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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1128	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Emballonura atrata	Emballonura atrata	Emballonura atrata	Emballonura atrata	Emballonura atrata	Emballonura atrata	Paremballonura atrata	Paremballonura atrata	Paremballonura atrata	Paremballonura atrata	Paremballonura atrata	Paremballonura atrata	Paremballonura atrata	Paremballonura atrata	Paremballonura atrata		[MSW3] atrata species group. Reviewed by Peterson et al. (1995).; [HMW] Emballonura atrata Peters, 1874 , “ aus dem Innern von Madagascar [= interior of Madagascar ].” This species is monotypic.; [batnames2022] Formerly part of Emballonura , but transferred to Paraemballonura by Goodman et al. (2012). Reviewed by Peterson et al. (1995).; [MDD2022] moved from Emballonura to Paremballonura; [IUCN] Following a recent taxonomic revision based on genetic and morphological comparisons, this species is now confined to the east of Madagascar, with the smaller E. tiavato occurring in the west (Goodman et al. 2006).; [batnames2023] Formerly part of Emballonura , but transferred to Paraemballonura by Goodman et al. (2012). Reviewed by Peterson et al. (1995).; [MDD2023] moved from Emballonura to Paremballonura; [MDD2025_2.0] moved from Emballonura to Paremballonura; [batnames2025_1.7] Formerly part of Emballonura, but transferred to Paraemballonura by Goodman et al. (2012). Reviewed by Peterson et al. (1995).; [MDD2025_2.2] moved from Emballonura to Paremballonura														atrata	Following a recent taxonomic revision based on genetic and morphological comparisons, this species is now confined to the east of Madagascar, with the smaller E. tiavato occurring in the west (Goodman et al. 2006).			atrata 	atrata 			atrata (W. C. H. Peters, 1875)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Peters' sheath-tailed bat	Madagascar	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Emballonura atrata	Madagascar.	Peters	1874	Monatsb. Preuss. Akad. Wiss. Berlin, p. 693.	Distribution: Confined to Madagas car.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Peters' sheath-tailed bat	Madagascar	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1874	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1874:693.		E and C Madagascar.	Madagascar.		PETERS	1874	Inner margin of tragus convex. Calcar longer than tibia. Size relatively small (forearm length, 37-39 mm). Postorbital crests not confluent with sagittal crest. Basisphenoid pits of medium depth and separated by a median septum, but not extending into ali sphenoids.	Distribution: Confined to Madagas car.	No subspecies.		44	species	E. atrata	PETERS	1874	Emballonura	subgenus	Emballonura atrata				Inner margin of tragus convex. Calcar longer than tibia. Size relatively small (forearm length, 37-39 mm). Postorbital crests not confluent with sagittal crest. Basisphenoid pits of medium depth and separated by a median septum, but not extending into ali sphenoids.	No subspecies.		2. E. atrata PETERS 1874 [atrata group],	2	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Emballonuridae	Emballonurinae		Emballonura atrata	Emballonura		atrata	Peters		1874		Monatsb. K. Preuss. Akad. Wiss. Berlin	1874		693		Peters's Sheath-tailed Bat	Madagascar, restricted to "interior of Madagascar" by Peterson et al. (1995).	Madagascar except for S region.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		atrata species group. Reviewed by Peterson et al. (1995).	03D587F2FFC24C09FF773D45F787FE30	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	362	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFC24C09FF773D45F787FE30.xml	Paremballonura atrata	Emballonuridae	Paremballonura	atrata		1875	Emballonure de Madagascar @fr | Östliche Madagaskar-reischwanzfledermaus @de | Embalonuro de Peters @es | German @en	Emballonura atrata Peters, 1874 , “ aus dem Innern von Madagascar [= interior of Madagascar ].” This species is monotypic.	E Madagascar, from Maroantsetra S to Tolagnaro (= Fort Dauphin), including Nosy Mangabé and Sainte Marie I.	