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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1129	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Paremballonura tiavato	Paremballonura tiavato	Paremballonura tiavato	Paremballonura tiavato	Paremballonura tiavato	Paremballonura tiavato	Paremballonura tiavato	Paremballonura tiavato	Paremballonura tiavato		[HMW] Emballonura tiavato Goodman et al., 2006, “ Madagascar : Province d’Antsiranana , Réserve Spéciale d’Ankarana, 2.6 km E of Andrafiabe, in forest near Andrafiabe Cave , 12°55.9’S , 49°03.4’E , ± 50 m . ” This species is monotypic.; [batnames2022] Formerly part of Emballonura , but transferred to the new genus Paraemballonura by Goodman et al. (2012).; [MDD2022] moved from Emballonura to Paremballonura; [IUCN] Recently described as a new species in a split from Emballonura atrata (Goodman et al. 2006).; [batnames2023] Formerly part of Emballonura , but transferred to the new genus Paraemballonura by Goodman et al. (2012).; [MDD2023] moved from Emballonura to Paremballonura; [MDD2025_2.0] moved from Emballonura to Paremballonura; [batnames2025_1.7] Formerly part of Emballonura, but transferred to the new genus Paraemballonura by Goodman et al. (2012).; [MDD2025_2.2] moved from Emballonura to Paremballonura														tiavato	Recently described as a new species in a split from Emballonura atrata (Goodman et al. 2006).			tiavato	tiavato			tiavato (S. M. Goodman, Cardiff, Ranivo, A. L. Russell, & Yoder, 2006)						N/A																																								NA																											03D587F2FFC34C09F8F73828FD72F82A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	361	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFC34C09F8F73828FD72F82A.xml	Paremballonura tiavato	Emballonuridae	Paremballonura	tiavato		2006	Western Sheath-tailed Bat @en | Emballonure des rochers @fr | Westliche Madagaskar-reischwanzfledermaus @de | Embalonuro occidental @es | Rock-dwelling Sheath-tailed Bat @en	Emballonura tiavato Goodman et al., 2006, “ Madagascar : Province d’Antsiranana , Réserve Spéciale d’Ankarana, 2.6 km E of Andrafiabe, in forest near Andrafiabe Cave , 12°55.9’S , 49°03.4’E , ± 50 m . ” This species is monotypic.	W Madagascar, from Daraina region to Tsingy de Bemaraha National Park, including Nosy Be and Nosy Romba, with an isolated record from Toliara at exfreme S of its distribution.	Head-body 40-46 mm, tail 15-18, ear 11-14 mm, hindfoot 5-6 mm, forearm 35-41 mm; weight 2-7-3-8 g. Dorsal fur of the Western Sheathtailed Bat is long, shaggy, and silky . It is uniformly pale to medium grayish brown. Basal one-quarter of hairs are lighter, approaching medium gray. Venter is paler buff-brown, with grayish brown cast, and basal one-third of hairs are distinctiy lighter and medium gray . Ear is long (11-15 mm), terminating with slightly pointed tip. Inner part of tragus is convex, with hatchet-shaped sharp anterior projection. Calcar is shorter than tibia. Cranium has distinct rostral expansion across nasal and maxilla bones. Postorbital processes are greatly reduced in size. Postorbital crest is not confluent with sagittal crest. Nasal bones are hourglass-shaped, with welldefined, deep, and broad nasal sulcus that terminates well before anterior margin of nasal. There is a narrow diastema between P1 and P2. Dental formula of species of ParembaUonura is I 2/3, C 1/1, P 2/2, M 3/3 (x2) = 34.	Lowland karst dry forests from sea level to elevations of c.350 m.	The Western Sheath-tailed Bat forages for insects, including moths, using a slow and delicate flight in forest understories and above small streams lined with vegetation.	Gestation ofWestern Sheath-tailed Bats occurs from December to February, and young are bom in February. All females examined had one embryo.	The Western Sheath-tailed Bat is crepuscular and begins to forage in twilight conditions in shaded forest understories. It roosts in cave entrances and rock overhangs in dim sunlight. It will use night roosts in buildings between bouts of foraging but does not use anthropomorphic structures for day roosts.	Roosting colonies of Western Sheath-tailed Bats are usually less than 20 individuals.	