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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1137	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene	Pharotis imogene		[MSW3] See Flannery (1995a) and Bonaccorso (1998).; [HMW] Pharotis imogene Thomas, 1914 , “Kamali, Lower Kemp Welch River, British New Guinea [= Central Province, Papua New Guinea ].” Pharotis imogene was initially included under Nyctophilus timoriensis (species inquirenda = doubtful identity or name under investigation) by O. Thomas in 1897 until Thomas in 1914 proposed the genus Pharotis for wmogene. Phylogenetic relationships of Pharotis to Nyctophilus and within Vespertilionidae remain unclear. Both genera have been placed in their own subfamily Nyctophilinae or in subfamily Vespertilioninae , sometimes in the distinct tribe Nyctophilini or Vespertilionini . Pharotis has not yet been included in any phylogenetic study, but N. gouldi and N. geoffroyi have nested in Vespertilionini as sister to some species of Chalinolobus and Vespadelus . M. Volleth and C. R. Tidemann in 1991 suggested that Nyctophilus belongs in Vespertilionini based on karyotypic data. Despite uncertainty as to which tribe Pharotis (and Nyctophilus ) belongsin,it is listed here under Vespertilionini . More research is needed to determine the real relationship of Pharotis and Nyctophilus . Monotypic.; [batnames2022] Although previously believed extinct (see Flannery, 1995 a and Bonaccorso et al., 2008), a single individual of P. imogene was captured in 2012 in the Abau district of Papua New Guinea.; [batnames2023] Although previously believed extinct (see Flannery, 1995 a and Bonaccorso et al., 2008), a single individual of P. imogene was captured in 2012 in the Abau district of Papua New Guinea.; [batnames2025_1.7] Although previously believed extinct (see Flannery, 1995a and Bonaccorso et al., 2008), a single individual of P. imogene was captured in 2012 in the Abau district of Papua New Guinea.														imogene				imogene 	imogene 			imogene O. Thomas, 1914		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Big-eared bat	SE New Guinea	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pharotis imogene	Papua New Guinea, Central Prov., Lower Kemp Welch River, Kamali.	Thomas	1914	Ann. Mag. Nat. Hist., ser. 8, 14:382.	Distribution: Same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Big-eared bat	SE New Guinea	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1914	Ann. Mag. Nat. Hist., ser. 8, 14:382.		SE New Guinea.	Papua New Guinea, Central Prov., Lower Kemp Welch River, Kamali.		THOMAS	1914	Size fairly small (forearm length, 37-38 mm).	Distribution: Same as for genus.	No subspecies.		131	species	P. imogene	THOMAS	1914	Pharotis	genus	Pharotis imogene				Size fairly small (forearm length, 37-38 mm).	No subspecies.		1. P. imogene THOMAS 1914.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nyctophilini	Pharotis imogene	Pharotis		imogene	Thomas		1914		Ann. Mag. Nat. Hist., ser. 8	14		382		Thomas's Big-eared Bat	Papua New Guinea, Central Prov., Lower Kemp Welch River, Kamali.	SE New Guinea.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Critically Endangered.		See Flannery (1995a) and Bonaccorso (1998).	4C3D87E8FFD06A6FFF8397C7189BBF12	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	801	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFD06A6FFF8397C7189BBF12.xml	Pharotis imogene	Vespertilionidae	Pharotis	imogene	Thomas	1914	Nyctophile imogene @fr | Neuguinea-Langohr @de | Pharotis de Nueva Guinea @es | Thomas's Big-eared Bat @en	Pharotis imogene Thomas, 1914 , “Kamali, Lower Kemp Welch River, British New Guinea [= Central Province, Papua New Guinea ].” Pharotis imogene was initially included under Nyctophilus timoriensis (species inquirenda = doubtful identity or name under investigation) by O. Thomas in 1897 until Thomas in 1914 proposed the genus Pharotis for wmogene. Phylogenetic relationships of Pharotis to Nyctophilus and within Vespertilionidae remain unclear. Both genera have been placed in their own subfamily Nyctophilinae or in subfamily Vespertilioninae , sometimes in the distinct tribe Nyctophilini or Vespertilionini . Pharotis has not yet been included in any phylogenetic study, but N. gouldi and N. geoffroyi have nested in Vespertilionini as sister to some species of Chalinolobus and Vespadelus . M. Volleth and C. R. Tidemann in 1991 suggested that Nyctophilus belongs in Vespertilionini based on karyotypic data. Despite uncertainty as to which tribe Pharotis (and Nyctophilus ) belongsin,it is listed here under Vespertilionini . More research is needed to determine the real relationship of Pharotis and Nyctophilus . Monotypic.	