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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1150	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus	Phyllostomus elongatus		[HMW] Phyllostoma elongatum E. Geoffroy Saint-Hilaire, 1810 , type locality not given. Restricted by A. Cabrera in 1958 to Rio Branco, Mato Grosso , Brazil . This species is monotypic.						ater.			ater			elongatus 	elongatus - ater 	elongatus, ater, lanceolatum		elongatus 	elongatus - ater 	elongatus, ater, lanceolatum	elongatus, ater, lanceolatum	elongatus 	elongatus - ater	elongatus É. Geoffroy Saint-Hilaire, 1810|ater (J. E. Gray, 1866)|lanceolatum von Pelzeln, 1883 [nomen nudum]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Colombia, Venezuela – E Peru, Bolivia, SE Brazil	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Phyllostomus elongatus	Brazil, Mato Grosso, Rio Branco.	E. Geoffroy	1810	Ann. Mus. Hist. Nat. Paris, 15:182.	Distribution: Ranging across tropical mainland South America from Colombia to eastern Brazil, but west of the Andes not south of Ecuador.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Colombia, Guianas – E Peru, Bolivia, SE Brazi	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	E. Geoffroy	1810	Ann. Mus. Hist. Nat. Paris, 15:182.		Bolivia, E Peru, Ecuador, and Colombia to Guianas and E Brazil.	Brazil, Mato Grosso, Rio Branco.		E. GEOFFROY	1810	Size fairly large (forearm length, 61-71 mm; condylobasal length, 24-28 mm). Calcar longer than hind foot. Sagittal crest well developed.	Distribution: Ranging across tropical mainland South America from Colombia to eastern Brazil, but west of the Andes not south of Ecuador.	No subspecies.		76	species	P. elongatus	E. GEOFFROY	1810	Phyllostomus	genus	Phyllostomus elongatus				Size fairly large (forearm length, 61-71 mm; condylobasal length, 24-28 mm). Calcar longer than hind foot. Sagittal crest well developed.	No subspecies.		3. P. elongatus (E. GEOFFROY 1810).	3	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Phyllostominae		Phyllostomus elongatus	Phyllostomus		elongatus	E. Geoffroy	y	1810		Ann. Mus. Natn. Hist. Nat. Paris	15		182		Lesser Spear-nosed Bat	Brazil, Mato Grosso, Rio Branco.	Bolivia, E Peru, Ecuador, and Colombia to Guianas and E Brazil.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	ater Gray, 1866.		03A687BCFFA6FFA613BFFC42FBE9FFC8	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	507	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFA6FFA613BFFC42FBE9FFC8.xml	Phyllostomus elongatus	Phyllostomidae	Phyllostomus	elongatus	E. Geoffroy Saint-Hilaire	1810	Phyllostome des sous-bois @fr | Kleine Lanzennase @de | Filostomapequeno @es	Phyllostoma elongatum E. Geoffroy Saint-Hilaire, 1810 , type locality not given. Restricted by A. Cabrera in 1958 to Rio Branco, Mato Grosso , Brazil . This species is monotypic.	Lowlands E of Andes in Colombia , Venezuela , the Guianas, Amazonian Basin to SE Peru and N & C Bolivia , and E Brazil , also an isolated population in W Colombia and NW Ecuador .	Head-body 65-100 mm, tail 15-24 mm, ear 25-32 mm, hindfoot 14-20 mm, forearm 58-71 mm; weight 30-57 g. The Lesser Spear-nosed Bat is large and robust. Its morphology is very similar to the Greater Spear-nosed Bat ( P. hastatus ), but it is much smaller and less robust in all aspects. Fur is short, soft, velvety, and glossy; dorsal fur is brown to reddish brown; head, neck, and shoulders are often paler than back; venter is paler than back to fairly dark. Head is large; snout is broad and robust; face and noseleaf are black to dark brown; noseleaf is medium-sized, broad, and pear-shaped, with base horseshoe-shaped and entire free edge separated from upper lip; ears are medium-sized, broad, and triangular, with rounded tips; eyes are large; and chin has small tubercles forming a Vsshape. Wing membranes are blackish, but tips are sometimes white; caudal membrane is developed; calcar is longer than foot (more than 14 mm ); and tail is short, reaching to middle of membrane or less. Teeth are strong and pointed. Chromosomal complement has 2n = 32 and FN = 58.	