Head-body 45-49 mm, tail 18-20 mm, ear 15-19 mm, hindfoot C .6 mm, forearm 37—41 mm; weight 5—7-1 g. Dorsal fur of Peters’s Sheathtailed Bat is uniformly dark slate-gray to black; venter is slightly paler than dorsum. Snout is quite pointed and extends well beyond lower lip. Ears are prominent, broad, and rounded, with distinct indentation near tips. Inner surface of pinna is heavily ribbed. Tragus is longer than broad, with rounded smooth margin. Eyes are large, with dark brown irises. Flight membranes are dark black-brown.	In or near to humid rainforest forests from sea level to elevations of C.900 m.	Peters’s Sheath-tailed Bat forages for insects within 8 m of the ground along forest edges and low over rivers.	Gestation of Peters’s Sheath-tailed Bats occurs in November, and females have one embryo. An observation of copulation in a limestone cave was reported from March.	Peters’s Sheath-tailed Bats roost in caves and small rock crevices and below exposed tree roots. In eastern central highlands, it also roosts in village houses. The species exclusively uses twilight zones near cave entrances.	Colony size rarely exceeds 30 individuals and more typically is 3-10 individuals; however, groups of 80—120 can occur. Peters’s Sheath-tailed Bat occurs in caves with Rufous Trident Bats (Triaenops menamend), Major’s Long-fingered Bats (Miniopterus majori), Lesser Long-fingered Bats ( M.fraterculus ) , and Glen’s Long-fingered Bats ( M gleni )— all of which inhabit deeper dark areas of caves.	Classified as Least Concern on The IUCN Red List. Peters’s Sheath-tailed Bat is relatively widespread in eastern Madagascar. Its population is decreasing, and its status should be periodically reviewed given that it is collected for bushmeat in eastern Madagascar and forest clearing for expanding agriculture threatens foraging and roosting sites. Peters’s Sheath-tailed Bat is protected in Parc National de Mantadia, Parc National de Midongy du Sud, and a network of protected sites around Baie de Antogil. It likely does not require specific conservation action as long as intact humid forests are protected, but local populations should be monitored at roosting sites.	Garbutt (2007) | Goodman, Cardiff et al. (2006) | Goodman, Puechmaille, Hizem et al. (2012) | Jenkins et al. (2007) | Kofoky et al. (2007) | Monadjem, Cardiff et al. (2017 a) | Randrianandrianinà st al. (2006) | Peters, 1867.)	https://zenodo.org/record/3747952/files/figure.png	28 . Peters’s Sheath-tailed Bat Paremballonura atrata French : Emballonure de Madagascar / German : Östliche Madagaskar-reischwanzfledermaus / Spanish : Embalonuro de Peters Taxonomy . Emballonura atrata Peters, 1874 , “ aus dem Innern von Madagascar [= interior of Madagascar ].” This species is monotypic. Distribution. E Madagascar, from Maroantsetra S to Tolagnaro (= Fort Dauphin), including Nosy Mangabé and Sainte Marie I. Descriptive notes. Head-body 45-49 mm, tail 18-20 mm, ear 15-19 mm, hindfoot C .6 mm, forearm 37—41 mm; weight 5—7-1 g. Dorsal fur of Peters’s Sheathtailed Bat is uniformly dark slate-gray to black; venter is slightly paler than dorsum. Snout is quite pointed and extends well beyond lower lip. Ears are prominent, broad, and rounded, with distinct indentation near tips. Inner surface of pinna is heavily ribbed. Tragus is longer than broad, with rounded smooth margin. Eyes are large, with dark brown irises. Flight membranes are dark black-brown. Habitat . In or near to humid rainforest forests from sea level to elevations of C.900 m. Food and Feeding . Peters’s Sheath-tailed Bat forages for insects within 8 m of the ground along forest edges and low over rivers. Breeding . Gestation of Peters’s Sheath-tailed Bats occurs in November, and females have one embryo. An observation of copulation in a limestone cave was reported from March. Activity patterns. Peters’s Sheath-tailed Bats roost in caves and small rock crevices and below exposed tree roots. In eastern central highlands, it also roosts in village houses. The species exclusively uses twilight zones near cave entrances. Movements, Home range and Social organization. Colony size rarely exceeds 30 individuals and more typically is 3-10 individuals; however, groups of 