Known only from two localities in Central Province, SE Papua New Guinea , including type locality and Oio village, Abau district, ¢. 120 km E of former. Former records are either misidentified Nyctophilus timoriensis or correspond to N. microdon .	Head—body 43-50-1 mm, tail 42-44 mm, ear 23-4-25 mm, hindfoot 7-8-9-4 mm, forearm 37-5-39-6 mm; weight 7-7 g (one specimen). Tibia lengths are 13-1-19 mm. Pharotis and Nyctophilus are the only genera of Vespertilionidae that combine large ears that arejoined by membranejust above forehead—except in the Papuan Long-eared Bat ( NN . microtis ) and the Pygmy Long-eared Bat ( NN . walkeri )—and presence of simple small noseleaf. Upperparts and underparts are dark brown. Wing membranes are uniformly brown and naked. Ears are brown, long, and bluntly pointed and have horizontally ribbing on inner surfaces. Tragus is irregularly lobed. Well-developed auricular band unites ears. Muzzle is strongly truncated and sparsely haired. Noseleaf has rounded anterior projecting flap and posterior cup oriented to anterior flap. Calcaris long (14-15 mm) and robust. Pharotis is distinguished from Nyctophilus as follows: convex dorsal margin of noseleaf above nostrils (vs. median concave margin), deep pit immediately posterior to posterior nasal prominence (vs. absent), lobe on inner margin of tragus located midway between each side of tragus (vs. located on outer margin of tragus), and skin between and above nostrils is completely naked (vs. covered with fine hairs). Baculum is similar to that of Nyctophilus (long and rather broad with scarcely expanded, bilobed base, and broad shaft tapering distally to blunt point, to produce shallow bifid tip; shaft is ventrally deeply fluted), but shaft is narrower, tapering to slightly expanded tip. Braincase is large, smooth with no ridges, and rounded without crests. Rostrum is very short, and nasal notch is deep. Interorbital region is very wide, with smoothly rounded edges. Tympanic bullae are medium-sized (c.3-5 mm); palate is shortened; greatest skull length is ¢. 15 mm ; and maxillary tooth row is c.4-5 mm. C' is slightly larger than that of the Small-toothed Long-eared Bat (N. microdon ); and upper incisors are noticeably procumbent and angled medially. Dental formulais11/3,C1/1,P1/2,M 3/3 (x2) = 30.	Uncertain but either savanna woodlands or lowland rainforest patches in savanna woodlands at upper elevation of ¢. 100 m . In Abau district, south-eastern Papua New Guinea , the single recorded female New Guinea Big-eared Bat was trapped in rainforest less than 100 m from boundary between extensive rainforest and large expanse of grassland (once a coconut plantation) and was within 12 km of the coast and extensive open country.	Morphology ofears,skull, and teeth strongly suggests that New Guinea Big-eared Bats forage by gleaning,likely on weevils, other beetles, ants, and moths.	No information.	New Guinea Big-eared Bats are nocturnal.	[Large number of specimens (45, with six females) originally collected suggests that it was probably a maternity colony. As a part of a broader study of impacts of selective rainforest logging on small mammals of the Cloudy Forest Management Area in Abau district, bats were captured with mist nets and harp traps, and their calls were recorded. A New Guinea Big-eared Bat was captured in this area together with Diadem Leaf-nosed Bats (Hipposideros diadema), Lesser Long-tongued Blossom Bats ( Macroglossus minimus), Large Long-fingered Bats ( Miniopterus magnater), Little Long-fingered Bats (M. australis ), Maluku Myotis ( Myotis moluccarum ), Papuan Long-eared Bats, New Guinea Pipistrelles ( Pipistrellus angulatus ), Lesser Papuan Pipistrelles ( P. papuanus ), and Watts’s Pipistrelles ( P. wattsi ).	