I* is bicuspid, I’ is longer than second cusp of I, canines are considerably large, P? is small but visible above gum, and lower molars are nyctalodont. Chromosomal complement has 2n = 44, FNa = 50, and FN = 54. Habitat. [Lowland to alpine habitats from sea level up to elevations of ¢. 2200 m in the Alps (probably single vagrant individuals). Habitats selected by Nathusius’s Pipistrelle include woodlands, forest edges, wetlands, open areas, and grasslands. In terms offorests, it occurs in deciduous, lowland, riparian, and coniferous forests and urban gardens with isolated trees. It tends to select those habitats surrounding ponds, lakes, or water bodies; thus,it is common in wetlands, swamps, and floodplains, especially during migration. Food and Feeding. Nathusius’s Pipistrelle reportedly eats Diptera and Lepidoptera , especially non-biting midges, mosquitoes, and black flies. Despite being slower than its congeners, it hunts insects (aerial hawker) at 3-20 m aboveground, commonly on forest edges, linear structures, and forest paths and over water bodies (from small ponds to large lakes), and occasionally around streetlamps. It has also been recorded foraging over sea up to 1-3 km from the coast. Dietary composition varies between maternity, swarming, and migration periods. In Latvia , during migration, Nathusius’s Pipistrelles seem to use a “fly-and-forage” feeding strategy, which consists in stopping several times during migration to feed and then continue the flight. Breeding. Mating of Nathusius’s Pipistrelles occurs in June-November, including when they roost in nursery colonies (June-August), during migration (August-November), and at their destination at swarming and hibernation sites (November). Males can be resident in swarming areas (which they defend from other males), visitors to specific mating locations, or sporadic visitors that try to mate opportunistically. Maternity colonies are only found in northern and Eastern Europe (Britain, eastern Germany , Baltic States, Belarus , Ukraine , and Russia ). During breeding season, Nathusius’s Pipistrelles congregate in small colonies of some tens to a few hundred individuals, roosting in tree holes, bat boxes, and buildings, usually in lowland forests and wetlands. They tend to give birth to twins in June and leave colonies by the end ofJuly. Young females become sexually active during their first year of life. Nathusius’s Pipistrelles can live up to 13 years. Activity patterns. Nathusius’s Pipistrelle emerges at dusk to hunt flying insects. Unlike other pipistrelles, its flight is not that erratic, being straighter and more predictable. It roosts in hollow trees, bird and bat boxes, rock crevices, expansion joints in bridges, houses, and a wide range of different buildings, timber claddings, or wooden churches. In winter,it tends to roost in rock crevices, cliffs, caves, mines, and buildings in urban areas (most likely benefiting from urban heat islands). It hibernates from mid-winter to early spring. Its echolocation is very similar to that of Kuhl’s Pipistrelle and has high FM sweeps with CF cue that resembles a hockey stick-shaped call when represented on sonograms. Peak frequencies are ¢.38—43 kHz. Echolocation calls of Nathusius’s Pipistrelle can be confused with those of Kuhl’s Pipistrelle (PF. kuhliz) and Savi’s Pipistrelle ( Hypsugo savii ). Calls are variable; in open spaces, FM component can disappear or be lower in frequency, making identification difficult; and in cluttered environments, pulses become highly modulated resembling Myotis echolocation. Pulses last up to 12 milliseconds. Socials calls are very distinct from those of Kuhl’s Pipistrelle and are typically used to reliably identify recorded free-flying bat calls. Movements, Home range and Social organization. Nathusius’s Pipistrelle is one of the few long-distance migratory bat species in Europe, for which migration patterns of more than 1900 km have been reported. In Europe, it tends to breed in north-eastern countries in summer and migrate toward south-western countries to hibernate. In these regions,it positively selects natural wetlands for foraging and avoids agricultural lands and pastures. During migration, it presumably follows coastlines (it has been captured or recorded in offshore platforms and fishing boats), but it has also been captured over high mountains or following waterways. It has also been recorded migrating from continental Europe to Great Britain . Females start migration before males. Migration extends from August to October or November. Nathusius’s Pipistrelles can fly up to 29-48 km/night and only occasionally up to 80 km /night. During the maternity period, foraging areas are relatively close to maternity