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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1158	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus	Pipistrellus inexspectatus		[MSW2] Subgenus Hypsugo.; [MSW3] Subgenus Pipistrellus. Often misspelled inexpectatus.; [HMW] Pipistrellus inexspectatus Aellen, 1959 , “Ngaouyanga, vallée de la haute Benoué, Cameroon .” Pipustrellus inexspectatus is probably closest to the P. kuhlii group of species. Records previously attributed to it from DR Congo and Kenya are now attributed to Hypsugo eisentrauti . Specific epithet is often misspelled as inexpectatus. Monotypic.; [batnames2022] Subgenus Pipistrellus . Often misspelled inexpectatus .; [IUCN] Taxonomy needs to be checked.; [batnames2023] Subgenus Pipistrellus . Often misspelled inexpectatus .; [batnames2025_1.7] Subgenus Pipistrellus. Often misspelled inexpectatus.														inexspectatus	Taxonomy needs to be checked.			inexspectatus 	inexspectatus 			inexspectatus Aellen, 1959|inexpectatus Zhukova, Yuzefovich, Lebedev, & Kruskop, 2025 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Aellen's pipistrelle	Cameroun, ? Sudan, ? Zaire	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pipistrellus inexspectatus	Cameroun, Upper Benoue Valley, Ngaaouyanga.	Aellen	1959	Arch. Sci. Phys. Nat. Geneve, 12:226.	Distribution: A poorly known species apparently confined to Benin, Cameroon, Zaire, Uganda, and Kenya.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Aellen's pipistrelle	Benin – Uganda, ? Sudan; ref. 4.74	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Aellen	1959	Arch. Sci. Phys. Nat. Geneve, 12:226.	Subgenus Hypsugo.	Benin, Cameroon, Zaire, Uganda, Kenya, perhaps Sudan.	Cameroon, Upper Benoue Valley, Ngaaouyanga.		AELLEN	1957	Size relatively small (forearm length, 31-32 mm). Inner upper incisor bicuspid. Rostrum relatively short and broad. Braincase relatively high.	Distribution: A poorly known species apparently confined to Benin, Cameroon, Zaire, Uganda, and Kenya.	No subspecies.		114	species	P. inexspectatus	AELLEN	1957	Pipistrellus	subgenus	Pipistrellus inexspectatus				Size relatively small (forearm length, 31-32 mm). Inner upper incisor bicuspid. Rostrum relatively short and broad. Braincase relatively high.	No subspecies.		31. P. inexspectatus AELLEN 1957 [kuhlii group],	31	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus inexspectatus	Pipistrellus	Pipistrellus	inexspectatus	Aellen		1959		Arch. Sci. Phys. Nat. Geneve	12		226		Aellen's Pipistrelle	Cameroon, Upper Benoue Valley, Ngaaouyanga.	Sierra Leone, Ghana, Benin, Cameroon, and Uganda. Specimens from Kenya and Dem. Rep. Congo previously referred to this species are now thought to represent eisentrauti; see Koopman et al. (1995). A possible record from Sudan cannot be confirmed as the specimen is too immature to identify (M. Happold, pers. comm.)	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Subgenus Pipistrellus. Often misspelled inexpectatus.	4C3D87E8FFE96A56FF4C9B761710BCFC	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	776	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFE96A56FF4C9B761710BCFC.xml	Pipistrellus inexspectatus	Vespertilionidae	Pipistrellus	inexspectatus	Aellen	1959	Pipistrelle d'Aellen @fr | Aellen-Zwergfledermaus @de | Pipistrela de Aellen @es | Aellen’s Pipistrelle Bat @en | Central Africa Pipistrelle @en	Pipistrellus inexspectatus Aellen, 1959 , “Ngaouyanga, vallée de la haute Benoué, Cameroon .” Pipustrellus inexspectatus is probably closest to the P. kuhlii group of species. Records previously attributed to it from DR Congo and Kenya are now attributed to Hypsugo eisentrauti . Specific epithet is often misspelled as inexpectatus. Monotypic.	W Africa in N Sierra Leone , SW Burkina Faso , C Ghana , N Benin , NW Nigeria , and NC Cameroon ; there are potential records from E Sudan and Uganda , but they require additional investigation.	