80—120 can occur. Peters’s Sheath-tailed Bat occurs in caves with Rufous Trident Bats (Triaenops menamend), Major’s Long-fingered Bats (Miniopterus majori), Lesser Long-fingered Bats ( M.fraterculus ) , and Glen’s Long-fingered Bats ( M gleni )— all of which inhabit deeper dark areas of caves. Status and Conservation . Classified as Least Concern on The IUCN Red List. Peters’s Sheath-tailed Bat is relatively widespread in eastern Madagascar. Its population is decreasing, and its status should be periodically reviewed given that it is collected for bushmeat in eastern Madagascar and forest clearing for expanding agriculture threatens foraging and roosting sites. Peters’s Sheath-tailed Bat is protected in Parc National de Mantadia, Parc National de Midongy du Sud, and a network of protected sites around Baie de Antogil. It likely does not require specific conservation action as long as intact humid forests are protected, but local populations should be monitored at roosting sites. Bibliography. Garbutt (2007), Goodman, Cardiff et al. (2006), Goodman, Puechmaille, Hizem et al. (2012), Jenkins et al. (2007), Kofoky et al. (2007), Monadjem, Cardiff et al. (2017 a ), Randrianandrianinà st al. (2006). Peters, 1867	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Paremballonura atrata	Paremballonura		atrata	Peters	1874	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1885:33:00	Peters's Sheath-tailed Bat	None.	Madagascar, restricted to "interior of Madagascar" by Peterson et al. (1995).	Madagascar except for S region.	Not listed.	Least Concern	Formerly part of Emballonura , but transferred to Paraemballonura by Goodman et al. (2012). Reviewed by Peterson et al. (1995).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Paremballonura atrata	23	Peters's Sheath-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	EMBALLONURINAE	EMBALLONURINI	Paremballonura	NA	atrata	W. Peters	1874	1	Emballonura_atrata	Peters, W. C. H. (1874). Mittheilung über eine neue Gattung un d zwei neue Arten von Säugethieren aus Madagascar. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1874, 693.	https://www.biodiversitylibrary.org/item/112817#page/779/mode/1up	ZMB 4692		‘‘aus dem Innern von Madagascar [= interior of Madagascar]."			atrata (W. Peters, 1874)	moved from Emballonura to Paremballonura	Goodman, S.M., Puechmaille, S.J., Friedli-Weyeneth, N., Gerlach, J., Ruedi, M., Schoeman, M.C., Stanley, W.T., Teeling, E.C. 2012. Phylogeny of Emballonurini (Emballonuridae) with descriptions of a new genus and species from Madagascar. Journal of Mammalogy 93(6): 1440-1455.	Madagascar	Africa	Afrotropic	LC	0	0	0	Paremballonura_atrata	0	oldname match	Emballonura_atrata	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	7671	Paremballonura atrata	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Paremballonura	atrata	(Peters, 1874)	Following a recent taxonomic revision based on genetic and morphological comparisons, this species is now confined to the east of Madagascar, with the smaller E. tiavato occurring in the west (Goodman et al. 2006).	20000000	Paremballonura atrata	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	This species is listed as Least Concern because it is relatively widespread in eastern Madagascar and although its population is certainly decreasing as the humid forest is converted into agricultural areas there is no evidence that the decrease has been rapid enough to warrant Near Threatened or threatened status.	Paremballonura atrata is usually found roosting in caves and small crevices in or near to relatively intact humid forest (Goodman et al. 2006, Randrianandrianina et al . 2006, Kofoky et al . 2007, Jenkins et al . 2007). It is considered to be dependent on relatively intact forest (Goodman et al . 2005, 2006).	This species is collected by people for bushmeat in one part of eastern Madagascar (D. Andriafidison pers. comm). Otherwise, forest clearance for expanding agriculture threatens the foraging and roosting sites for this species (Jenkins et al. 2007).	