colonies (less than 6-5 km). Maternity colonies usually share space with the Common Pipistrelle, Brandt's Myotis ( Myotis brandtii ), and the Pond Myotis ( M. dasycneme ). Status and Conservation. Classified as Least Concern on The IUCN Red List. Although not very common, Nathusius’s Pipistrelle is quite widespread. Although it has been more common in northern countries,it is now more frequently reported in the south, and during the last few decades, it has been reported or established in the Netherlands , UK , Switzerland , Italy , and Spain . Because it is one of the few long-distance migratory bat species in Europe, modification or fragmentation of habitat and commuting routes might compromise its movements. Roost disturbance and destruction, loss of mature trees in wetlands, and direct persecution are recognized as direct threats. Bibliography. Arslan & Zima (2014), Benda, Georgiakakis et al. (2008), Bogdanowicz (1999a), Boshamer & Bekker (2008), Brosset (1990), Dietz & Kiefer (2016), Flaguer, Puig-Montserrat et al. (2009), Flaquer, Ruiz-Jarillo et al. (2005), Harris &Yalden (2008), Jahelkova et al. (2008), Kriiger et al. (2014), Limpens & Schulte (2000), Lundy et al. (2010), Pacifici et al. (2013), Paunovi¢ & Juste (2016), Pétersons (2004), Rodriguez-Munoz et al. (1993), Russ & Racey (2007), Russ, Hutson et al. (2001), Russ, O'Neill & Montgomery (1998), Sachanowicz et al. (2019), Schober & Grimmberger (1998), Speakman et al. (1991), Suba et al. (2012), Trujillo & Garcia (2009), Vierhaus (2004), Voigt et al. (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pipistrellus nathusii	Pipistrellus	Pipistrellus	nathusii	Keyserling and Blasius	1839	1	Arch. Naturgesch.	5(1): 320	Nathusius' Pipistrelle	 unicolor Fatio, 1905.	Germany, Berlin.	W Europe to Urals and Caucasus, and W Asia Minor; S England.	Not listed.	Least Concern	Subgenus Pipistrellus .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pipistrellus nathusii	23	Nathusius's Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	nathusii	Keyserling & Blasius	1839	1						Berlin, Germany.			nathusii (Keyserling & Blasius, 1839)|unicolor (Fatio, 1905)	NA	NA	Ireland|United Kingdom|Spain|France|Luxembourg|Belgium|Netherlands|Germany|Denmark|Norway|Sweden|Finland|Switzerland|Liechtenstein|Italy|Austria|Czech Republic|Poland|Slovakia|Hungary|Slovenia|Croatia|Bosnia & Herzegovina|Serbia|Kosovo|Montenegro|Albania|North Macedonia|Greece|Bulgaria|Turkey|Romania|Moldova|Ukraine|Belarus|Estonia|Latvia|Lithuania|Russia|Georgia|Armenia|Azerbaijan|Kazakhstan	Asia|Europe	Palearctic	LC	0	0	0	Pipistrellus_nathusii	0	sciname match	Pipistrellus_nathusii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17316	Pipistrellus nathusii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pipistrellus	nathusii	(Keyserling &; Blasius, 1839)		20000000	Pipistrellus nathusii	Least Concern		2016	2016-04-25 00:00:00 UTC	3.1	English	The species is widespread and abundant, and there is no evidence of current significant population decline. Consequently the species is assessed as Least Concern.	Nathusius' Pipistrelle (Pipistrellus nathusii ) ;forages over a range of habitats including woodland, edge, wetlands (among which natural ones are preferred, Flaquer et al. 2009), ;and open parkland. It feeds mainly on Diptera and Lepidoptera (Krüger ;et al. 2014). ;Summer roosts are located in tree holes, buildings, and bat boxes, mainly in woodland areas. Winter roost sites include crevices in cliffs, buildings and around the entrance of caves, often in relatively cold, dry, and exposed sites. It is a migratory species, with movements of up to 1,905 km recorded (Petersons 2004). Migrations typically follow a NE-SW route (Bogdanowicz 1999).</span>	Although not major threats, the species is affected by habitat fragmentation on migration routes, loss of and disturbance to roosts in buildings, loss of mature trees with cavities and/or loose bark, etc., and water quality changes which may affect food supply.	Nathusius' Pipistrelle ;is abundant in northern parts of range, and less common but increasingly recorded in southern and western parts of its range. Summer maternity colonies of up to 200 individuals have been recorded, but large winter aggregations are not known. Increase in higher minimum temperature seems to be a relevant factor for colonisation of new areas by ;Nathusius' Pipistrelle, thus climate change may allow this species to expand its current range in the future (Lundy et al. 2010).	