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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1155	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Pipistrellus creticus	Pipistrellus creticus	Pipistrellus creticus	Pipistrellus hanaki creticus	Pipistrellus creticus	Pipistrellus creticus	Pipistrellus creticus	Pipistrellus creticus	Pipistrellus creticus		[HMW] Pipustrellus hanaki creticus Benda in Benda et al, 2009, “ Greece , Crete , Rouva oak forest ; a pasture with creek at the Agiou Ioanni chapel, Prefecture of Irakleio [= Heraklion]; 35° 11’ N , 24° 55’ E , 940 m asl.” Pipistrellus creticus was originally described as a subspecies of P. hanaki ; however, it was found to be sister to P. pygmaeus and is now recognized as a distinct species. Nevertheless, earlier studies determined that P. creticus and P. hanaki were sister species, but this was notstatistically supported and only cytochrome-b data were used. Monotypic.; [batnames2022] Previously considered a subspecies of hanaki , but distinct; see Benda et al. (2014).; [MDD2022] recently described as a subspecies of P. hanaki, but has been subsequently recognized as a distinct species; [batnames2023] Previously considered a subspecies of hanaki , but distinct; see Benda et al. (2014).; [MDD2023] recently described as a subspecies of P. hanaki, but has been subsequently recognized as a distinct species; [MDD2025_2.0] recently described as a subspecies of P. hanaki, but has been subsequently recognized as a distinct species; [batnames2025_1.7] Previously considered a subspecies of hanaki, but distinct; see Benda et al. (2014).; [MDD2025_2.2] recently described as a subspecies of P. hanaki, but has been subsequently recognized as a distinct species														creticus				creticus	creticus			creticus Benda in Benda, Georgiakakis, Dietz, Hanák, Galanaki, Markantonatou, Chudárková, Hulva, & Horáček, 2008						N/A																																								NA																											4C3D87E8FFF56A4AFA58977D1959B266	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	772	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFF56A4AFA58977D1959B266.xml	Pipistrellus creticus	Vespertilionidae	Pipistrellus	creticus	Benda	2008	Pipistrelle de Crete @fr | Kreta-Zwergfledermaus @de | Pipistrelade Creta @es	Pipustrellus hanaki creticus Benda in Benda et al, 2009, “ Greece , Crete , Rouva oak forest ; a pasture with creek at the Agiou Ioanni chapel, Prefecture of Irakleio [= Heraklion]; 35° 11’ N , 24° 55’ E , 940 m asl.” Pipistrellus creticus was originally described as a subspecies of P. hanaki ; however, it was found to be sister to P. pygmaeus and is now recognized as a distinct species. Nevertheless, earlier studies determined that P. creticus and P. hanaki were sister species, but this was notstatistically supported and only cytochrome-b data were used. Monotypic.	Crete .	Head-body 37-43 mm,tail 34-37 mm, ear 11-3-12-7 mm, forearm 29-30-3 mm; weight 3-3-3-9 g. The Crete Pipistrelle is externally similar to Hanak’s Pipistrelle ( P. hanaki ) but significantly smaller in body and skull sizes. Dorsal pelage is rusty brown (darkest along back and top of head and becoming lighter along sides); ventral pelage is paler brown; and face, wing membranes, ears, and tragus are dark brown, being slightly paler around eyes and bases of ears. Colors are more vibrant in reproductive condition, especially around face and behind ears, which are more reddish. Ears are rounded at tops, and tragus has rounded tip and curves forward near top. Uropatagium stretches from calcar at ankles to tail tip (although tail extends c.0-5— 1 mm beyond uropatagium) and is the same color as wing membranes. Glans penis is simple and has more or less obvious pale medialstripe dorsally and foreskin that is covered in grayish brown hairs. Skull is clearly very small, and braincase is relatively broad and high compared with that of Hanak’s Pipistrelle, also having shorter and narrow rostrum on average; tooth row is shorter than in Hanak’s Pipistrelle.	