Head—body 46-8-47-2 mm,tail 30-37-3 mm, ear 10-13 mm, hindfoot 5-6-6 mm, forearm 31-33 mm; weight 3-5-5 g. Dorsum of Aellen’s Pipistrelle is bright reddish brown to dark brown; venter is reddish cream. Ears are brown and relatively short, with rounded tips; tragus is probably just under 50% the ear length and has concave or straight anterior margin, more or less smoothly convex posterior margin with pronounced basal lobule, and broadly rounded tip. Wings are brown, with distinct white hind border. Uropatagium is paler brown than wing membranes, tail extendsslightly past uropatagium, and postcalcarial lobe is hardly visible. Skull is of medium size and rather robust compared with other African Pipistrellus ; braincase is relatively low with moderate breadth; interorbital region is relatively narrow; rostrum is of moderate length and breadth and has well-developed lateral depression just above infraorbital foramina; forehead region is nearly straight to barely concave; I? is long, bicuspid, and straight; I” is small (c.50% the height of I?); P* is tiny and almost never visible above gum; C' and P* touch; and lower molars are myotodont.	Dry and moist savannas and woodlands. One Aellen’s Pipistrelle in Maroua, northern Cameroon , was captured over a riverbed, and two individuals from Burkina Faso were captured in wooded savanna along a rupicolous bar and in a gallery forest.	No information.	No information.	Shape of Aellen’s Pipistrelle’s call is very steep (near vertical), broadband FM; a call reported from Maroua had peak frequency of 54-7 kHz, minimum frequency of 47-7 kHz, mean frequency of 51-2 kHz, knee frequency of 54-7 kHz, characteristic frequency of 47-7 kHz, and duration of 0-46 milliseconds.	No information.	Classified as Data Deficient on The IUCN Red List. Virtually nothing is known about ecology of Aellen’s Pipistrelle. Known distribution is veryscattered, and additional collecting efforts are needed to clarify its conversation status.	ACR (2018) | Aellen (1959) | Fahr (2008e) | Grubb et al. (1998) | Hill & Harrison (1987) | Kangoye, Ouéda, Granjon et al. (2015) | Kangoyé, Ouéda, Thiombiano & Guenda (2012) | Manga Mongombe (2012) | Monadjem, Richards, Taylor & Stoffberg (2013) | Van Cakenberghe & Happold (2013j)	https://zenodo.org/record/6397828/files/figure.png	28. Aellen’s Pipistrelle Pipistrellus inexspectatus French: Pipistrelle d'Aellen / German: Aellen-Zwergfledermaus / Spanish: Pipistrela de Aellen Other common names: Aellen’s Pipistrelle Bat , Central Africa Pipistrelle Taxonomy. Pipistrellus inexspectatus Aellen, 1959 , “Ngaouyanga, vallée de la haute Benoué, Cameroon .” Pipustrellus inexspectatus is probably closest to the P. kuhlii group of species. Records previously attributed to it from DR Congo and Kenya are now attributed to Hypsugo eisentrauti . Specific epithet is often misspelled as inexpectatus. Monotypic. Distribution. W Africa in N Sierra Leone , SW Burkina Faso , C Ghana , N Benin , NW Nigeria , and NC Cameroon ; there are potential records from E Sudan and Uganda , but they require additional investigation. Descriptive notes. Head—body 46-8-47-2 mm,tail 30-37-3 mm, ear 10-13 mm, hindfoot 5-6-6 mm, forearm 31-33 mm; weight 3-5-5 g. Dorsum of Aellen’s Pipistrelle is bright reddish brown to dark brown; venter is reddish cream. Ears are brown and relatively short, with rounded tips; tragus is probably just under 50% the ear length and has concave or straight anterior margin, more or less smoothly convex posterior margin with pronounced basal lobule, and broadly rounded tip. Wings are brown, with distinct white hind border. Uropatagium is paler brown than wing membranes, tail extendsslightly past uropatagium, and postcalcarial lobe is hardly visible. Skull