Mainly evergreen forest habitats but also humid tropical to low subtropical forests, Andean foothills mainly in Amazonia, and humid tropical forest in Choco Ecoregion at elevations of 60-1100 m. The Lesser Spearnosed Bat is occasionally found in drier habitats near streams, semi-humid forests, and flooded savannas. It also occurs in terra firma forest; flooded forests; swamps; primary, secondary, selectively logged and gallery forests; forest edges; cultivated areas; and pastures. It usually flies low over streams and rivers.	Diet of the Lesser Spear-nosed Bat includes small vertebrates, flower parts, nectar, fruits, and large insects. In Peru , pollen has been found on it. In Bolivia , several families of insects occur in its diet ( Blattidae , Gryllidae , Reduviidae , Carabidae , Curculionidae , Hydrophylidae, Scarabaeidae , and Formicidae ), along with orthopterans, coleopterans, and other undetermined arthropods. In Brazil , fruit of Rollinia mucosa ( Magnoliales , Annonaceae ) was eaten. It is occasionally carnivorous; in Amazonas, Brazil , it was reported eating ajuvenile of Seba’s Short-tailed Bat ( Carollia perspicillata ).	Reproductive female Lesser Spear-nosed Bats have been found in June in Colombia and July-August in Peru . In western Ecuador , mothers with young attached to their nipples have been recorded in November and February. In northern Brazil , several pregnant and lactating females were caught at beginning of rainy season.	The Lesser Spear-nosed Bat is nocturnal and aerial. It emerges from its roosts at complete darkness and forage for c.2 hours before returning to roosts. In Bolivia , it has been captured in mist net at 18:00-20:00 h. Echolocation calls are 51-4-78-2 kHz. It roosts in hollow trees, culverts, and tunnels and under bridges.	Lesser Spear-nosed Bats roost in aggregations, divided up into smaller social units consisting of permanent harems of a single male and up to twelve females and their young and groups of non-breeding males. This social structure is very similar to that of the Greater Spear-nosed Bat. In Esmeraldas , Ecuador , a colony of ¢.200 individuals was found sharing space with Seba’s Short-tailed Bats. In Beni , Bolivia , an individual was found roosting in a tree hollow of Gallesia integrifolia ( Phytolaccaceae ), sharing space with Seba’s Short-tailed Bats, Common Vampire Bats ( Desmodus rotundus ), and Pallas’s Long-tongued Bats ( Glossophaga soricina ).	Classified as Least Concern on The IUCN Red List. The Lesser Spear-nosed Batis fairly common and widespread. Its conservation status is considered stable and is found at several protected areas.	Aguirre (2002) | Aguirre & Teran (2007) | Aguirre, Lens & Matthysen (2003) | Albuja (1999) | Cabrera (1958) | Fischer et al. (1997) | Gardner (1977b) | Handley (1976) | Hill (1965a) | Nogueira, Peracchi & Moratelli (2007) | Rivera et al. (2015) | Tirira (2017) | Tuttle (1970) | Williams & Genoways (2008)	https://zenodo.org/record/6458683/files/figure.png	40. Lesser Spear-nosed Bat Phyllostomus elongatus French: Phyllostome des sous-bois / German: Kleine Lanzennase / Spanish: Filostoma pequeno Taxonomy. Phyllostoma elongatum E. Geoffroy Saint-Hilaire, 1810 , type locality not given. Restricted by A. Cabrera in 1958 to Rio Branco, Mato Grosso , Brazil . This species is monotypic. Distribution. Lowlands E of Andes in Colombia , Venezuela , the Guianas, Amazonian Basin to SE Peru and N & C Bolivia , and E Brazil , also an isolated population in W Colombia and NW Ecuador . Descriptive notes. Head-body 65-100 mm, tail 15-24 mm, ear 25-32 mm, hindfoot 14-20 mm, forearm 58-71 mm; weight 30-57 g. The Lesser Spear-nosed Bat is large and robust. Its morphology is very similar to the Greater Spear-nosed Bat ( P. hastatus ), but it is much smaller and less robust in all aspects. Fur is short, soft, velvety, and glossy; dorsal fur is brown to reddish brown; head, neck, and shoulders are often paler than back; venter is paler than back to fairly dark. Head is large; snout is broad and robust; face and noseleaf are black to dark brown; noseleaf is medium-sized, broad, and pear-shaped, with base horseshoe-shaped and entire free edge separated from upper lip; ears are medium-sized, broad, and triangular, with rounded tips; eyes are large; and chin has small tubercles forming a Vsshape. Wing membranes are blackish, but tips are sometimes white; caudal membrane is developed; calcar is longer than foot (more than 14 mm ); and tail is short, reaching to middle of membrane or less. Teeth