Mediterranean woodland dominated by Quercus ( Fagaceae ) and Cupressus ( Cupressaceae ) from sea level up to elevations of ¢. 940 m . Crete Pipistrelles have also been found in sweet chestnut ( Castanea sativa, Fagaceae ) plantations.	Crete Pipistrelles are insectivorous, probably feeding by aerialhawking in open areas. Feces of eight individuals (55 pellets) were dominated by Brachycera, Auchenorrhyncha, and Lepidoptera and had smaller amounts of Hymenoptera (Formicoidea and Ichneumonoidea) , Nematocera ( Culicidae , Chironomidae ), and Coleoptera .	No information.	The Crete Pipistrelle is nocturnal. Foraging call shape is FM-QCEF. Start frequency averages 59-5 kHz (46-2-101-1 kHz), end frequency 48-8 kHz (43-6 55-2 kHz), and pulse duration 4-3 milliseconds (1-5-8-6 milliseconds). Two types of social calls have also been recorded: A type (peak frequency of ¢.27-7 kHz and duration of 19-9 milliseconds) and B type (peak frequency of ¢.20-9 kHz and duration of 24-8 milliseconds). B type calls were more prevalent in July-October; A type calls decreased in use from July to November.	No information.	Not assessed on The IUCN Red List. Taxonomic status of the Crete Pipistrelle was only recently confirmed as distinct. It is common on Crete but only found there, which might indicate some need for protection. Additional research is needed to clarify its conservation status.	Benda, Georgiakakis et al. (2008) | Benda, Spitzenberger et al. (2014) | Evin et al. (2011) | Georgiakakis & Russo (2012) | Zdarska (2013)	https://zenodo.org/record/6397812/files/figure.png	20. Crete Pipistrelle Pipistrellus creticus French: Pipistrelle de Crete / German: Kreta-Zwergfledermaus / Spanish: Pipistrela de Creta Taxonomy. Pipustrellus hanaki creticus Benda in Benda et al, 2009, “ Greece , Crete , Rouva oak forest ; a pasture with creek at the Agiou Ioanni chapel, Prefecture of Irakleio [= Heraklion]; 35° 11’ N , 24° 55’ E , 940 m asl.” Pipistrellus creticus was originally described as a subspecies of P. hanaki ; however, it was found to be sister to P. pygmaeus and is now recognized as a distinct species. Nevertheless, earlier studies determined that P. creticus and P. hanaki were sister species, but this was notstatistically supported and only cytochrome-b data were used. Monotypic. Distribution. Crete . Descriptive notes. Head-body 37-43 mm,tail 34-37 mm, ear 11-3-12-7 mm, forearm 29-30-3 mm; weight 3-3-3-9 g. The Crete Pipistrelle is externally similar to Hanak’s Pipistrelle ( P. hanaki ) but significantly smaller in body and skull sizes. Dorsal pelage is rusty brown (darkest along back and top of head and becoming lighter along sides); ventral pelage is paler brown; and face, wing membranes, ears, and tragus are dark brown, being slightly paler around eyes and bases of ears. Colors are more vibrant in reproductive condition, especially around face and behind ears, which are more reddish. Ears are rounded at tops, and tragus has rounded tip and curves forward near top. Uropatagium stretches from calcar at ankles to tail tip (although tail extends c.0-5— 1 mm beyond uropatagium) and is the same color as wing membranes. Glans penis is simple and has more or less obvious pale medialstripe dorsally and foreskin that is covered in grayish brown hairs. Skull is clearly very small, and braincase is relatively broad and high compared with that of Hanak’s Pipistrelle, also having shorter and narrow rostrum on average; tooth row is shorter than in Hanak’s Pipistrelle. Habitat. Mediterranean woodland dominated by Quercus ( Fagaceae ) and Cupressus ( Cupressaceae ) from sea level up to elevations of ¢. 