Unknown	Nathusius' Pipistrelle (Pipistrellus nathusii ) ;is a western Palearctic migratory species. It is restricted to Europe, Asia Minor and Transcaucasia where it is found at latitudes of up to ca ;37-63°N. In the Mediterranean, it is generally widespread across southern Europe although apparently absent from most of Iberia (although there have been some new recent records in Spain which extend the known range) and Fennoscandia. ;<span lang="EN-US">It has been found recently in the island of Mallorca in the Balearic islands (Trujillo and García 2009). ;Recent records included also Crete in its range (Benda et al. ;2009). With few exceptions, maternity colonies are confined to northeastern Europe (eastern Germany, Baltic states, Belarus, Ukraine and Russia: Vierhaus 2004) ;<span class="msoIns">and ;<span class="msoIns">occur ;<span class="msoIns">also in England and Northern Ireland<span class="msoIns"> ;(<span class="msoIns">Russ ;et al . ;in ;<span class="msoIns">Harris and ;<span class="SpellE">Yalden<span class="msoIns"> ;2008). The first observation of ;Nathusius' Pipistrelle ;breeding in Finland was made in 2006 (H. Henttonen pers. comm. 2006) when a colony of ca 10 individuals, including lactating females, was found close to the southern coast some 50-60 km east of Helsinki. The species is typically associated with lowland areas but has been recorded up to 2,200 m asl in the Alps (Bogdanowicz 1999).</span></span></span></span>		Terrestrial	It is protected under national law in most range states. It is also protected under international law through the Bonn Convention (Eurobats) and Bern Convention in parts of its range where these apply, and is included in Annex IV of the EU Habitats and Species Directive. It is regarded as a species of special concern by Eurobats. Proposals for the conservation of the species in Europe (including research requirements) were made by Limpens and Schulte (2000) following a workshop in Germany in 1998. They recommended a Europe-wide census involving assessment of the species' population status and trends, identification of its mating, hibernation and maternity areas, investigation of migration routes, and identification of preferred resting areas on migration routes.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pipistrellus	Pipistrellus	nathusii	Keyserling and Blasius	1839	1	Arch. Naturgesch.	5(1): 320	Nathusius' Pipistrelle	 unicolor Fatio, 1905.	Germany, Berlin.	W Europe to Urals and Caucasus, and W Asia Minor; S England.	Not listed.	Least Concern	Subgenus Pipistrellus .	Pipistrellus nathusii	1005628	23	Nathusius's Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	nathusii	Keyserling & Blasius	1839	1						Berlin, Germany.			nathusii (Keyserling & Blasius, 1839)|unicolor (Fatio, 1905)	NA	NA				Ireland|United Kingdom|Spain|France|Luxembourg|Belgium|Netherlands|Germany|Denmark|Norway|Sweden|Finland|Switzerland|Liechtenstein|Italy|Austria|Czech Republic|Poland|Slovakia|Hungary|Slovenia|Croatia|Bosnia & Herzegovina|Serbia|Kosovo|Montenegro|Albania|North Macedonia|Greece|Bulgaria|Turkey|Romania|Moldova|Ukraine|Belarus|Estonia|Latvia|Lithuania|Russia|Georgia|Armenia|Azerbaijan|Kazakhstan	Asia|Europe	Palearctic	LC	0	0	0	Pipistrellus_nathusii	0	sciname match	Pipistrellus_nathusii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pipistrellus_nathusii	1005628	23	Nathusius's Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus	Nannugo	nathusii	von Keyserling & J. H. Blasius	1	Vesperugo Nathusii	Keyserling, A. von and Blasius, J.H. 1839. Uebersicht der Gattungs- und Artcharaktere der europäischen Fledermäuse. Archiv für Naturgeschichte 1839(1):293-331.	https://www.biodiversitylibrary.org/page/24904231				Berlin, Germany.			NA	NA				Ireland|United Kingdom|Spain|France|Luxembourg|Belgium|Netherlands|Germany|Denmark|Norway|Sweden|Finland|Switzerland|Liechtenstein|Italy|Austria|Czech Republic|Poland|Slovakia|Hungary|Slovenia|Croatia|Bosnia and Herzegovina|Serbia|Kosovo|Montenegro|Albania|North Macedonia|Greece|Bulgaria|Turkey|Romania|Moldova|Ukraine|Belarus|Estonia|Latvia|Lithuania|Russia|Georgia|Armenia|Azerbaijan|Kazakhstan	Asia|Europe	Palearctic	LC	0	0	0	Pipistrellus_nathusii	0	sciname match	Pipistrellus_nathusii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pipistrellus	Pipistrellus	nathusii	Keyserling & Blasius	1839	1	Arch. Naturgesch.	5(1): 320	Nathusius' Pipistrelle	unicolor Fatio, 1905.	Germany, Berlin.	W Europe to Urals and Caucasus, and W Asia Minor; S England.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17316/22132621/' target='_blank'>Least Concern</a>	Subgenus Pipistrellus.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pipistrellus nathusii; Pipistrellus nathusii; Pipistrellus nathusii; Pipistrellus nathusii; Pipistrellus nathusii; Pipistrellus nathusii; unicolor; unicolor; nathusii; unicolor; Pipistrelle de Nathusius; Rauhautfledermaus; Pipistrela de Nathusius; Nathusius's Pipistrelle; Nathusius's Pipistrelle; Nathusius' Pipistrelle; P. nathusii