are strong and pointed. Chromosomal complement has 2n = 32 and FN = 58. Habitat. Mainly evergreen forest habitats but also humid tropical to low subtropical forests, Andean foothills mainly in Amazonia, and humid tropical forest in Choco Ecoregion at elevations of 60-1100 m. The Lesser Spearnosed Bat is occasionally found in drier habitats near streams, semi-humid forests, and flooded savannas. It also occurs in terra firma forest; flooded forests; swamps; primary, secondary, selectively logged and gallery forests; forest edges; cultivated areas; and pastures. It usually flies low over streams and rivers. Food and Feeding. Diet of the Lesser Spear-nosed Bat includes small vertebrates, flower parts, nectar, fruits, and large insects. In Peru , pollen has been found on it. In Bolivia , several families of insects occur in its diet ( Blattidae , Gryllidae , Reduviidae , Carabidae , Curculionidae , Hydrophylidae, Scarabaeidae , and Formicidae ), along with orthopterans, coleopterans, and other undetermined arthropods. In Brazil , fruit of Rollinia mucosa ( Magnoliales , Annonaceae ) was eaten. It is occasionally carnivorous; in Amazonas, Brazil , it was reported eating ajuvenile of Seba’s Short-tailed Bat ( Carollia perspicillata ). Breeding. Reproductive female Lesser Spear-nosed Bats have been found in June in Colombia and July-August in Peru . In western Ecuador , mothers with young attached to their nipples have been recorded in November and February. In northern Brazil , several pregnant and lactating females were caught at beginning of rainy season. Activity patterns. The Lesser Spear-nosed Bat is nocturnal and aerial. It emerges from its roosts at complete darkness and forage for c.2 hours before returning to roosts. In Bolivia , it has been captured in mist net at 18:00-20:00 h. Echolocation calls are 51-4-78-2 kHz. It roosts in hollow trees, culverts, and tunnels and under bridges. Movements, Home range and Social organization. Lesser Spear-nosed Bats roost in aggregations, divided up into smaller social units consisting of permanent harems of a single male and up to twelve females and their young and groups of non-breeding males. This social structure is very similar to that of the Greater Spear-nosed Bat. In Esmeraldas , Ecuador , a colony of ¢.200 individuals was found sharing space with Seba’s Short-tailed Bats. In Beni , Bolivia , an individual was found roosting in a tree hollow of Gallesia integrifolia ( Phytolaccaceae ), sharing space with Seba’s Short-tailed Bats, Common Vampire Bats ( Desmodus rotundus ), and Pallas’s Long-tongued Bats ( Glossophaga soricina ). Status and Conservation. Classified as Least Concern on The IUCN Red List. The Lesser Spear-nosed Batis fairly common and widespread. Its conservation status is considered stable and is found at several protected areas. Bibliography. Aguirre (2002), Aguirre & Teran (2007), Aguirre, Lens & Matthysen (2003), Albuja (1999), Cabrera (1958), Fischer et al. (1997), Gardner (1977b), Handley (1976), Hill (1965a), Nogueira, Peracchi & Moratelli (2007), Rivera et al. (2015), Tirira (2017), Tuttle (1970), Williams & Genoways (2008).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Phyllostomus elongatus	Phyllostomus		elongatus	E. Geoffroy	1810	1	Ann. Mus. Natn. Hist. Nat. Paris	0.7514	Lesser Spear-nosed Bat	 ater Gray, 1866.	Brazil, Mato Grosso, Rio Branco.	Bolivia, E Peru, Ecuador, and Colombia to Guianas and E Brazil.	Not listed.	Least Concern		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Phyllostomus elongatus	23	Lesser Spear-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	PHYLLOSTOMINAE	PHYLLOSTOMINI	Phyllostomus	NA	elongatus	É. Geoffroy Saint-Hilaire	1810	1	Phyllostoma_elongatum	Geoffroy Saint-Hilaire, É. (1810). Sur les Phyllostomes et les Mégadermes, deux genres de la famille des Chauve-souris. Annales du Muséum d'histoire naturelle, 15, 182.	https://www.biodiversitylibrary.org/item/23421#page/206/mode/1up	MNHN 1986-1044		type locality not given. Restricted by A. Cabrera in 1958 to Rio Branco, Mato Grosso, Brazil.			