Classified as Critically Endangered (Possibly Extinct) on The IUCN Red Last. If the New Guinea Big-eared Bat still exists,it is likely to have a very small population and small distribution that is subject to a continued decline in extent of occurrence, area of occupancy, and extent and quality of habitat. It is one of the top ten island endemic bat (IEB) species as identified by the research priority metric (RPM), ranked in order of research priority. New Guinea Big-eared Bats and Bulmer’s Fruit Bats ( Aproteles bulmerae ) are listed among the top 100 mammals in the EDGE (Evolutionary Distinct and Globally Endangered) program. The New Guinea Big-eared Bat is threatened by habitat loss around human population centers (e.g. Kamali, Central Province, Papua New Guinea ), reduction of area and quality of small rainforest patches in savanna woodlands, encroachmentof fire, and human activities, particularly firewood collection. It was rediscovered in 2012 (a single individual) from coastal district Abau, 120 km east of the only previous known locality at Kamali. C. Hughes and colleagues in 2014 recommend the following steps to addressit conservation status: bat surveys using harp traps in Abau district and adjoining lowland region to determine local distribution and abundance; whether it can be identified from echolocation calls; and radio-tracking during different stages of reproductive cycle to define habitat use, roosting requirements, and movements.	Amador etal. (2018) | Boitani et al. (2006) | Bonaccorso (1998) | Bonaccorso, Hamilton & Parnaby (2008a) | Broken-Brow & Hughes (2012) | Conenna et al. (2017) | Flannery (1995b) | Hill & Harrison (1987) | Hoofer & Van Den Bussche (2003) | Hughes et al. (2014) | Isaac et al. (2007) | Koopman (1993, 1994) | Koopman & Jones (1970) | Nowak (1999) | Roehrs et al. (2010) | Simmons (2005) | Tate (1941e, 1942b) | Thomas (1897b, 1914c) | Volleth & Tidemann (1991)	https://zenodo.org/record/6397934/files/figure.png	78. New Guinea Big-eared Bat Pharotis imogene French: Nyctophile imogene / German: Neuguinea-Langohr / Spanish: Pharotis de Nueva Guinea Other common names: Thomas's Big-eared Bat Taxonomy. Pharotis imogene Thomas, 1914 , “Kamali, Lower Kemp Welch River, British New Guinea [= Central Province, Papua New Guinea ].” Pharotis imogene was initially included under Nyctophilus timoriensis (species inquirenda = doubtful identity or name under investigation) by O. Thomas in 1897 until Thomas in 1914 proposed the genus Pharotis for wmogene. Phylogenetic relationships of Pharotis to Nyctophilus and within Vespertilionidae remain unclear. Both genera have been placed in their own subfamily Nyctophilinae or in subfamily Vespertilioninae , sometimes in the distinct tribe Nyctophilini or Vespertilionini . Pharotis has not yet been included in any phylogenetic study, but N. gouldi and N. geoffroyi have nested in Vespertilionini as sister to some species of Chalinolobus and Vespadelus . M. Volleth and C. R. Tidemann in 1991 suggested that Nyctophilus belongs in Vespertilionini based on karyotypic data. Despite uncertainty as to which tribe Pharotis (and Nyctophilus ) belongsin,it is listed here under Vespertilionini . More research is needed to determine the real relationship of Pharotis and Nyctophilus . Monotypic. Distribution. Known only from two localities in Central Province, SE Papua New Guinea , including type locality and Oio village, Abau district, ¢. 120 km E of former. Former records are either misidentified Nyctophilus timoriensis or correspond to N. microdon . Descriptive notes. Head—body 43-50-1 mm, tail 42-44 mm, ear 23-4-25 mm, hindfoot 7-8-9-4 mm, forearm 37-5-39-6 mm; weight 7-7 g (one specimen). Tibia lengths are 13-1-19 mm. Pharotis and Nyctophilus are the only genera of Vespertilionidae that combine large ears that arejoined by membranejust above forehead—except in the Papuan Long-eared Bat ( NN . microtis ) and the Pygmy Long-eared Bat ( NN . walkeri )—and presence of simple small noseleaf. Upperparts and underparts are dark brown. Wing membranes are uniformly brown and naked. Ears are brown, long, and bluntly pointed and have horizontally ribbing on inner surfaces. Tragus is irregularly lobed. Well-developed auricular band unites ears. Muzzle is strongly truncated and sparsely haired. Noseleaf has rounded anterior