is of medium size and rather robust compared with other African Pipistrellus ; braincase is relatively low with moderate breadth; interorbital region is relatively narrow; rostrum is of moderate length and breadth and has well-developed lateral depression just above infraorbital foramina; forehead region is nearly straight to barely concave; I? is long, bicuspid, and straight; I” is small (c.50% the height of I?); P* is tiny and almost never visible above gum; C' and P* touch; and lower molars are myotodont. Habitat. Dry and moist savannas and woodlands. One Aellen’s Pipistrelle in Maroua, northern Cameroon , was captured over a riverbed, and two individuals from Burkina Faso were captured in wooded savanna along a rupicolous bar and in a gallery forest. Food and Feeding. No information. Breeding. No information. Activity patterns. Shape of Aellen’s Pipistrelle’s call is very steep (near vertical), broadband FM; a call reported from Maroua had peak frequency of 54-7 kHz, minimum frequency of 47-7 kHz, mean frequency of 51-2 kHz, knee frequency of 54-7 kHz, characteristic frequency of 47-7 kHz, and duration of 0-46 milliseconds. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Virtually nothing is known about ecology of Aellen’s Pipistrelle. Known distribution is veryscattered, and additional collecting efforts are needed to clarify its conversation status. Bibliography. ACR (2018), Aellen (1959), Fahr (2008e), Grubb et al. (1998), Hill & Harrison (1987), Kangoye, Ouéda, Granjon et al. (2015), Kangoyé, Ouéda, Thiombiano & Guenda (2012), Manga Mongombe (2012) , Monadjem, Richards, Taylor & Stoffberg (2013), Van Cakenberghe & Happold (2013j).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pipistrellus inexspectatus	Pipistrellus	Pipistrellus	inexspectatus	Aellen	1959	0	Arch. Sci. Phys. Nat. Geneve	0.6569	Aellen's Pipistrelle	None.	Cameroon, Upper Benoue Valley, Ngaaouyanga.	Sierra Leone, Ghana, Benin, Cameroon, and Uganda. Specimens from Kenya and Dem. Rep. Congo previously referred to this species are now thought to represent eisentrauti; see Koopman et al. (1995). A possible record from Sudan cannot be confirmed as the specimen is too immature to identify (M. Happold, pers. comm.)	Not listed.	Data Deficient	Subgenus Pipistrellus . Often misspelled inexpectatus .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pipistrellus inexspectatus	23	Aellen's Pipistrelle	Aellen's Pipistrelle Bat|Central Africa Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	inexspectatus	Aellen	1959	0	Pipistrellus_inexspectatus	Aellen, V. (1959). Chiroptères nouveaux d'Afrique. Archives des Sciences Physiques et Naturelles Genève, 12(2), 226.	https://www.e-periodica.ch/digbib/view?pid=ads-002%3A1959%3A12%3A%3A140#237	MHNC (Original No.) 954		"Ngaouyanga, vallée de la haute Bénoué, Cameroon."			inexspectatus Aellen, 1959	NA	NA	Sierra Leone|Burkina Faso|Ghana|Benin|Nigeria|Cameroon|Sudan?|South Sudan?|Uganda?	Africa	Afrotropic	DD	0	0	0	Pipistrellus_inexspectatus	0	sciname match	Pipistrellus_inexspectatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17343	Pipistrellus inexspectatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pipistrellus	inexspectatus	Aellen, 1959	Taxonomy needs to be checked.	20000000	Pipistrellus inexspectatus	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Data Deficient in view of the absence of sufficient information on its extent of occurrence, natural history, threats and conservation status.	This species has been recorded from both dry and moist savanna habitats. The type locality is reportedly from Guinea woodland (Grubb et al. 1998).	The threats to this species are poorly known.	Little information is available on the population abundance or size of this species.	