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authorityParentheses originalNameCombination authoritySpeciesCitation authoritySpeciesLink typeVoucher typeKind typeVoucherURIs typeLocality typeLocalityLatitude typeLocalityLongitude taxonomyNotes taxonomyNotesCitation distributionNotes distributionNotesCitation subregionDistribution countryDistribution continentDistribution biogeographicRealm iucnStatus extinct domestic flagged CMW_sciName diffSinceCMW MSW3_matchtype MSW3_sciName diffSinceMSW3 docOrigin_batnames2025 Family Genus Subgenus Species Author Date Parentheses (1=author & date in parentheses) Citation Pages Common Name Synonyms Type Locality Distribution CITES IUCN Comments column3781 column3791 subtribe CONCAT_ALTNAMES line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1127 application/vnd.openxmlformats-officedocument.spreadsheetml.sheet N/A N/A N/A N/A N/A N/A Paratriaenops pauliani Paratriaenops pauliani Paratriaenops pauliani Paratriaenops pauliani Paratriaenops pauliani Paratriaenops pauliani Paratriaenops pauliani Paratriaenops pauliani Paratriaenops pauliani [HMW] Triaenopr paulianí Goodman 8c Ranivo, 2008 , “Aldabra Atoll, Picard lsland, c. 09 24 ’ S , 46 12 ’ E ," Seychelles . Although known for a century, I’. pauliani wasjust described in 2008; it was formerly included in Rfunula. Monotypic.; [MDD2022] recently described; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae; [IUCN] On the basis of a few available specimens from the western Seychelles, this recently described species was previously identified as the Malagasy species Triaenops furculus or T. auritus , which today are placed in the genus Paratriaenops. On the basis of morphological data it has been shown to be distinct from P. furculus and P. auritus , and, hence, endemic to the western Seychelles (Goodman and Ranivo 2008).; [MDD2023] recently described; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae; [MDD2025_2.0] recently described; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae; [MDD2025_2.2] recently described; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae furinea, pauliani On the basis of a few available specimens from the western Seychelles, this recently described species was previously identified as the Malagasy species Triaenops furculus or T. auritus , which today are placed in the genus Paratriaenops. On the basis of morphological data it has been shown to be distinct from P. furculus and P. auritus , and, hence, endemic to the western Seychelles (Goodman and Ranivo 2008). furinea, pauliani furinea, pauliani furinea (Tate, 1941) [nomen nudum]|pauliani (S. M. Goodman & Ranivo, 2008) N/A NA 0383245F2224977B8B0BF635F70EF2B9 Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Rhinonycteridae_194.pdf.imf hash://md5/ffba5c272223977c8e73f509ffb1ffd0 209 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/83/24/0383245F2224977B8B0BF635F70EF2B9.xml Paratriaenops pauliani Rhinonycteridae Paratriaenops pauliani Goodman & Ranivo 2008 Triaenops de Paulian @fr | Pauliari-Dreizahnblattnase @de | Rinonicteno de Paulian @es Triaenopr paulianí Goodman 8c Ranivo, 2008 , “Aldabra Atoll, Picard lsland, c. 09 24 ’ S , 46 12 ’ E ," Seychelles . Although known for a century, I’. pauliani wasjust described in 2008; it was formerly included in Rfunula. Monotypic. Known only from Picard Aldabra Atoll, Western Seychelles . Tail 17-21 mm, ear 14-& 15-4 mm, hindfoot 5-7-6-2 mm, forearm 41 -4-45-1 mm; weight 6 -5 g (single individual). No speciﬁc data are available for hindfoot length. Paulian's Trident Bat is the smallest species of Pamlnkıenops Pelage has slightly darker tinge dorsally than ventrally, being pale gray, brownish gray, or dull gray. Noseleaf is large (length 3-4-4 -0 mm), relatively broad, and rounded, with straplike, longitudinal, and anteriorly shallowly bifurcated projection, acres anterior part of anterior leaf, above anterior medial emargination. Three tall pointed projections are present on posterior part of posterior leaf, its anterior part has in medial position short, laterally ﬂattened, slightly upward directed projection. Tom] width of three posterior projections is almost entire width of posterior leaf. Posterior leaf is satlloped by numerous cells, small and more or less equal in size, ananged in two transversal rows. separated from each other by ﬂeshy septa, except central pair in poste rior row, which are separated by incomplete septum. Tall posterior projections are not emarginated at their bases, lateral two projections are slightly curved medially, and medial projection is markedly broader at its basis than lateral projections. There are no lateral supplementary leaﬂets, but thickened ridge occurs beneath margin