Classified as Least Concern on The IUCN Red List. The Western Sheath-tailed Bat currently has a large distribution, but it has experienced some population decline due to loss of foraging habitat. It is not abundant anywhere in its distribution, and close population monitoring and periodic reassessment are needed. It is threatened by habitat loss from slash-and-burn agriculture, logging, and charcoal collecting. Other potential threats include cave disturbance from tourism activities, fire, and mining. It is protected in Parc National Tsingy de Bemaraha, Réserve Spéciale d’Ankarana and Réserve Spéciale d’Analamerana.	Cardiff (2006) | Goodman, Andriafidison et al. (2005) | Goodman, Cardiff et al. (2006) | Jenkins et al. (2007) | Kofoky et al. (2007) | Monadjem , Cardiff et al. (2017b) | Peterson et al. (1995) | Razakarivony et al. (2005) | Robinson et al. (2006)	https://zenodo.org/record/3747950/files/figure.png	27 . Western Sheath-tailed Bat Paremballonura tiavato French: Emballonure des rochers /German: Westliche Madagaskar-reischwanzfledermaus / Spanish: Embalonuro occidental Other common names: Rock-dwelling Sheath-tailed Bat Taxonomy . Emballonura tiavato Goodman et al., 2006, “ Madagascar : Province d’Antsiranana , Réserve Spéciale d’Ankarana, 2.6 km E of Andrafiabe, in forest near Andrafiabe Cave , 12°55.9’S , 49°03.4’E , ± 50 m . ” This species is monotypic. Distribution. W Madagascar, from Daraina region to Tsingy de Bemaraha National Park, including Nosy Be and Nosy Romba, with an isolated record from Toliara at exfreme S of its distribution. Descriptive notes. Head-body 40-46 mm, tail 15-18, ear 11-14 mm, hindfoot 5-6 mm, forearm 35-41 mm; weight 2-7-3-8 g. Dorsal fur of the Western Sheathtailed Bat is long, shaggy, and silky . It is uniformly pale to medium grayish brown. Basal one-quarter of hairs are lighter, approaching medium gray. Venter is paler buff-brown, with grayish brown cast, and basal one-third of hairs are distinctiy lighter and medium gray . Ear is long (11-15 mm), terminating with slightly pointed tip. Inner part of tragus is convex, with hatchet-shaped sharp anterior projection. Calcar is shorter than tibia. Cranium has distinct rostral expansion across nasal and maxilla bones. Postorbital processes are greatly reduced in size. Postorbital crest is not confluent with sagittal crest. Nasal bones are hourglass-shaped, with welldefined, deep, and broad nasal sulcus that terminates well before anterior margin of nasal. There is a narrow diastema between P1 and P2. Dental formula of species of ParembaUonura is I 2/3, C 1/1, P 2/2, M 3/3 (x2) = 34. Habitat . Lowland karst dry forests from sea level to elevations of c.350 m. Food and Feeding . The Western Sheath-tailed Bat forages for insects, including moths, using a slow and delicate flight in forest understories and above small streams lined with vegetation. Breeding . Gestation ofWestern Sheath-tailed Bats occurs from December to February, and young are bom in February. All females examined had one embryo. Activity patterns. The Western Sheath-tailed Bat is crepuscular and begins to forage in twilight conditions in shaded forest understories. It roosts in cave entrances and rock overhangs in dim sunlight. It will use night roosts in buildings between bouts of foraging but does not use anthropomorphic structures for day roosts. Movements, Home range and Social organization. Roosting colonies of Western Sheath-tailed Bats are usually less than 20 individuals. Status and Conservation . Classified as Least Concern on The IUCN Red List. The Western Sheath-tailed Bat currently has a large distribution, but it has experienced some population decline due to loss of foraging habitat. It is not abundant anywhere in its distribution, and close population monitoring and periodic reassessment are needed. It is threatened by habitat loss from slash-and-burn agriculture, logging, and charcoal collecting. Other potential threats include cave disturbance from tourism activities, fire, and mining. It is protected in Parc National Tsingy de Bemaraha, Réserve Spéciale d’Ankarana and Réserve Spéciale d’Analamerana. Bibliography. Cardiff (2006), Goodman, Andriafidison et al. (2005), Goodman, Cardiff et al. (2006), Jenkins et al. (2007), Kofoky et al. (2007), Monadjem , Cardiff et al. (2017b), Peterson et al. (1995), Razakarivony et al. (2005), Robinson et al. (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Paremballonura tiavato	Paremballonura		tiavato	Goodman, Cardiff, Ranivo, Russell & Yoder	2006	1	Am. Mus. Novitates	######	Rock-loving Sheath-tailed Bat	None.	