There is no information on population size. This species is rarely trapped in flight or detected acoustically and information on populations and distribution is mainly from bats found in caves. Colony size in caves rarely exceeds 20 individuals (Jenkins et al. 2007).	Unknown	This species is restricted to the humid zone of eastern Madagascar and has been recorded from around Maroantsetra in the north to Tolagnaro (Fort Dauphin) in the south (Goodman et al. 2006). It is known from a wide elevational range, from as low as 30 m (Goodman et al . 2006) to at least 900 m (Randrianandriananina et al . 2006).		Terrestrial	This species has been recorded from Parc National de Mantadia (Randrianandrianina et al. 2006) and Parc National de Midongy du Sud (Goodman et al . 2006) as well as from a network of protected sites around the Baie de Antogil (Goodman et al . 2006). It probably does not require specific conservation action as long as forest vegetation is protected, but local populations should be monitored at roosting sites.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Paremballonura		atrata	Peters	1874	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1885:33:00	Peters's Sheath-tailed Bat	None.	Madagascar, restricted to "interior of Madagascar" by Peterson et al. (1995).	Madagascar except for S region.	Not listed.	Least Concern	Formerly part of Emballonura , but transferred to Paraemballonura by Goodman et al. (2012). Reviewed by Peterson et al. (1995).	Paremballonura atrata	1004811	23	Peters's Sheath-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	EMBALLONURINAE	EMBALLONURINI	Paremballonura	NA	atrata	W. Peters	1874	1	Emballonura_atrata	Peters, W. C. H. (1874). Mittheilung über eine neue Gattung un d zwei neue Arten von Säugethieren aus Madagascar. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1874, 693.	https://www.biodiversitylibrary.org/item/112817#page/779/mode/1up	ZMB 4692		‘‘aus dem Innern von Madagascar [= interior of Madagascar]."			atrata (W. Peters, 1874)	moved from Emballonura to Paremballonura	Goodman, S.M., Puechmaille, S.J., Friedli-Weyeneth, N., Gerlach, J., Ruedi, M., Schoeman, M.C., Stanley, W.T., Teeling, E.C. 2012. Phylogeny of Emballonurini (Emballonuridae) with descriptions of a new genus and species from Madagascar. Journal of Mammalogy 93(6): 1440-1455.				Madagascar	Africa	Afrotropic	LC	0	0	0	Paremballonura_atrata	0	oldname match	Emballonura_atrata	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Paremballonura_atrata	1004811	23	Peters's Sheath-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Emballonurinae	Emballonurini	Paremballonura	NA	atrata	W. C. H. Peters	1	Emballonura atrata	Peters, W.C.H. 1875. Ãœber eine neue Gattung und zwei neue Arten von Säugethieren aus Madagascar. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1874:690-694.	https://www.biodiversitylibrary.org/page/36627059	ZMB 4692	holotype		‘‘aus dem Innern von Madagascar [= interior of Madagascar]."			moved from Emballonura to Paremballonura	Goodman, S.M., Puechmaille, S.J., Friedli-Weyeneth, N., Gerlach, J., Ruedi, M., Schoeman, M.C., Stanley, W.T., Teeling, E.C. 2012. Phylogeny of Emballonurini (Emballonuridae) with descriptions of a new genus and species from Madagascar. Journal of Mammalogy 93(6): 1440-1455.				Madagascar	Africa	Afrotropic	LC	0	0	0	Paremballonura_atrata	0	oldname match	Emballonura_atrata	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Paremballonura		atrata	Peters	1874	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1885:33:00	Peters's Sheath-tailed Bat	None.	Madagascar, restricted to "interior of Madagascar" by Peterson et al. (1995).	Madagascar except for S region.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/7671/22135427/' target='_blank'>Least Concern</a>	Formerly part of Emballonura, but transferred to Paraemballonura by Goodman et al. (2012). Reviewed by Peterson et al. (1995).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Emballonura atrata; Paremballonura atrata; Paremballonura atrata; Paremballonura atrata; Paremballonura atrata; Paremballonura atrata; atrata; Emballonure de Madagascar; Östliche Madagaskar-reischwanzfledermaus; Embalonuro de Peters; German; Peters's Sheath-tailed Bat; Peters's Sheath-tailed Bat; Peters's Sheath-tailed Bat; P. atrata