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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L114	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Carollia manu	Carollia manu	Carollia manu	Carollia manu	Carollia manu	Carollia manu	Carollia manu	Carollia manu	Carollia manu		[HMW] Carollia manu Pacheco, Solari & Velazco, 2004 , “Morro Leguia, Paucartambo-Pillcopata road, km. 134, 2250 m , Paucartambo Province, Cuzco Department, Peru , at approximately 13°11'52” S , 71°34'36" W .” Carolia manu was called Carolliasp.? (3) by R. H. Pine in 1972. In her morphometric analyses, L. J. McLellan in 1984 included Pine’s single specimen in C. perspicillata . Monotypic.; [MDD2022] recently described; [IUCN] This is a recently described species and is easily confused with C. perspicillata (Aguirre pers. comm.).; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														manu	This is a recently described species and is easily confused with C. perspicillata (Aguirre pers. comm.).			manu	manu			manu Pacheco, Solari, & Velazco, 2004						N/A																																								NA																											03A687BCFF85FF851396F88EF889F395	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	536	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF85FF851396F88EF889F395.xml	Carollia manu	Phyllostomidae	Carollia	manu	Pacheco, Solari & Velazco	2004	Carollia du Manu @fr | Manu-Kurzschwanzblattnase @de | Carolia de Manu @es	Carollia manu Pacheco, Solari & Velazco, 2004 , “Morro Leguia, Paucartambo-Pillcopata road, km. 134, 2250 m , Paucartambo Province, Cuzco Department, Peru , at approximately 13°11'52” S , 71°34'36" W .” Carolia manu was called Carolliasp.? (3) by R. H. Pine in 1972. In her morphometric analyses, L. J. McLellan in 1984 included Pine’s single specimen in C. perspicillata . Monotypic.	SE Peru ( Madre de Dios , Cusco , and Puno departments) and W & C Bolivia (La Paz Department).	Head-body 59-66 mm, tail 7-10 mm, ear 18-22 mm, hindfoot 13-15 mm, forearm 41-2-44-4 mm; weight 17-21 g. The Manu Short-tailed Batis a large species of Carollia , with soft, long, and fluffy dorsal fur. Dorsal hairs are tricolored, with grayish brown basal band, buff-to-whitish brown medial band, and brown band at tips. Ventral hairs are short, brown-tipped, and tricolored on pectoral region and bicolored on abdominal and inguinal region. Dorsal and ventral pelage is not counter shaded and looks brown overall. Forearm is long, and proximal one-halfis well furred. Uropatagium is wide, enclosing shorttail, and has a deep notch. Lower lip has central papilla surrounded by smaller warts in a U-shape. Ears are moderately large, broad, and triangular, with pointed tips. Rostrum is short and wide; anteorbital region is inflated, with low sagittal crest; and zygomatic arches are incomplete. C' is large and divergent. Upper and lower molars are broad and robust.	Tropical montane forests (Cloud forests), along eastern slope of Andes in south-eastern Peru and northern to central Bolivia , at elevations of 1300-2250 m. These forests have abundant epiphytes, hepatics ( Hepatica , Ranunculaceae ), and tree ferns. Most Manu Short-tailed Bats were collected on steep terrain, with mature and secondary forests, and usually near small streams. At higher elevations, forest trees were heavily covered with epiphytes, and understory was dominated by bamboo. Manu Short-tailed Bats have been found in sympatry with Silky Short-tailed Bats (C. brevicaudum) and Seba’s Short-tailed Bats ( C. perspicillata ).	There is no specific information available for this species, but fruits are expected to be the primary food of the Manu Short-tailed Bat.	No information.	No information.	No information.	Classified as Least Concern on The IUCN Red List. The Manu Short-tailed Bat might have a wider distribution than is currently known. It occurs in some protected areas in south-eastern Peru .	