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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1141	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Phoniscus jagori	Phoniscus jagori	Phoniscus jagorii	Kerivoula jagori	Kerivoula jagorii	Phoniscus jagorii	Phoniscus jagorii	Phoniscus jagorii	Phoniscus jagorii	Phoniscus jagorii	Phoniscus jagorii	Phoniscus jagorii	Phoniscus jagorii	Phoniscus jagorii	Phoniscus jagorii		[MSW2] Subgenus Phoniscus.; [MSW3] See Hill (1965) and Kingston et al. (1997). Specimens from Laos are slightly smaller than those reported from elsewhere (Robinson and Webber, 2000), and may represent a distinct taxon. Sometimes spelled jagori.; [HMW] Vespertilio (Kerivoula) jagorii Peters, 1866 , “ Insel Samar,” Philippines . Specimens of P. jagorii from Laos are apparently slightly smaller than elsewhere and might be a distinct taxon. Up to three subspecies ( jagorii , javanus , and rapax) have been recognized, but additional studies are needed to determine their validity. Monotypic.; [IUCN] The taxonomy of this species requires review.				javanus, rapax		javana, rapax (see Hill, 1965:549-550).	javanus, jagorii, rapax		javana, rapax			jagorii 	jagorii - javana, rapax	jagorii, javana, rapax	The taxonomy of this species requires review.	jagorii 	jagorii - javana, rapax	jagorii, javana, rapax	jagorii, javanus, rapax, jagori	jagorii	jagorii - javana, rapax	jagorii (W. C. H. Peters, 1866)|javanus (O. Thomas, 1880)|rapax G. S. Miller, 1931|jagori Corbet & J. Edwards Hill, 1980 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Peter's trumpet-eared bat	Samar I, Philippines, Java, ? Celebes	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Phoniscus jagori	Philippines, Samar.	Peters	1866	Monatsb. Preuss. Akad. Wiss. Berlin, p. 399.	Distribution: Known from Java and Borneo to the Philippines and Celebes.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Peters' trumpet-eared bat	Samar I, Philippines, Borneo, Java, Sulawesi	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1866	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1866:399.	Subgenus Phoniscus.	Borneo, Java, Bali, Sulawesi, and Lesser Sunda Isis, Samar Isl (Philippines).	Philippines, Samar.		PETERS	1866	Angular concavity on posterior margin of ear near apex well developed. Anterior palatal emargination rounded. Basioccipital pits well developed. Size relatively large (forearm length, 37-39 mm; condylobasal length, 15-16 mm).	Distribution: Known from Java and Borneo to the Philippines and Celebes.	Three subspecies are recognized:	K.j.javanus (Java, Bali, Borneo), K. j. jagorii (Samar in the Philippines), K.j. rapax (Celebes).	100	species	K. jagorii	PETERS	1866	Phoniscus	subgenus	Kerivoula jagorii				Angular concavity on posterior margin of ear near apex well developed. Anterior palatal emargination rounded. Basioccipital pits well developed. Size relatively large (forearm length, 37-39 mm; condylobasal length, 15-16 mm).	Three subspecies are recognized:		18. K.jagorii (PETERS 1866).	18	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Kerivoulinae		Phoniscus jagorii	Phoniscus		jagorii	Peters	y	1866		Monatsb. K. Preuss. Akad. Wiss. Berlin	1866		399		Peters's Trumpet-eared Bat	Philippines, Samar Isl.	Laos; Peninsular Malaysia, Borneo, Java, Bali, Sulawesi, and Lesser Sunda Isls, Samar Isl (Philippines).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc) as Kerivoula jagori.	javana Thomas, 1880; rapax Miller, 1931.	See Hill (1965) and Kingston et al. (1997). Specimens from Laos are slightly smaller than those reported from elsewhere (Robinson and Webber, 2000), and may represent a distinct taxon. Sometimes spelled jagori.	4C3D87E8FF7C6AC3FF889CA91699BCF2	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	895	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF7C6AC3FF889CA91699BCF2.xml	Phoniscus jagorii	Vespertilionidae	Phoniscus	jagorii		1866	Kérivoule de Peters @fr | Peters-Trompetenohr @de | Fonisco de Peters @es | Common Trumpet-eared Bat @en | Frosted Groove-toothed Bat @en	Vespertilio (Kerivoula) jagorii Peters, 1866 , “ Insel Samar,” Philippines . Specimens of P. jagorii from Laos are apparently slightly smaller than elsewhere and might be a distinct taxon. Up to three subspecies ( jagorii , javanus , and rapax) have been recognized, but additional studies are needed to determine their validity. Monotypic.	