940 m . Crete Pipistrelles have also been found in sweet chestnut ( Castanea sativa, Fagaceae ) plantations. Food and Feeding. Crete Pipistrelles are insectivorous, probably feeding by aerialhawking in open areas. Feces of eight individuals (55 pellets) were dominated by Brachycera, Auchenorrhyncha, and Lepidoptera and had smaller amounts of Hymenoptera (Formicoidea and Ichneumonoidea) , Nematocera ( Culicidae , Chironomidae ), and Coleoptera . Breeding. No information. Activity patterns. The Crete Pipistrelle is nocturnal. Foraging call shape is FM-QCEF. Start frequency averages 59-5 kHz (46-2-101-1 kHz), end frequency 48-8 kHz (43-6 55-2 kHz), and pulse duration 4-3 milliseconds (1-5-8-6 milliseconds). Two types of social calls have also been recorded: A type (peak frequency of ¢.27-7 kHz and duration of 19-9 milliseconds) and B type (peak frequency of ¢.20-9 kHz and duration of 24-8 milliseconds). B type calls were more prevalent in July-October; A type calls decreased in use from July to November. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The IUCN Red List. Taxonomic status of the Crete Pipistrelle was only recently confirmed as distinct. It is common on Crete but only found there, which might indicate some need for protection. Additional research is needed to clarify its conservation status. Bibliography. Benda, Georgiakakis et al. (2008), Benda, Spitzenberger et al. (2014), Evin et al. (2011), Georgiakakis & Russo (2012), Zdarska (2013).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pipistrellus creticus	Pipistrellus	Pipistrellus	creticus	Benda	2009	0	Acta Soc. Zool. Bohem.	74:33:00	Crete Pipistrelle	None.	Greece, Crete, Rouva oak forest; a pasture with creek at the Agiou Ioanni chapel, Prefecture of Irakleio	Crete (Greece)	Not listed.	Vulnerable under Pipistrellus hanaki 	Previously considered a subspecies of hanaki , but distinct; see Benda et al. (2014).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pipistrellus creticus	23	Crete Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	creticus	Benda in Benda, Georgiakakis, Dietz, Hanák, Galanaki, Markantonatou, Chudárková, Hulva, & Horáček	2009	0	Pipistrellus_hanaki_creticus	Benda, P., Georgiakakis, P., Dietz, C., Hanák, V., Galanaki, K., Markantonatou, V., Chudárková, A., Hulva, P. & Horáček, I. (2008). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 7. The bat fauna of Crete, Greece. Acta Societatis Zoologicae Bohemicae, 72, 105-190.	http://nm-cz.nm.netservis.cz/download/pm/zoo/benda_lit/Benda2009aszb.pdf	NMP 92350		"Greece, Crete, Rouva oak forest; a pasture with creek at the Agiou Ioanni chapel, Prefecture of Irakleio [= Heraklion]; 35° 11' N, 24° 55' E, 940 m a.s.l."	35.183	24.917	creticus Benda in Benda, Georgiakakis, Dietz, Hanák, Galanaki, Markantonatou, Chudárková, Hulva, & Horáček, 2009	recently described as a subspecies of P. hanaki, but has been subsequently recognized as a distinct species	Benda, P., Georgiakakis, P., Dietz, C., Hanák, V., Galanaki, K., Markantonatou, V., ... & Horáček, I. (2008). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 7. The bat fauna of Crete, Greece. Acta Societatis Zoologicae Bohemicae, 72, 105-190.|Benda, P., Spitzenberger, F., Hanák, V., Andreas, M., Reiter, A., Å evčík, M., ... & Uhrin, M. (2014). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 11. On the bat fauna of Libya II. Acta Societatis Zoologicae Bohemicae, 78, 1-162.	Greece	Europe	Palearctic	NA	0	0	0	Pipistrellus_creticus	0	unmatched	NA	1																																			Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pipistrellus	Pipistrellus	creticus	Benda	2009	0	Acta Soc. Zool. Bohem.	74:33:00	Crete Pipistrelle	None.	Greece, Crete, Rouva oak forest; a pasture with creek at the Agiou Ioanni chapel, Prefecture of Irakleio	Crete (Greece)	Not listed.	Vulnerable under Pipistrellus hanaki 	Previously considered a subspecies of hanaki , but distinct; see Benda et al. (2014).	