elongatus (É. Geoffroy Saint-Hilaire, 1810)|ater (J. E. Gray, 1866)|lanceolatum (Pelzeln, 1883) [nomen nudum]	NA	NA	Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia	South America	Neotropic	LC	0	0	0	Phyllostomus_elongatus	0	sciname match	Phyllostomus_elongatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17217	Phyllostomus elongatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Phyllostomus	elongatus	(É. Geoffroy, 1810)		20000000	Phyllostomus elongatus	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern because it is widely distributed, relatively common, and unlikely to be declining at a rate which would qualify the species for inclusion in a threatened category.	This species strongly prefers multistratal tropical evergreen forest although it is occasionally taken in dryer habitats near streams. It roosts in small groups in tree cavities, or large groups in caves. These roosting congregations are divided up into smaller permanent social units consisting of harems of a single male and several females and their young, and groups of non-breeding bachelors. Apparently it is strongly disposed to feed on fruits, but it may also take nectar and pollen and large insects (Gardner 1977). It also hunts aerially above the forest canopy. These bats emerge from their roosts at complete darkness and forage for about two hours before returning to the roost.	Deforestation occurs in some parts of its range though this is not considered a major threat.	It is moderately common.	Unknown	This species ranges across Colombia, Venezuela, the Guianas, Ecuador, Peru, Bolivia and southeastern Brazil (Eisenberg 1989, Lim and Engstrom 2000) in the lowlands only.	This species is not used.	Terrestrial	It is recommended to reduce loss of forest habitats.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Phyllostomus		elongatus	É. Geoffroy Saint-Hilaire	1810	1	Ann. Mus. Natn. Hist. Nat. Paris	0.751389	Lesser Spear-nosed Bat	 ater Gray, 1866.	Brazil, Mato Grosso, Rio Branco.	Bolivia, E Peru, Ecuador, and Colombia to Guianas and E Brazil.	Not listed.	Least Concern		Phyllostomus elongatus	1004981	23	Lesser Spear-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	PHYLLOSTOMINAE	PHYLLOSTOMINI	Phyllostomus	NA	elongatus	É. Geoffroy Saint-Hilaire	1810	1	Phyllostoma_elongatum	Geoffroy Saint-Hilaire, É. (1810). Sur les Phyllostomes et les Mégadermes, deux genres de la famille des Chauve-souris. Annales du Muséum d'histoire naturelle, 15, 182.	https://www.biodiversitylibrary.org/item/23421#page/206/mode/1up	MNHN 1986-1044		type locality not given. Restricted by A. Cabrera in 1958 to Rio Branco, Mato Grosso, Brazil.			elongatus (É. Geoffroy Saint-Hilaire, 1810)|ater (J. E. Gray, 1866)|lanceolatum (Pelzeln, 1883) [nomen nudum]	NA	NA				Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia	South America	Neotropic	LC	0	0	0	Phyllostomus_elongatus	0	sciname match	Phyllostomus_elongatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Phyllostomus_elongatus	1004981	23	Lesser Spear-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Phyllostominae	Phyllostomini	Phyllostomus	NA	elongatus	É. Geoffroy Saint-Hilaire	0	Phyllostoma elongatum	Geoffroy Saint-Hilaire, É. 1810. Sur les phyllostomes et les mégadermes, deux genres de la famille des chauve-souris. Annales du Muséum d'histoire naturelle 15:157-198.	https://www.biodiversitylibrary.org/page/3546867 | https://www.biodiversitylibrary.org/page/3546876	MNHN-ZM-MO-1986-1044 (= MNHN A 2)	holotype	http://coldb.mnhn.fr/catalognumber/mnhn/zm/mo-1986-1044	type locality not given. Restricted by A. Cabrera in 1958 to Rio Branco, Mato Grosso, Brazil.			NA	NA				Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia	South America	Neotropic	LC	0	0	0	Phyllostomus_elongatus	0	sciname match	Phyllostomus_elongatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Phyllostomus		elongatus	É. Geoffroy Saint-Hilaire	1810	1	Ann. Mus. Natn. Hist. Nat. Paris	0.751389	Lesser Spear-nosed Bat	ater Gray, 1866.	Brazil, Mato Grosso, Rio Branco.	Bolivia, E Peru, Ecuador, and Colombia to Guianas and E Brazil.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17217/22135836/' target='_blank'>Least Concern</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Phyllostomus elongatus; Phyllostomus elongatus; Phyllostomus elongatus; Phyllostomus elongatus; Phyllostomus elongatus; Phyllostomus elongatus; ater; ater; elongatus; ater; lanceolatum; Phyllostome des sous-bois; Kleine Lanzennase; Filostomapequeno; Lesser Spear-nosed Bat; Lesser Spear-nosed Bat; Lesser Spear-nosed Bat; P. elongatus