E Thailand , Laos , NW & SW Vietnam , C Peninsular Malaysia (along with Penang I), NE & SW Borneo, W Java, Bali I, N Sulawesi, and Philippines (Samar and Bohol Is); possibly in N Cambodia .	Head-body 39-8 45- 2 mm , tail 36-49- 45 mm , ear 12- 3- 17 mm , hindfoot 8:5-9- 1 mm , forearm 35—42- 5 mm ; weight 5-8- 11 g . Dorsal pelage is mixed brown and black, with frosted gold throughout (hairs banded with dark gray-brown bases, middle buff and dark brown bands, and goldentips); venteris paler, with slightly grayertips. Short tiny yellow hairs are along bones of wings,tail, and legs. Ears are funnel-shaped, with rounded tips; tragusis long and conspicuously white, with sharply pointed tip and distinct and deep notch near base on posterior margin. Tail is long, and calcar is strong and curved. Basioccipital pits appear to be deeper than in the Groove-toothed Trumpet-eared Bat ( P. atrox ). I? is unicuspid and pointed; I? is distinctly smaller and shorter; C' has two distinct longitudinal lateral grooves on outer side; P* is broader than P° but shorter anteroposteriorly; P* is somewhat rectangular shaped with an anterointernal elongation; and lower premolars are narrow, especially P.	Primary and moderately disturbed semi-evergreen forests in limestone karst, rainforests, and dry dipterocarp forests at low elevations.	Peters’s Trumpet-eared Bats are maneuverable fliers and appearto forage in dense vegetation where preferred orb-weaving spiders live. They might become toxic themselves by sequestering poison from spider prey. Maximum bite force was 2:6 N in individuals from Malaysia .	No information.	Day roosts of Peters’s Trumpet-eared Bat appear most commonly in abandoned nests of broadbills (Eurylaimussp.); they make entrance holes in bottoms of the abandoned nests. Calls are broad FM sweep, with start frequencies of 137-140 kHz (mean 137-6 kHz), end frequencies of 77-85-7 kHz (82-9 kHz), peak frequencies of 87-5-88-2 kHz (88 kHz), and durations of 1:4—1-6 milliseconds. In Malaysia , a larger sample size reported start frequencies of 154-4-184-8 kHz (mean 169-3 kHz), end frequencies of 616-76 kHz (70-4 kHz), peak frequencies of 79-2-117-6 kHz (94-4 kHz), and durations of 1:6—3-2 milliseconds (2-2 milliseconds).	Peters’s Trumpet-eared Bat usually roosts alone but sometimes in small groups.	Classified as Least Concern on The IUCN Red List. Peters’s Trumpet-eared Bat is widespread but not particularly common. It does not currently seem to face any major threats, but deforestation is likely a problem in some regions.	Francis (2008a) | Heaney et al. (1998) | Hill (1965b) | Kingston , Francis, Rosell-Ambal & Tabaranza (2008) | Kingston , Francis, Zubaid & Kunz (2003) | Kingston , Kunz etal. (1997) | Kruskop (2013a) | Phillipps & Phillipps (2016) | Ryan (1965) | Senawi et al. (2015) | Thong Vu Dinh et al. (2006)	https://zenodo.org/record/6398434/files/figure.png	294. Peters’s Trumpet-eared Bat Phoniscus jagorii French: Kérivoule de Peters / German: Peters-Trompetenohr / Spanish: Fonisco de Peters Other common names: Common Trumpet-eared Bat , Frosted Groove-toothed Bat Taxonomy. Vespertilio (Kerivoula) jagorii Peters, 1866 , “ Insel Samar,” Philippines . Specimens of P. jagorii from Laos are apparently slightly smaller than elsewhere and might be a distinct taxon. Up to three subspecies ( jagorii , javanus , and rapax) have been recognized, but additional studies are needed to determine their validity. Monotypic. Distribution. E Thailand , Laos , NW & SW Vietnam , C Peninsular Malaysia (along with Penang I), NE & SW Borneo, W Java, Bali I, N Sulawesi, and Philippines (Samar and Bohol Is); possibly in N Cambodia . Descriptive notes. Head-body 39-8 45- 2 mm , tail 36-49- 45 mm , ear 12- 3- 17 mm , hindfoot 8:5-9- 1 mm , forearm 35—42- 5 mm ; weight 5-8- 11 g . Dorsal pelage is mixed brown and black, with frosted gold throughout (hairs banded with dark gray-brown bases, middle buff and dark brown bands, and goldentips); venteris paler, with slightly grayertips. Short tiny yellow hairs are along bones of wings,tail, and