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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1126	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Triaenops furculus	Triaenops furculus	Triaenops furculus	Triaenops furculus	Triaenops furculus	Triaenops furculus	Paratriaenops furcula	Paratriaenops furcula	Paratriaenops furcula	Paratriaenops furculus	Paratriaenops furcula	Paratriaenops furcula	Paratriaenops furcula	Paratriaenops furcula	Paratriaenops furcula		[MSW2] Includes aurita; see Hayman and Hill (1971:30).; [MSW3] Does not includes auritus; see Peterson et al. (1995), but also see Hayman and Hill (1971). Originally spelled furcula but emended to furculus by Hill (1982), presumably to agree in gender with the generic epithet. May include furinea Tate, 1941, possibly a lapsus for furcula (see discussion in Hill, 1982).; [HMW] Tfiaenops fiımıla Trouessart, 1906 , “Grotte de Sarondrana (baie Saint-Augustin), cote Ouest de Madagascar [St. Augustine Bay, east coast of Madagascar ]. Widely used specific epithet ƒumdus has been replaced by the original epithet jinrula because it is a noun in apposition and does not need to be changed. Monotypic.; [batnames2022] Does not include auritus ; see Peterson et al. (1995), but also see Hayman and Hill (1971). Originally spelled furcula but emended to furculus by Hill (1982), presumably to agree in gender with the generic epithet. However, HMW pointed out that this correction was uneccessary as furcula is a noun in apposition and therefore does not change endings to agree with the genus name. May include furinea Tate, 1941, possibly a lapsus for furcula (see discussion in Hill, 1982).; [MDD2022] generally spelt furculus, but the name has been changed to furcula to match the generic gender; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae; [batnames2023] Does not include auritus ; see Peterson et al. (1995), but also see Hayman and Hill (1971). Originally spelled furcula but emended to furculus by Hill (1982), presumably to agree in gender with the generic epithet. However, HMW pointed out that this correction was uneccessary as furcula is a noun in apposition and therefore does not change endings to agree with the genus name. May include furinea Tate, 1941, possibly a lapsus for furcula (see discussion in Hill, 1982).; [MDD2023] generally spelt furculus, but the name has been changed to furcula because it is a noun in apposition and does not change with generic gender; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae; [MDD2025_2.0] generally spelt furculus, but the name has been changed to furcula because it is a noun in apposition and does not change with generic gender; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae; [batnames2025_1.7] Does not include auritus; see Peterson et al. (1995), but also see Hayman and Hill (1971). Originally spelled furcula but emended to furculus by Hill (1982), presumably to agree in gender with the generic epithet. However, HMW pointed out that this correction was uneccessary as furcula is a noun in apposition and therefore does not change endings to agree with the genus name. May include furinea Tate, 1941, possibly a lapsus for furcula (see discussion in Hill, 1982).; [MDD2025_2.2] generally spelt furculus, but the name has been changed to furcula because it is a noun in apposition and does not change with generic gender; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae				aurita		aurita.								furcula				furcula	furcula, furculus			furcula (Trouessart, 1906)|furculus (Corbet & J. Edwards Hill, 1980) [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Trouessart's trident bat	Madagascar, Aldabra, Cosmoledo Is	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Triaenops furculus	Madagascar, near Tulear, St. Augustine Bay.	Trouessart	1906	Bull. Mus. Hist. Nat. Paris, 1906, 7:446.	Distribution: Confined to Madagascar and the Aldabra group to the north west.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Trouessart's trident bat	Madagascar, Aldabra, Picard, Cosmoledo Is	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Trouessart	1906	Bull. Mus. Hist. Nat. Paris, 1906, 7:446.	Includes aurita; see Hayman and Hill (1971:30).	N and W Madagascar, Aldabra Isl.	Madagascar, near Tulear, St. Augustine Bay.		TROUESSART	1906	Noseleaf with a thickened supplementary ridge. Median anter ior process forked anteriorly. Outer margins of lateral projections of posterior noseleaf basally smoothly convex. Anterior margin of ear evenly rounded. Rostrum with lateral profile of nares nearly vertical. Size fairly small (forearm length, 43-46 mm).	