of anterior leaf. Fara are large and pointed, and anterior margin is evenly rounded. Skull has very prominent nasal swellings laterally and dorsally. Braincase is slightly higher than rostrum, and there is depression between rostrum and braincase, with low but long sagittal crest. Wide plates are present on dorsal sides of zygomatic bones. Greatest skull length is 17-1 mm and zygo matic width is 8 -7 mm (single individual). F is bilobed; C' is slender, with slight cingulum; posterior secondary cusp is represented by ridge extending one-third the crown height; P' is small and extruded; C' and P‘ are in or nearly in contact; and M‘ is only little reduced, with metacone. is bicuspid, 1,2 is tricuspid and larger than ll, C, is slender, P? has about twothirds the crown area and one-half the height of P., and M, is unneduced. Baculum has not been described, but it is expected to be same general fonn and size of congeners. No information. Paulian's Trident Bat is nocturnal. Although not conﬁrmed, day roosts of Paulian's Trident Bat are expected to occur in sea caves, cavities in coral cliffs along the seacoast, and crevices in mushroom-shaped formations (Champignons) of derived coral rocks. No information. Classiﬁed as Data Deﬁcient on 771e IUCN Rai List. Paulian`s Trident Bat is only confirmed on the small Picard lsland (9 -3 km’), and all records come from the vicinity of an abandoned settlement or the research station of the Seychelles lslands Foundation. The Aldabra Atoll (155-4 km’) is a Nature Reserve and a UNESCO World Heritage Site and still represents one of the last remaining intact ecosystems in the world. Despite the enonnous rarity of Paulian`s Trident Bat. which mightjustify classiﬁcation as Endangered, it is not under an imminent threat due to the strict ecosystem protection from the Seychelles government. was reported to occur in the Cosmoledo Atoll (Western Seychelles ), c. 115 km east-southeast of the Aldabra Atoll, but these reports are considered incorrect, supposedly caused by mislabeling of museum specimens collected by]. G. F. Fryer during the Percy Sladen Trust Expedition to the atolls of the Western Seychelles in 1908 and 1909. Paulian's Trident Bat is known only from six museum specimens: four collected in 1908-1909 and two in 1977 and 2004. An observation of one individual in 1992 without further details is also available. Nevertheless, it is possible that Paulian`s Trident Bat inhabits most of or even the entire Aldabra Atoll and notjust the 6 % of its area represented by Picard lsland. von Brandıs (2004) | Goodman à Ramvo (2008) | Hutson (2004) https://zenodo.org/record/6611845/files/figure.png 9. Paulian ’ s Trident Bat Paratriaenops pauliani French: Triaenops de Paulian / German: Pauliari-Dreizahnblattnase / Spanish: Rinonicteno de Paulian Other common nunca Pauharfs Triple Lent-nosed Bat Taxonomy. Triaenopr paulianí Goodman 8c Ranivo, 2008 , “Aldabra Atoll, Picard lsland, c. 09 24 ’ S , 46 12 ’ E ," Seychelles . Although known for a century, I’. pauliani wasjust described in 2008; it was formerly included in Rfunula. Monotypic. Distribution. Known only from Picard Aldabra Atoll, Western Seychelles . Descriptive notes. Tail 17-21 mm, ear 14-& 15-4 mm, hindfoot 5-7-6-2 mm, forearm 41 -4-45-1 mm; weight 6 -5 g (single individual). No speciﬁc data are available for hindfoot length. Paulian's Trident Bat is the smallest species of Pamlnkıenops Pelage has slightly darker tinge dorsally than ventrally, being pale gray, brownish gray, or dull gray. Noseleaf is large (length 3-4-4 -0 mm), relatively broad, and rounded, with straplike, longitudinal, and anteriorly shallowly bifurcated projection, acres anterior part of anterior leaf, above anterior medial emargination. Three tall pointed projections are present on posterior part of posterior leaf, its anterior part has in medial position short, laterally ﬂattened, slightly upward directed projection. Tom] width of three posterior projections is almost entire width of posterior leaf. Posterior leaf is satlloped by numerous cells, small and more or less equal in size, ananged in two transversal rows. separated from each other by ﬂeshy septa, except central pair in poste rior row, which are separated by incomplete septum. Tall posterior projections are not emarginated at their bases, lateral two projections are slightly curved medially, and medial projection is markedly broader at its basis than lateral projections. There are no lateral supplementary leaﬂets, but thickened ridge occurs beneath margin of anterior leaf. Fara are large and pointed, and anterior margin is evenly rounded. Skull has very prominent nasal swellings laterally and dorsally. Braincase is slightly higher than