legs. Ears are funnel-shaped, with rounded tips; tragusis long and conspicuously white, with sharply pointed tip and distinct and deep notch near base on posterior margin. Tail is long, and calcar is strong and curved. Basioccipital pits appear to be deeper than in the Groove-toothed Trumpet-eared Bat ( P. atrox ). I? is unicuspid and pointed; I? is distinctly smaller and shorter; C' has two distinct longitudinal lateral grooves on outer side; P* is broader than P° but shorter anteroposteriorly; P* is somewhat rectangular shaped with an anterointernal elongation; and lower premolars are narrow, especially P. Habitat. Primary and moderately disturbed semi-evergreen forests in limestone karst, rainforests, and dry dipterocarp forests at low elevations. Food and Feeding. Peters’s Trumpet-eared Bats are maneuverable fliers and appearto forage in dense vegetation where preferred orb-weaving spiders live. They might become toxic themselves by sequestering poison from spider prey. Maximum bite force was 2:6 N in individuals from Malaysia . Breeding. No information. Activity patterns. Day roosts of Peters’s Trumpet-eared Bat appear most commonly in abandoned nests of broadbills (Eurylaimussp.); they make entrance holes in bottoms of the abandoned nests. Calls are broad FM sweep, with start frequencies of 137-140 kHz (mean 137-6 kHz), end frequencies of 77-85-7 kHz (82-9 kHz), peak frequencies of 87-5-88-2 kHz (88 kHz), and durations of 1:4—1-6 milliseconds. In Malaysia , a larger sample size reported start frequencies of 154-4-184-8 kHz (mean 169-3 kHz), end frequencies of 616-76 kHz (70-4 kHz), peak frequencies of 79-2-117-6 kHz (94-4 kHz), and durations of 1:6—3-2 milliseconds (2-2 milliseconds). Movements, Home range and Social organization. Peters’s Trumpet-eared Bat usually roosts alone but sometimes in small groups. Status and Conservation. Classified as Least Concern on The IUCN Red List. Peters’s Trumpet-eared Bat is widespread but not particularly common. It does not currently seem to face any major threats, but deforestation is likely a problem in some regions. Bibliography. Francis (2008a), Heaney et al. (1998), Hill (1965b), Kingston , Francis, Rosell-Ambal & Tabaranza (2008), Kingston , Francis, Zubaid & Kunz (2003), Kingston , Kunz etal. (1997), Kruskop (2013a), Phillipps & Phillipps (2016), Ryan (1965), Senawi et al. (2015), Thong Vu Dinh et al. (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Phoniscus jagorii	Phoniscus		jagorii	Peters	1866	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1872:39:00	Peters' Trumpet-eared Bat	 javana Thomas, 1880; rapax Miller, 1931.	Philippines, Samar Isl.	Laos; Peninsular Malaysia, Borneo, Java, Bali, Sulawesi, and Lesser Sunda Isls, Samar Isl (Philippines).	Not listed.	Least Concern		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Phoniscus jagorii	23	Peters's Trumpet-eared Bat	Common Trumpet-eared Bat|Frosted Groove-toothed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Phoniscus	NA	jagorii	W. Peters	1866	1	Vespertilio_(Kerivoula)_Jagorii	Peters, W. C. H. (1866). Mittheilung über neue oder ungenügend bekannte Flederthiere (Vampyrops, Uroderma, Chiroderma, Ametrida, Tylostoma, Vespertilio, Vesperugo) und Nager (Tylomys, Lasiomys). Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1866, 399.	https://www.biodiversitylibrary.org/item/120607#page/439/mode/1up	ZMB 2541		"Insel Samar," Philippines.			jagorii (W. Peters, 1866)|javana (O. Thomas, 1880)|rapax G. S. Miller, 1931	NA	NA	Thailand|Myanmar|Laos|Cambodia?|Vietnam|Malaysia|Indonesia|Philippines	Asia	Indomalaya	LC	0	0	0	Phoniscus_jagorii	0	sciname match	Phoniscus_jagorii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10976	Phoniscus jagorii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Phoniscus	jagorii	(Peters, 1866)	The taxonomy of this species requires review.	20000000	Phoniscus jagorii	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	It is associated with primary and moderately disturbed semi-evergreen forest in limestone karst areas. It has been recorded from primary forest in Malaysia (Kingston et al . 2003) and disturbed forest in Thailand (S. Bumrungsri pers. comm. 2006). One individual was collected from a degraded forest in Putao in the northern part of Myanmar in 2016.	