Castano et al. (2018) | McLellan (1984) | McLellan & Koopman (2008) | Pacheco et al. (2004) | Pine (1972) | Solari et al. (2006) | Velazco (2013)	https://zenodo.org/record/6458823/files/figure.png	106. Manu Short-tailed Bat Carollia manu French: Carollia du Manu / German: Manu-Kurzschwanzblattnase / Spanish: Carolia de Manu Taxonomy. Carollia manu Pacheco, Solari & Velazco, 2004 , “Morro Leguia, Paucartambo-Pillcopata road, km. 134, 2250 m , Paucartambo Province, Cuzco Department, Peru , at approximately 13°11'52” S , 71°34'36" W .” Carolia manu was called Carolliasp.? (3) by R. H. Pine in 1972. In her morphometric analyses, L. J. McLellan in 1984 included Pine’s single specimen in C. perspicillata . Monotypic. Distribution. SE Peru ( Madre de Dios , Cusco , and Puno departments) and W & C Bolivia (La Paz Department). Descriptive notes. Head-body 59-66 mm, tail 7-10 mm, ear 18-22 mm, hindfoot 13-15 mm, forearm 41-2-44-4 mm; weight 17-21 g. The Manu Short-tailed Batis a large species of Carollia , with soft, long, and fluffy dorsal fur. Dorsal hairs are tricolored, with grayish brown basal band, buff-to-whitish brown medial band, and brown band at tips. Ventral hairs are short, brown-tipped, and tricolored on pectoral region and bicolored on abdominal and inguinal region. Dorsal and ventral pelage is not counter shaded and looks brown overall. Forearm is long, and proximal one-halfis well furred. Uropatagium is wide, enclosing shorttail, and has a deep notch. Lower lip has central papilla surrounded by smaller warts in a U-shape. Ears are moderately large, broad, and triangular, with pointed tips. Rostrum is short and wide; anteorbital region is inflated, with low sagittal crest; and zygomatic arches are incomplete. C' is large and divergent. Upper and lower molars are broad and robust. Habitat. Tropical montane forests (Cloud forests), along eastern slope of Andes in south-eastern Peru and northern to central Bolivia , at elevations of 1300-2250 m. These forests have abundant epiphytes, hepatics ( Hepatica , Ranunculaceae ), and tree ferns. Most Manu Short-tailed Bats were collected on steep terrain, with mature and secondary forests, and usually near small streams. At higher elevations, forest trees were heavily covered with epiphytes, and understory was dominated by bamboo. Manu Short-tailed Bats have been found in sympatry with Silky Short-tailed Bats (C. brevicaudum) and Seba’s Short-tailed Bats ( C. perspicillata ). Food and Feeding. There is no specific information available for this species, but fruits are expected to be the primary food of the Manu Short-tailed Bat. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Manu Short-tailed Bat might have a wider distribution than is currently known. It occurs in some protected areas in south-eastern Peru . Bibliography. Castano et al. (2018), McLellan (1984), McLellan & Koopman (2008), Pacheco et al. (2004), Pine (1972), Solari et al. (2006), Velazco (2013).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Carollia manu	Carollia		manu	Pacheco, Solari, & Velazco	2004	0	Occas. Pap. Mus. Texas Tech Univ.	######	Manu Short-tailed Bat	None.	Morro Leguía, Paucartambo Province, Cuzco Department, Peru	Andean S Peru to N Bolivia	Not listed.	Least Concern		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Carollia manu	23	Manu Short-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	CAROLLIINAE	NA	Carollia	NA	manu	Pacheco, Solari, & Velazco	2004	0	Carollia_manu	Pacheco, V., Solari, S., & Velazco, P. M. (2004). A new species of Carollia (Chiroptera: Phyllostomidae) from the Andes of Peru and Bolivia. Occasional Papers, Museum of Texas Tech University, 236, 3.	https://www.biodiversitylibrary.org/item/263193#page/3/mode/1up	MUSM 8802		"Morro Leguía, Paucartambo-Pillcopata road, km. 134, 2250 m, Paucartambo Province, Cuzco Department, Peru, at approximately 13°11'52" S, 71°34'36" W."	-13.2	-71.58	manu Pacheco, Solari, & Velazco, 2004	recently described	Pacheco, V., Solari, S., & Velazco, P. M. (2004). A new species of Carollia (Chiroptera: Phyllostomidae) from the Andes of Peru and Bolivia. Occasional Papers, Museum of Texas Tech University, 236., 1-15.	