rostrum, and there is depression between rostrum and braincase, with low but long sagittal crest. Wide plates are present on dorsal sides of zygomatic bones. Greatest skull length is 17-1 mm and zygo matic width is 8 -7 mm (single individual). F is bilobed; C' is slender, with slight cingulum; posterior secondary cusp is represented by ridge extending one-third the crown height; P' is small and extruded; C' and P‘ are in or nearly in contact; and M‘ is only little reduced, with metacone. is bicuspid, 1,2 is tricuspid and larger than ll, C, is slender, P? has about twothirds the crown area and one-half the height of P., and M, is unneduced. Baculum has not been described, but it is expected to be same general fonn and size of congeners. Habitat. Known only from autochthonous dry shrubland of various fonnations and introduced coconut plantations. Paulian ‘ Trident Bat is conﬁned Picard s reportedly to lsland, with semiarid climate and rainy season in November-April. Food and Feeding Feeding. Paulian ‘ Trident Bat is insectivorous. Breeding. No information. Activity patterns. Paulian's Trident Bat is nocturnal. Although not conﬁrmed, day roosts of Paulian's Trident Bat are expected to occur in sea caves, cavities in coral cliffs along the seacoast, and crevices in mushroom-shaped formations (Champignons) of derived coral rocks. Movements, Home range and Social organization. No information. Status and Conservation. Classiﬁed as Data Deﬁcient on 771e IUCN Rai List. Paulian`s Trident Bat is only confirmed on the small Picard lsland (9 -3 km’), and all records come from the vicinity of an abandoned settlement or the research station of the Seychelles lslands Foundation. The Aldabra Atoll (155-4 km’) is a Nature Reserve and a UNESCO World Heritage Site and still represents one of the last remaining intact ecosystems in the world. Despite the enonnous rarity of Paulian`s Trident Bat. which mightjustify classiﬁcation as Endangered, it is not under an imminent threat due to the strict ecosystem protection from the Seychelles government. was reported to occur in the Cosmoledo Atoll (Western Seychelles ), c. 115 km east-southeast of the Aldabra Atoll, but these reports are considered incorrect, supposedly caused by mislabeling of museum specimens collected by]. G. F. Fryer during the Percy Sladen Trust Expedition to the atolls of the Western Seychelles in 1908 and 1909. Paulian's Trident Bat is known only from six museum specimens: four collected in 1908-1909 and two in 1977 and 2004. An observation of one individual in 1992 without further details is also available. Nevertheless, it is possible that Paulian`s Trident Bat inhabits most of or even the entire Aldabra Atoll and notjust the 6 % of its area represented by Picard lsland. Bibliography. von Brandıs (2004), Goodman à Ramvo (2008), Hutson (2004) Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022. Rhinonycteridae Paratriaenops pauliani Paratriaenops pauliani Goodman & Ranivo 2008 1 Zoosystema 30(3): 685 Paulian's Trident Bat None. W Seychelles Aldabra Atoll, Picard Island, Seychelles Not listed. Data Deficient Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Paratriaenops pauliani 23 Paulian's Trident Bat Paulian's Triple Leaf-nosed Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA RHINONYCTERIDAE NA NA Paratriaenops NA pauliani Goodman & Ranivo 2008 1 Triaenops_pauliani Goodman, S. M., & Ranivo, J. (2008). A new species of Triaenops (Mammalia, Chiroptera, Hipposideridae) from Aldabra Atoll, Picard Island (Seychelles). Zoosystema, 30(3), 685. https://sciencepress.mnhn.fr/en/periodiques/zoosystema/30/3/une-nouvelle-espece-de-triaenops-mammalia-chiroptera-hipposideridae-de-l-atoll-d-aldabra-ile-picard-seychelles BM 1913.2.18.1 "Aldabra Atoll, Picard Island, c. 09Â°24'S, 46Â°12'E," Seychelles. -9.4 46.2 furinea (Tate, 1941) [nomen nudum]|pauliani (Goodman & Ranivo, 2008) recently described; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae Goodman, S. M., & Ranivo, J. (2008). A new species of Triaenops (Mammalia, Chiroptera, Hipposideridae) from Aldabra Atoll, Picard Island (Seychelles). Zoosystema, 30(3), 681-693.|Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333. Seychelles Africa Afrotropic DD 0 0 0 Paratriaenops_pauliani 0 unmatched NA 1 IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022]. 80000000 Paratriaenops pauliani ANIMALIA CHORDATA MAMMALIA CHIROPTERA HIPPOSIDERIDAE Paratriaenops pauliani Goodman et Ranivo, 2008 On the basis of a few available specimens from the western Seychelles, this recently described species was previously identified as the Malagasy species Triaenops furculus or T. auritus , which today are placed in the genus Paratriaenops. On the basis of morphological data it has been shown to be distinct from P. furculus and P. auritus , and, hence, endemic to the western Seychelles (Goodman and Ranivo 2008). 100000000 Paratriaenops pauliani Data Deficient 2017 2016-05-02 00:00:00 UTC 3.1 English