projecting flap and posterior cup oriented to anterior flap. Calcaris long (14-15 mm) and robust. Pharotis is distinguished from Nyctophilus as follows: convex dorsal margin of noseleaf above nostrils (vs. median concave margin), deep pit immediately posterior to posterior nasal prominence (vs. absent), lobe on inner margin of tragus located midway between each side of tragus (vs. located on outer margin of tragus), and skin between and above nostrils is completely naked (vs. covered with fine hairs). Baculum is similar to that of Nyctophilus (long and rather broad with scarcely expanded, bilobed base, and broad shaft tapering distally to blunt point, to produce shallow bifid tip; shaft is ventrally deeply fluted), but shaft is narrower, tapering to slightly expanded tip. Braincase is large, smooth with no ridges, and rounded without crests. Rostrum is very short, and nasal notch is deep. Interorbital region is very wide, with smoothly rounded edges. Tympanic bullae are medium-sized (c.3-5 mm); palate is shortened; greatest skull length is ¢. 15 mm ; and maxillary tooth row is c.4-5 mm. C' is slightly larger than that of the Small-toothed Long-eared Bat (N. microdon ); and upper incisors are noticeably procumbent and angled medially. Dental formulais11/3,C1/1,P1/2,M 3/3 (x2) = 30. Habitat. Uncertain but either savanna woodlands or lowland rainforest patches in savanna woodlands at upper elevation of ¢. 100 m . In Abau district, south-eastern Papua New Guinea , the single recorded female New Guinea Big-eared Bat was trapped in rainforest less than 100 m from boundary between extensive rainforest and large expanse of grassland (once a coconut plantation) and was within 12 km of the coast and extensive open country. Food and Feeding. Morphology ofears,skull, and teeth strongly suggests that New Guinea Big-eared Bats forage by gleaning,likely on weevils, other beetles, ants, and moths. Breeding. No information. Activity patterns. New Guinea Big-eared Bats are nocturnal. Movements, Home range and Social organization. [Large number of specimens (45, with six females) originally collected suggests that it was probably a maternity colony. As a part of a broader study of impacts of selective rainforest logging on small mammals of the Cloudy Forest Management Area in Abau district, bats were captured with mist nets and harp traps, and their calls were recorded. A New Guinea Big-eared Bat was captured in this area together with Diadem Leaf-nosed Bats (Hipposideros diadema), Lesser Long-tongued Blossom Bats ( Macroglossus minimus), Large Long-fingered Bats ( Miniopterus magnater), Little Long-fingered Bats (M. australis ), Maluku Myotis ( Myotis moluccarum ), Papuan Long-eared Bats, New Guinea Pipistrelles ( Pipistrellus angulatus ), Lesser Papuan Pipistrelles ( P. papuanus ), and Watts’s Pipistrelles ( P. wattsi ). Status and Conservation. Classified as Critically Endangered (Possibly Extinct) on The IUCN Red Last. If the New Guinea Big-eared Bat still exists,it is likely to have a very small population and small distribution that is subject to a continued decline in extent of occurrence, area of occupancy, and extent and quality of habitat. It is one of the top ten island endemic bat (IEB) species as identified by the research priority metric (RPM), ranked in order of research priority. New Guinea Big-eared Bats and Bulmer’s Fruit Bats ( Aproteles bulmerae ) are listed among the top 100 mammals in the EDGE (Evolutionary Distinct and Globally Endangered) program. The New Guinea Big-eared Bat is threatened by habitat loss around human population centers (e.g. Kamali, Central Province, Papua New Guinea ), reduction of area and quality of small rainforest patches in savanna woodlands, encroachmentof fire, and human activities, particularly firewood collection. It was rediscovered in 2012 (a single individual) from coastal district Abau, 120 km east of the only previous known locality at Kamali. C. Hughes and colleagues in 2014 recommend the following steps to addressit conservation status: bat surveys using harp traps in Abau district and adjoining lowland region to determine local distribution and abundance; whether it can be identified from echolocation calls; and radio-tracking during different stages of reproductive cycle to define habitat use, roosting requirements, and movements. Bibliography. Amador etal. (2018), Boitani et al. (2006), Bonaccorso (1998), Bonaccorso, Hamilton & Parnaby (2008a), Broken-Brow & Hughes (2012), Conenna et al. (2017), Flannery (1995b), Hill & Harrison (1987), Hoofer & Van Den Bussche (2003), Hughes et al. (2014), Isaac et al. (2007), Koopman (1993, 1994), Koopman & Jones (1970), Nowak (1999), Roehrs et al. (2010), Simmons (2005), Tate (1941e, 1942b), Thomas (1897b, 1914c), Volleth & Tidemann (1991).