Madagascar, Province d&#39;Antisiranana, Reserve Special d&#39;Ankarana	NW Madagascar	Not listed.	Least Concern	Formerly part of Emballonura , but transferred to the new genus Paraemballonura by Goodman et al. (2012).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Paremballonura tiavato	23	Western Sheath-tailed Bat	Rock-dwelling Sheath-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	EMBALLONURINAE	EMBALLONURINI	Paremballonura	NA	tiavato	Goodman, Cardiff, Ranivo, Russel, & Yoder	2006	1	Emballonura_tiavato	Goodman, S. M., Cardiff, S. G., Ranivo, J., Russell, A. L., & A. D. Yoder. (2006). A new species of Emballonura (Chiroptera: Emballonuridae) from the dry regions of Madagascar. American Museum Novitates, 3538, 6.	https://bioone.org/journals/american-museum-novitates/volume-2006/issue-3538/0003-0082(2006)3538[1:ANSOEC]2.0.CO;2/A-New-Species-of-Emballonura-Chiroptera--Emballonuridae-from-the/10.1206/0003-0082(2006)3538[1:ANSOEC]2.0.CO;2.short	FMNH 169705		"Madagascar: Province d'Antsiranana, Réserve Spéciale d'Ankarana, 2.6 km E of Andrafiabe, in forest near Andrafiabe Cave, 12°55.9'S, 49°03.4'E, ± 50 m."	-12.93	49.057	tiavato (Goodman, Cardiff, Ranivo, Russel, & Yoder, 2006)	moved from Emballonura to Paremballonura	Goodman, S.M., Puechmaille, S.J., Friedli-Weyeneth, N., Gerlach, J., Ruedi, M., Schoeman, M.C., Stanley, W.T., Teeling, E.C. 2012. Phylogeny of Emballonurini (Emballonuridae) with descriptions of a new genus and species from Madagascar. Journal of Mammalogy 93(6): 1440-1455.	Madagascar	Africa	Afrotropic	LC	0	0	0	Paremballonura_tiavato	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136835	Paremballonura tiavato	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Paremballonura	tiavato	Goodman, Cardiff, Ranivo, Russell &; Yoder, 2006	Recently described as a new species in a split from Emballonura atrata (Goodman et al. 2006).	20000000	Paremballonura tiavato	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	This bat is listed as Least Concern because although this species is declining due to the impacts of ongoing forest loss due to shifting agriculture and logging the reduction of suitable habitat is not thought to be sufficient to warrant listing in a higher category of threat. However, this species is not abundant within its range and requires close population monitoring and a reassessment may be needed in the near future.	It appears to be associated with relatively intact forest and is believed to be forest dependent (Goodman et al. 2005, 2006). It is usually found roosting in the entrances to narrow caves and overhangs on rocks that receive weak sunlight (Cardiff 2006, Goodman et al . 2006, Kofoky et al . 2007). At night it has been observed resting on buildings (Goodman et al . 2006). It feeds in forest understorey and over small streams with bank side vegetation (Robinson et al . 2006). Its diet includes Lepidoptera (Razakarivony et al . 2005).	The species is threatened by habitat loss due to slash-and-burn agriculture (tavy) and from charcoal collecting and logging. Other potential threats come from the disturbance of roosting caves from tourists (Kofoky et al. 2007), fire (Jenkins et al . 2007) or mining (Cardiff 2006).	Roosting colonies of P. tiavato are usually relatively small (&lt; 20 individuals).	Unknown	This recently described species is endemic to Madagascar (Goodman et al. 2006). It is restricted to the karstic lowland areas in the western part of the island, extending from the drier areas in the north in Diana Region to Parc National Tsingy de Bemaraha in the west (Goodman et al . 2005). A record from Toliara in the extreme south has also been attributed to this species (Peterson et al . 1995), but the specimen was not examined as part of the recent reassessment of Emballonura taxonomy on Madagascar (Goodman et al . 