This newly described species is known from two different islands in the extreme western Seychelles, which include the Aldabra and Cosmoledo Atolls. These two atolls are separate by over 100 km of seas and the occurrence of this species on the latter atoll has been questioned (Goodman and Ranivo 2008). As nothing is known about this species' ecology and most records (observations and specimens) are of animals in human structures on Aldabra Atoll, further field data is needed to properly assess this taxon. ;

As the vast majority of records are based on individuals that entered human-built structures, little is known and the habitat use and ecology of this species. The main threats to this species are not known. Population size and trends are not known, but based on very irregular records on Aldabra Atoll (Brandis 2004, Hutson 2004), presumably it is not very common Unknown This species is only known from Aldabra Atoll, specifically Picard Island. The collection locality of specimens obtained on the nearby Cosmoledo Atoll has been called into question (Goodman and Ranivo 2008). Terrestrial The Aldabra Atoll has the status of a UNESCO World Heritage site since late 1982 and as such, its terrestrial habitats are well protected. Afrotropical FALSE FALSE Global Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Rhinonycteridae Paratriaenops pauliani Goodman & Ranivo 2008 1 Zoosystema 30(3): 685 Paulian's Trident Bat None. W Seychelles Aldabra Atoll, Picard Island, Seychelles Not listed. Data Deficient Paratriaenops pauliani 1004764 23 Paulian's Trident Bat Paulian's Triple Leaf-nosed Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA Rhinonycteridae NA NA Paratriaenops NA pauliani Goodman & Ranivo 2008 1 Triaenops_pauliani Goodman, S. M., & Ranivo, J. (2008). A new species of Triaenops (Mammalia, Chiroptera, Hipposideridae) from Aldabra Atoll, Picard Island (Seychelles). Zoosystema, 30(3), 685. https://sciencepress.mnhn.fr/en/periodiques/zoosystema/30/3/une-nouvelle-espece-de-triaenops-mammalia-chiroptera-hipposideridae-de-l-atoll-d-aldabra-ile-picard-seychelles BM 1913.2.18.1 "Aldabra Atoll, Picard Island, c. 09Â°24'S, 46Â°12'E," Seychelles. -9.4 46.2 furinea (Tate, 1941) [nomen nudum]|pauliani (Goodman & Ranivo, 2008) recently described; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae Goodman, S. M., & Ranivo, J. (2008). A new species of Triaenops (Mammalia, Chiroptera, Hipposideridae) from Aldabra Atoll, Picard Island (Seychelles). Zoosystema, 30(3), 681-693.|Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333. Seychelles Africa Afrotropic DD 0 0 0 Paratriaenops_pauliani 0 unmatched NA 1 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Paratriaenops_pauliani 1004764 23 Paulian's Trident Bat Paulian's Triple Leaf-nosed Bat Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yinpterochiroptera NA NA Rhinolophoidea Rhinonycteridae NA NA Paratriaenops NA pauliani S. M. Goodman & Ranivo 1 Triaenops pauliani Goodman, S.M. and Ranivo, J. 2008. A new species of _Triaenops_ (Mammalia, Chiroptera, Hipposideridae) from Aldabra Atoll, Picard Island (Seychelles). Zoosystema 30(3):681-693. https://sciencepress.mnhn.fr/en/periodiques/zoosystema/30/3/une-nouvelle-espece-de-triaenops-mammalia-chiroptera-hipposideridae-de-l-atoll-d-aldabra-ile-picard-seychelles BMNH:Mamm:1913.2.18.1 holotype https://data.nhm.ac.uk/object/2192b2e8-0bca-4c28-b6fa-ba508f90ee41 "Aldabra Atoll, Picard Island, c. 09Â°24'S, 46Â°12'E," Seychelles. -9.4 46.2 recently described; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae Goodman, S. M., & Ranivo, J. (2008). A new species of Triaenops (Mammalia, Chiroptera, Hipposideridae) from Aldabra Atoll, Picard Island (Seychelles). Zoosystema, 30(3), 681-693.|Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333. Seychelles Africa Afrotropic DD 0 0 0 Paratriaenops_pauliani 0 unmatched NA 1 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Rhinonycteridae Paratriaenops pauliani Goodman & Ranivo 2008 1 Zoosystema 30(3): 685 Paulian's Trident Bat None. W Seychelles Aldabra Atoll, Picard Island, Seychelles Not Listed Data Deficient Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Paratriaenops pauliani; Paratriaenops pauliani; Paratriaenops pauliani; Paratriaenops pauliani; Paratriaenops pauliani; furinea; pauliani; Triaenops de Paulian; Pauliari-Dreizahnblattnase; Rinonicteno de Paulian; Paulian's Trident Bat; Paulian's Triple Leaf-nosed Bat; Paulian's Trident Bat; P. pauliani