Unknown	This little known species has been recorded from a few localities in West and Central Africa. It has been reported from Sierra Leone, Ghana, Benin and Nigeria and Cameroon. There are also uncertain records from east Sudan and Uganda, although the specimens in question are difficult to certainly allocate to Pipistrellus inexspectatus (Koopman 1975). The records of this species from the Democratic Republic of the Congo and Kenya are now allocated to Pipistrellus eisentrauti (Koopman et al. 1995; Simmons 2005).		Terrestrial	It is not known if the species is present in any protected areas. Further field surveys are needed to better understand the distribution, natural history and possible threats to this little known bats species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pipistrellus	Pipistrellus	inexspectatus	Aellen	1959	0	Arch. Sci. Phys. Nat. Geneve	0.656944	Aellen's Pipistrelle	None.	Cameroon, Upper Benoue Valley, Ngaaouyanga.	Sierra Leone, Ghana, Benin, Cameroon, and Uganda. Specimens from Kenya and Dem. Rep. Congo previously referred to this species are now thought to represent eisentrauti; see Koopman et al. (1995). A possible record from Sudan cannot be confirmed as the specimen is too immature to identify (M. Happold, pers. comm.)	Not listed.	Data Deficient	Subgenus Pipistrellus . Often misspelled inexpectatus .	Pipistrellus inexspectatus	1005621	23	Aellen's Pipistrelle	Aellen's Pipistrelle Bat|Central Africa Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	inexspectatus	Aellen	1959	0	Pipistrellus_inexspectatus	Aellen, V. (1959). Chiroptères nouveaux d'Afrique. Archives des Sciences Physiques et Naturelles Genève, 12(2), 226.	https://www.e-periodica.ch/digbib/view?pid=ads-002%3A1959%3A12%3A%3A140#237	MHNC (Original No.) 954		"Ngaouyanga, vallée de la haute Bénoué, Cameroon."			inexspectatus Aellen, 1959	NA	NA				Sierra Leone|Burkina Faso|Ghana|Benin|Nigeria|Cameroon|Sudan?|South Sudan?|Uganda?	Africa	Afrotropic	DD	0	0	0	Pipistrellus_inexspectatus	0	sciname match	Pipistrellus_inexspectatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pipistrellus_inexspectatus	1005621	23	Aellen's Pipistrelle	Aellen's Pipistrelle Bat|Central Africa Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus	incertae sedis	inexspectatus	Aellen	0	Pipistrellus inexspectatus	Aellen, V. 1959. Chiroptères nouveaux d'Afrique. Archives des Sciences 12(2):217-240.	https://www.e-periodica.ch/digbib/view?pid=ads-002%3A1959%3A12%3A%3A140#237	MHNLCF (number not known)	holotype		"Ngaouyanga, vallée de la haute Bénoué, Cameroon."			NA	NA				Sierra Leone|Burkina Faso|Ghana|Benin|Nigeria|Cameroon|Sudan?|South Sudan?|Uganda?	Africa	Afrotropic	DD	0	0	0	Pipistrellus_inexspectatus	0	sciname match	Pipistrellus_inexspectatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pipistrellus	Pipistrellus	inexspectatus	Aellen	1959	0	Arch. Sci. Phys. Nat. Geneve	0.656944	Aellen's Pipistrelle	None.	Cameroon, Upper Benoue Valley, Ngaaouyanga.	Sierra Leone, Ghana, Benin, Cameroon, and Uganda. Specimens from Kenya and Dem. Rep. Congo previously referred to this species are now thought to represent eisentrauti; see Koopman et al. (1995). A possible record from Sudan cannot be confirmed as the specimen is too immature to identify (M. Happold, pers. comm.)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17343/22128783/' target='_blank'>Data Deficient</a>	Subgenus Pipistrellus. Often misspelled inexpectatus.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pipistrellus inexspectatus; Pipistrellus inexspectatus; Pipistrellus inexspectatus; Pipistrellus inexspectatus; Pipistrellus inexspectatus; Pipistrellus inexspectatus; inexspectatus; Pipistrelle d'Aellen; Aellen-Zwergfledermaus; Pipistrela de Aellen; Aellen’s Pipistrelle Bat; Central Africa Pipistrelle; Aellen's Pipistrelle; Aellen's Pipistrelle Bat; Central Africa Pipistrelle; Aellen's Pipistrelle; Aellen's Pipistrelle; P. inexspectatus