Pipistrellus creticus	1005616	23	Crete Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	creticus	Benda in Benda, Georgiakakis, Dietz, Hanák, Galanaki, Markantonatou, Chudárková, Hulva, & Horáček	2009	0	Pipistrellus_hanaki_creticus	Benda, P., Georgiakakis, P., Dietz, C., Hanák, V., Galanaki, K., Markantonatou, V., Chudárková, A., Hulva, P. & Horáček, I. (2008). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 7. The bat fauna of Crete, Greece. Acta Societatis Zoologicae Bohemicae, 72, 105-190.	http://nm-cz.nm.netservis.cz/download/pm/zoo/benda_lit/Benda2009aszb.pdf	NMP 92350		"Greece, Crete, Rouva oak forest; a pasture with creek at the Agiou Ioanni chapel, Prefecture of Irakleio [= Heraklion]; 35° 11' N, 24° 55' E, 940 m a.s.l."	35.18333	24.91667	creticus Benda in Benda, Georgiakakis, Dietz, Hanák, Galanaki, Markantonatou, Chudárková, Hulva, & Horáček, 2009	recently described as a subspecies of P. hanaki, but has been subsequently recognized as a distinct species	Benda, P., Georgiakakis, P., Dietz, C., Hanák, V., Galanaki, K., Markantonatou, V., ... & Horáček, I. (2008). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 7. The bat fauna of Crete, Greece. Acta Societatis Zoologicae Bohemicae, 72, 105-190.|Benda, P., Spitzenberger, F., Hanák, V., Andreas, M., Reiter, A., Å evčík, M., ... & Uhrin, M. (2014). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 11. On the bat fauna of Libya II. Acta Societatis Zoologicae Bohemicae, 78, 1-162.				Greece	Europe	Palearctic	NA	0	0	0	Pipistrellus_creticus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pipistrellus_creticus	1005616	23	Crete Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus	Pipistrellus	creticus	Benda in Benda, Georgiakakis, Dietz, Hanák, Galanaki, Markantonatou, Chudárková, Hulva, & Horáček	0	Pipistrellus hanaki creticus	Benda, P., Georgiakakis, P., Dietz, C., Hanák, V., Galanaki, K., Markantonatou, V., Chudárková, A., Hulva, P. and Horáček, I. 2008. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 7. The bat fauna of Crete, Greece. Acta Societatis Zoologicae Bohemicae 72:105-190.	https://www.researchgate.net/publication/228668708_BatsMammalia_Chiroptera_of_the_Eastern_Mediterranean_and_Middle_East_Part_7_The_bat_fauna_of_Crete_Greece	NMPr P6V-092350	holotype		"Greece, Crete, Rouva oak forest; a pasture with creek at the Agiou Ioanni chapel, Prefecture of Irakleio [= Heraklion]; 35° 11' N, 24° 55' E, 940 m a.s.l."	35.18333	24.91667	recently described as a subspecies of P. hanaki, but has been subsequently recognized as a distinct species	Benda, P., Georgiakakis, P., Dietz, C., Hanák, V., Galanaki, K., Markantonatou, V., ... & Horáček, I. (2008). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 7. The bat fauna of Crete, Greece. Acta Societatis Zoologicae Bohemicae, 72, 105-190.|Benda, P., Spitzenberger, F., Hanák, V., Andreas, M., Reiter, A., Å evčík, M., ... & Uhrin, M. (2014). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 11. On the bat fauna of Libya II. Acta Societatis Zoologicae Bohemicae, 78, 1-162.				Greece	Europe	Palearctic	NE	0	0	0	Pipistrellus_creticus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pipistrellus	Pipistrellus	creticus	Benda	2009	0	Acta Soc. Zool. Bohem.	74:33:00	Crete Pipistrelle	None.	Greece, Crete, Rouva oak forest; a pasture with creek at the Agiou Ioanni chapel,    Prefecture of Irakleio	Crete (Greece)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	Not Evaluated	Previously considered a subspecies of hanaki, but distinct; see Benda et al. (2014).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pipistrellus creticus; Pipistrellus creticus; Pipistrellus creticus; Pipistrellus creticus; creticus; Pipistrelle de Crete; Kreta-Zwergfledermaus; Pipistrelade Creta; Crete Pipistrelle; Crete Pipistrelle; P. creticus