Peru|Bolivia	South America	Neotropic	LC	0	0	0	Carollia_manu	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136782	Carollia manu	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Carollia	manu	Pacheco, Solari &; Velazco, 2004	This is a recently described species and is easily confused with C. perspicillata (Aguirre pers. comm.).	20000000	Carollia manu	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern. Although at present it does not have a wide distribution, it may occur more widely than currently known. It has presumed large population and the population trend is likely to be stable.	Little is known of the behaviour of this species, but it is probably similar to other members of the genus. It occurs in humid montane forest (Velazco and Aguirre pers. comm.)	Habitat destruction, due to coffee and coca plantations, may threaten the species in parts of its range; it occurs in a fragile ecosystem and is sensitive (Velazco and Aguirre pers. comm.).	It is possible that the population is stable (Velazco and Aguirre pers. comm.).	Stable	This species is known from Department of Cuzco, Madre de Dios, and Puno, in southeastern Peru, and the Department of La Paz in northern Bolivia. The elevational range is from 1,300 to 2,250 m asl (Pacheco et al. 2004).	This species is not used.	Terrestrial	It occurs in some protected areas in Peru, but not in Bolivia (Velazco and Aguirre pers. comm.).	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Carollia		manu	Pacheco, Solari, & Velazco	2004	0	Occas. Pap. Mus. Texas Tech Univ.	236:03:00	Manu Short-tailed Bat	None.	Morro Leguía, Paucartambo Province, Cuzco Department, Peru	Andean S Peru to N Bolivia	Not listed.	Least Concern		Carollia manu	1004875	23	Manu Short-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	CAROLLIINAE	NA	Carollia	NA	manu	Pacheco, Solari, & Velazco	2004	0	Carollia_manu	Pacheco, V., Solari, S., & Velazco, P. M. (2004). A new species of Carollia (Chiroptera: Phyllostomidae) from the Andes of Peru and Bolivia. Occasional Papers, Museum of Texas Tech University, 236, 3.	https://www.biodiversitylibrary.org/item/263193#page/3/mode/1up	MUSM 8802		"Morro Leguía, Paucartambo-Pillcopata road, km. 134, 2250 m, Paucartambo Province, Cuzco Department, Peru, at approximately 13°11'52" S, 71°34'36" W."	-13.1978	-71.5767	manu Pacheco, Solari, & Velazco, 2004	recently described	Pacheco, V., Solari, S., & Velazco, P. M. (2004). A new species of Carollia (Chiroptera: Phyllostomidae) from the Andes of Peru and Bolivia. Occasional Papers, Museum of Texas Tech University, 236., 1-15.				Peru|Bolivia	South America	Neotropic	LC	0	0	0	Carollia_manu	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Carollia_manu	1004875	23	Manu Short-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Carolliinae	NA	Carollia	NA	manu	Pacheco, Solari, & Velazco	0	Carollia manu	Pacheco, V.R., Solari, S. and Velazco, P.M. 2004. A new species of _Carollia_ (Chiroptera: Phyllostomidae) from the Andes of Peru and Bolivia. Occasional Papers, Museum of Texas Tech University 236:1-16.	https://www.biodiversitylibrary.org/page/57663618	MUSM 8802	holotype		"Morro Leguía, Paucartambo-Pillcopata road, km. 134, 2250 m, Paucartambo Province, Cuzco Department, Peru, at approximately 13°11'52" S, 71°34'36" W."	-13.1978	-71.5767	recently described	Pacheco, V., Solari, S., & Velazco, P. M. (2004). A new species of Carollia (Chiroptera: Phyllostomidae) from the Andes of Peru and Bolivia. Occasional Papers, Museum of Texas Tech University, 236., 1-15.				Peru|Bolivia	South America	Neotropic	LC	0	0	0	Carollia_manu	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Carollia		manu	Pacheco, Solari & Velazco	2004	0	Occas. Pap. Mus. Texas Tech Univ.	236:03:00	Manu Short-tailed Bat	None.	Morro Leguía, Paucartambo Province, Cuzco Department, Peru	Andean S Peru to N Bolivia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136782/22033116/' target='_blank'>Least Concern</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Carollia manu; Carollia manu; Carollia manu; Carollia manu; Carollia manu; manu; Carollia du Manu; Manu-Kurzschwanzblattnase; Carolia de Manu; Manu Short-tailed Bat; Manu Short-tailed Bat; C. manu