2006). The distribution of this species would be significantly reduced if the Toliara specimen was omitted. It is currently known to range between 10 and 330 m elevation (Goodman et al . 2006).		Terrestrial	It is known from three protected areas, Parc National Tsingy de Bemaraha, Réserve Spéciale d’Ankarana, Réserve Spéciale d'Analamerana (Goodman et al. 2005).	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Paremballonura		tiavato	Goodman, Cardiff, Ranivo, Russell & Yoder	2006	1	Am. Mus. Novitates	########	Rock-loving Sheath-tailed Bat	None.	Madagascar, Province d&#39;Antisiranana, Reserve Special d&#39;Ankarana	NW Madagascar	Not listed.	Least Concern	Formerly part of Emballonura , but transferred to the new genus Paraemballonura by Goodman et al. (2012).	Paremballonura tiavato	1004812	23	Western Sheath-tailed Bat	Rock-dwelling Sheath-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	EMBALLONURINAE	EMBALLONURINI	Paremballonura	NA	tiavato	Goodman, Cardiff, Ranivo, Russel, & Yoder	2006	1	Emballonura_tiavato	Goodman, S. M., Cardiff, S. G., Ranivo, J., Russell, A. L., & A. D. Yoder. (2006). A new species of Emballonura (Chiroptera: Emballonuridae) from the dry regions of Madagascar. American Museum Novitates, 3538, 6.	https://bioone.org/journals/american-museum-novitates/volume-2006/issue-3538/0003-0082(2006)3538[1:ANSOEC]2.0.CO;2/A-New-Species-of-Emballonura-Chiroptera--Emballonuridae-from-the/10.1206/0003-0082(2006)3538[1:ANSOEC]2.0.CO;2.short	FMNH 169705		"Madagascar: Province d'Antsiranana, Réserve Spéciale d'Ankarana, 2.6 km E of Andrafiabe, in forest near Andrafiabe Cave, 12°55.9'S, 49°03.4'E, ± 50 m."	-12.9317	49.05667	tiavato (Goodman, Cardiff, Ranivo, Russel, & Yoder, 2006)	moved from Emballonura to Paremballonura	Goodman, S.M., Puechmaille, S.J., Friedli-Weyeneth, N., Gerlach, J., Ruedi, M., Schoeman, M.C., Stanley, W.T., Teeling, E.C. 2012. Phylogeny of Emballonurini (Emballonuridae) with descriptions of a new genus and species from Madagascar. Journal of Mammalogy 93(6): 1440-1455.				Madagascar	Africa	Afrotropic	LC	0	0	0	Paremballonura_tiavato	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Paremballonura_tiavato	1004812	23	Western Sheath-tailed Bat	Rock-dwelling Sheath-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Emballonurinae	Emballonurini	Paremballonura	NA	tiavato	S. M. Goodman, Cardiff, Ranivo, A. L. Russell, & Yoder	1	Emballonura tiavato	Goodman, S.M., Cardiff, S.G., Ranivo, J., Russell, A.L. and Yoder, A.D. 2006-10-19. A new species of _Emballonura_ (Chiroptera, Emballonuridae) from the dry regions of Madagascar. American Museum Novitates 3538:1-24.	https://www.biodiversitylibrary.org/page/59824373	FMNH:Mamm:169705	holotype	https://collections-zoology.fieldmuseum.org/catalogue/2668473	"Madagascar: Province d'Antsiranana, Réserve Spéciale d'Ankarana, 2.6 km E of Andrafiabe, in forest near Andrafiabe Cave, 12°55.9'S, 49°03.4'E, ± 50 m."	-12.9317	49.05667	moved from Emballonura to Paremballonura	Goodman, S.M., Puechmaille, S.J., Friedli-Weyeneth, N., Gerlach, J., Ruedi, M., Schoeman, M.C., Stanley, W.T., Teeling, E.C. 2012. Phylogeny of Emballonurini (Emballonuridae) with descriptions of a new genus and species from Madagascar. Journal of Mammalogy 93(6): 1440-1455.				Madagascar	Africa	Afrotropic	LC	0	0	0	Paremballonura_tiavato	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Paremballonura		tiavato	Goodman, Cardiff, Ranivo, Russell & Yoder	2006	1	Am. Mus. Novitates	########	Rock-loving Sheath-tailed Bat	None.	Madagascar, Province d&#39;Antisiranana, Reserve Special d&#39;Ankarana	NW Madagascar	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136835/22040708/' target='_blank'>Least Concern</a>	Formerly part of Emballonura, but transferred to the new genus Paraemballonura by Goodman et al. (2012).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Paremballonura tiavato; Paremballonura tiavato; Paremballonura tiavato; Paremballonura tiavato; Paremballonura tiavato; tiavato; Western Sheath-tailed Bat; Emballonure des rochers; Westliche Madagaskar-reischwanzfledermaus; Embalonuro occidental; Rock-dwelling Sheath-tailed Bat; Western Sheath-tailed Bat; Rock-dwelling Sheath-tailed Bat; Rock-loving Sheath-tailed Bat; P. tiavato