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pharotis imogene	Pharotis		imogene	Thomas	1914	0	Ann. Mag. Nat. Hist.	ser. 8, 14: 382	Thomas' Big-eared Bat	None.	Papua New Guinea, Central Prov., Lower Kemp Welch River, Kamali.	SE New Guinea.	Not listed.	Critically Endangered	Although previously believed extinct (see Flannery, 1995 a and Bonaccorso et al., 2008), a single individual of P. imogene was captured in 2012 in the Abau district of Papua New Guinea.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pharotis imogene	23	New Guinea Big-eared Bat	Thomas's Big-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Pharotis	NA	imogene	O. Thomas	1914	0	Pharotis_imogene	Thomas, O. (1914). A new genus of bats allied to Nyctophilus. Annals and Magazine of Natural History, ser. 8, 14, 382.	https://www.biodiversitylibrary.org/item/61720#page/394/mode/1up	BM 1891.9.10.2		"Kamali, Lower Kemp Welch River, British New Guinea [= Central Province, Papua New Guinea]."			imogene O. Thomas, 1914	NA	NA	Papua New Guinea	Oceania	Australasia/Oceania	CR	0	0	0	Pharotis_imogene	0	sciname match	Pharotis_imogene	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	16887	Pharotis imogene	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pharotis	imogene	Thomas, 1914		200000000	Pharotis imogene	Critically Endangered	D	2021	2019-07-18 00:00:00 UTC	3.1	English	Pharotis Imogene is listed as Critically Endangered under criterion D as the global population is currently conservatively estimated to be fewer than 50 mature individuals. The only evidence for its continued existence is a single individual collected in 2012.	Almost nothing is known about the ecology of Pharotis , but given their long ears and rudimentary noseleaf they are likely to be similar in ecology to Nyctophilus, using passive listening and low amplitude broadband echolocation calls to glean insects from vegetation. They roost communally or sometimes as individuals, probably in trees (Bonaccorso 1998). Roosting and foraging habitat is likely to be within lowland sclerophyll woodland and woodland with rainforest patches. The individual captured near Oio was trapped in rainforest near the boundary between extensive rainforest to the north and a large expanse of grassland, and was within 12 km of the coast (Hughes et al. 2014).	<p>This species is threatened by loss of woodland and forest habitat in general, especially in lowland regions near the coast. Around human population centres, including in the Kamali district where it has been recorded in the distant past, encroachment by fire, logging and human activities, particularly fire wood collection, are likely to be threatening processes (S. Hamilton pers. comm). </p>	<p>There has been an increase in the past decade of exploratory field surveys for bats across Papua New Guinea as part of biodiversity assessments for industry projects and conservation organisations, which have included the use of harp traps that are ideal for capturing long-eared bats. However, the record of Hughes et al. (2014) is the only indication of the persistence of Phraotis imogene . Further surveys are needed in similar lowland habitat to obtain a better understanding of its extent of occurrence and how common it might be locally.</p>	Stable	<p>The most recent capture of this species was from the sustainable rainforest logging concession, Cloudy Bay Forestry Management Area, near Oio Village, north-east of Abau, Central Province, Papua New Guinea (Hughes et al. 2014). Prior to this, there is only one other confirmed record of occurrence, when it was first discovered at Kamali on the lower Kemp Welch River, in the Central Province of south-eastern New Guinea in 1890 (Thomas 1897). A possible maternity colony of 45 individuals was collected by Dr. L. Loria, but only six of these specimens can now be accounted for (Flannery 1995, Hughes et al. 2014).  