There are no major threats to this species throughout its range, although deforestation may be affecting the species in parts of its range.	The status of the species is unknown in the Philippines and it is uncommon but widespread elsewhere.	Unknown	This species is widespread throughout Southeast Asia including mainland Thailand (Thong et al . 2006; S. Bumrungsri pers. comm. 2006), Malaysia (Kingston et al . 2006), Viet Nam (Thong et al . 2006), Lao PDR (Francis et al . 1999), Indonesia (found on Bali, Borneo, Java and Sulawesi), and the Philippines (known only from Samar) (Heaney et al . 1998). It can be expected in northern Cambodia, but this has not yet been confirmed. One individual was collected from a degraded forest in Putao in the northern part of Myanmar in 2016. The elevation is under 1,000 m asl (Oo et al. in prep). Moreover, this species was recorded in the Hkakaborazi Landscape in Myanmar and it is also included in the proposed Southern Extension of Hkakaborazi National Park.		Terrestrial	The species occurs in a number of protected areas throughout its range. Further studies are needed into the taxonomy, distribution, abundance, natural history and threats to this species.	Australasian|Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Phoniscus		jagorii	Peters	1866	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1872:39:00	Peters' Trumpet-eared Bat	 javana Thomas, 1880; rapax Miller, 1931.	Philippines, Samar Isl.	Laos; Peninsular Malaysia, Borneo, Java, Bali, Sulawesi, and Lesser Sunda Isls, Samar Isl (Philippines).	Not listed.	Least Concern		Phoniscus jagorii	1005312	23	Peters's Trumpet-eared Bat	Common Trumpet-eared Bat|Frosted Groove-toothed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Phoniscus	NA	jagorii	W. Peters	1866	1	Vespertilio_(Kerivoula)_Jagorii	Peters, W. C. H. (1866). Mittheilung über neue oder ungenügend bekannte Flederthiere (Vampyrops, Uroderma, Chiroderma, Ametrida, Tylostoma, Vespertilio, Vesperugo) und Nager (Tylomys, Lasiomys). Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1866, 399.	https://www.biodiversitylibrary.org/item/120607#page/439/mode/1up	ZMB 2541		"Insel Samar," Philippines.			jagorii (W. Peters, 1866)|javana (O. Thomas, 1880)|rapax G. S. Miller, 1931	NA	NA				Thailand|Myanmar|Laos|Cambodia?|Vietnam|Malaysia|Indonesia|Philippines	Asia	Indomalaya	LC	0	0	0	Phoniscus_jagorii	0	sciname match	Phoniscus_jagorii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Phoniscus_jagorii	1005312	23	Peters's Trumpet-eared Bat	Common Trumpet-eared Bat|Frosted Groove-toothed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Phoniscus	NA	jagorii	W. C. H. Peters	1	Vespertilio (Kerivoula) Jagorii	Peters, W.C.H. 1866. Hr. W. Peters machte eine Mittheilung über neue oder ungenügend bekannte Flederthiere (_Vampyrops_, _Uroderma_, _Chiroderma_, _Ametrida_, _Tylostoma_, _Vespertilio_, _Vesperugo_) und Nager (_Tylomys_, _Lasiomys_). Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1866:392-411.	https://www.biodiversitylibrary.org/page/39052961	ZMB 2541	holotype		"Insel Samar," Philippines.			NA	NA				Thailand|Myanmar|Laos|Cambodia?|Vietnam|Malaysia|Indonesia|Philippines	Asia	Indomalaya	LC	0	0	0	Phoniscus_jagorii	0	sciname match	Phoniscus_jagorii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Phoniscus		jagorii	Peters	1866	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1872:39:00	Peters' Trumpet-eared Bat	javana Thomas, 1880; rapax Miller, 1931.	Philippines, Samar Isl.	Laos; Peninsular Malaysia, Borneo, Java, Bali, Sulawesi, and Lesser Sunda Isls, Samar Isl (Philippines).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10976/21974660/' target='_blank'>Least Concern</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Phoniscus jagorii; Phoniscus jagorii; Phoniscus jagorii; Phoniscus jagorii; Phoniscus jagorii; Phoniscus jagorii; javana; rapax; javana; rapax; jagorii; javana; rapax; Kérivoule de Peters; Peters-Trompetenohr; Fonisco de Peters; Common Trumpet-eared Bat; Frosted Groove-toothed Bat; Peters's Trumpet-eared Bat; Common Trumpet-eared Bat; Frosted Groove-toothed Bat; Peters's Trumpet-eared Bat; Peters' Trumpet-eared Bat; P. jagorii