There are a further two records of this species mentioned in Bonaccorso (1998). A specimen was collected at Kapa Kapa by Dr. Loria, as reported by Thomas (1897), but it was initially identified as Nyctophilus timoriensis , and the identity was not confirmed as Pharotis prior to the specimen having been lost in flooding of the collection in the Museo Civico di Storia Naturale in Genoa. In addition, there is a report of a single specimen collected by T. Anderson in 1985 from the crown of a Pandanus tree in Rogut Village along the Tuman River in Central Province. This specimen (PM 25374 at the Papua New Guinea National Museum) is listed as being destroyed in 1992. Prior to this in 1988, careful examination and measurement of this specimen was undertaken by H. Parnaby as part of a revision of the genus Nyctophilus , which confirmed the identity as Nyctophilus microdon (Hughes et al. 2014). Both confirmed records of the species have been from below 100 m asl.</p>		Terrestrial	<p>This species has not been recorded from any protected areas. Further targeted field effort in suitable habitat is needed to build an understanding of extent of occurrence, and intensive study in the vicinity of its most recent capture would be useful to provide ecological data, including reference echolocation calls that might help with acoustics-based field surveys further afield. Areas known to contain roosting and foraging habitat should be protected.</p>	Australasian	1890	FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pharotis		imogene	Thomas	1914	0	Ann. Mag. Nat. Hist.	ser. 8, 14: 382	Thomas' Big-eared Bat	None.	Papua New Guinea, Central Prov., Lower Kemp Welch River, Kamali.	SE New Guinea.	Not listed.	Critically Endangered	Although previously believed extinct (see Flannery, 1995 a and Bonaccorso et al., 2008), a single individual of P. imogene was captured in 2012 in the Abau district of Papua New Guinea.	Pharotis imogene	1005768	23	New Guinea Big-eared Bat	Thomas's Big-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Pharotis	NA	imogene	O. Thomas	1914	0	Pharotis_imogene	Thomas, O. (1914). A new genus of bats allied to Nyctophilus. Annals and Magazine of Natural History, ser. 8, 14, 382.	https://www.biodiversitylibrary.org/item/61720#page/394/mode/1up	BM 1891.9.10.2		"Kamali, Lower Kemp Welch River, British New Guinea [= Central Province, Papua New Guinea]."			imogene O. Thomas, 1914	NA	NA				Papua New Guinea	Oceania	Australasia/Oceania	CR	0	0	0	Pharotis_imogene	0	sciname match	Pharotis_imogene	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Pharotis_imogene	1005768	23	New Guinea Big-eared Bat	Thomas's Big-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Pharotis	NA	imogene	O. Thomas	0	Pharotis imogene	Thomas, O. 1914-11-01. A new genus of bats allied to _Nyctophilus_. Annals and Magazine of Natural History (8)14(83):381-383.	https://www.biodiversitylibrary.org/page/18616617	BMNH:Mamm:1891.9.10.2	holotype	https://data.nhm.ac.uk/object/10e34094-3d78-4040-9fb7-36b4bbec78e5	"Kamali, Lower Kemp Welch River, British New Guinea [= Central Province, Papua New Guinea]."			NA	NA				Papua New Guinea	Oceania (Continent)	Australasia	CR	0	0	0	Pharotis_imogene	0	sciname match	Pharotis_imogene	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pharotis		imogene	Thomas	1914	0	Ann. Mag. Nat. Hist.	ser. 8, 14: 382	Thomas' Big-eared Bat	None.	Papua New Guinea, Central Prov., Lower Kemp Welch River, Kamali.	SE New Guinea.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/16887/209524728/' target='_blank'>Critically Endangered</a>	Although previously believed extinct (see Flannery, 1995a and Bonaccorso et al., 2008), a single individual of P. imogene was captured in 2012 in the Abau district of Papua New Guinea.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pharotis imogene; Pharotis imogene; Pharotis imogene; Pharotis imogene; Pharotis imogene; Pharotis imogene; imogene; Nyctophile imogene; Neuguinea-Langohr; Pharotis de Nueva Guinea; Thomas's Big-eared Bat; New Guinea Big-eared Bat; Thomas's Big-eared Bat; Thomas's Big-eared Bat; Thomas' Big-eared Bat; P. imogene