Distribution: Confined to Madagascar and the Aldabra group to the north west.	No subspecies.		68	species	T. furculus	TROUESSART	1906	Triaenops	genus	Triaenops furculus				Noseleaf with a thickened supplementary ridge. Median anter ior process forked anteriorly. Outer margins of lateral projections of posterior noseleaf basally smoothly convex. Anterior margin of ear evenly rounded. Rostrum with lateral profile of nares nearly vertical. Size fairly small (forearm length, 43-46 mm).	No subspecies.		1. T. furculus TROUESSART 1906.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Triaenops furculus	Triaenops		furculus	Trouessart		1906		Bull. Mus. Natn. Hist. Nat. Paris	1906	7	446		Trouessart's Trident Bat	Madagascar, near Tulear (= Toliara), St. Augustine Bay, Grotte de Sarondrano.	N and W Madagascar, Aldabra and Cosmoledo Isls (Seychelles).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Does not includes auritus; see Peterson et al. (1995), but also see Hayman and Hill (1971). Originally spelled furcula but emended to furculus by Hill (1982), presumably to agree in gender with the generic epithet. May include furinea Tate, 1941, possibly a lapsus for furcula (see discussion in Hill, 1982).	0383245F2224977B8E0DF4DFF949FD3B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinonycteridae_194.pdf.imf	hash://md5/ffba5c272223977c8e73f509ffb1ffd0	209	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/83/24/0383245F2224977B8E0DF4DFF949FD3B.xml	Paratriaenops furcula	Rhinonycteridae	Paratriaenops	furcula		1906	Ti aenops deTrouessartl German @fr | Rinonictenode Trouessart @es	Tfiaenops fiımıla Trouessart, 1906 , “Grotte de Sarondrana (baie Saint-Augustin), cote Ouest de Madagascar [St. Augustine Bay, east coast of Madagascar ]. Widely used specific epithet ƒumdus has been replaced by the original epithet jinrula because it is a noun in apposition and does not need to be changed. Monotypic.	Endemic to Madagascar , where occurs in a narrow belt along most of W coast.	Head-body 50-60 mm, tail 19-27 mm, forearm 4249 mm , ear 15-20 mm; weight 3-9 g. No specific data are available for hindfoot length. Trouessarfs Trident Bat is small. Pelage has slightly darker tinge dorsally than ventrally, it is gray, grayish brown. dull brown, or occasionally pale yellowish gray. \Ming membranes are dark brown, and ears and noseleaf are pale gray or grayish brown. Noseleaf is large (width 6-5-8-1 mm), narrow, and rounded, with straplike, longitudinal, and anteriorly deeply bifurcated projection across anterior part of anterior leaf, above deep anterior medial emargination. Three tall pointed projections are present on posterior part of posterior leaf, and anterior part has in medial position short, laterally flattened, slightly upward directed projection. Total width of three posterior projections is almost entire width of posterior leaf. Posterior noseleaf is scalloped by numerous cells, small and more or les equal in size, arranged in two transversal rows, separated from each other by fleshy septa, except central pair of cells of posterior row that are separated by incomplete septum. Tall posterior projections are not emarginated at their bases. There are no lateral supplementary leaflets, but thickened ridge is beneath margin of anterior noseleaf. Ears are large and pointed, with anterior margin evenly rounded. Skull has laterally and dorsally very prominent nasal swellings. Braincase is slightly higher than rostrum, and there is depression between rostrum and braincase, with low but long sagittal crest. Wide plates are present on dorsal sides of zygomatic bones. Greatest skull lengths are 15-9-18 mm, condylo-canine lengths are 14-1-15-5 mm, zygomatic widths are 8-9 mm, and upper tooth row (C'-M‘) lengths are 5 -6-6-3 mm. I’ is bilobed; C' is slender, with slight cingulum; posterior secondary cusp is represented lather as a ridge extending one-third the crown height; P’ is small and extruded; C‘ and P‘ are in or nearly in contact; and M’ is only little reduced, with metacone. is bicuspid, 12 is tricuspid and larger than Il, C, is slender, P? has about twothirds crown area and one-half height of P‘, and M‘ is unreduced. Baculum is simple, very narrow, and 1 -5-1 -9 mm long, with pointed distal tip and broad and triangular proximal epiphysis; baculum widths are 0-2-0-5 mm (c.l 4—27% of baculum length).	Coastal lowland dry forests from sea level to elevations of c. l 40 m . Trouessart`s Trident Bats most frequently occupy dry deciduous forest limestone karst; however, they are found in numerous other dry vegetation fonnations on limestone karst (dry open palm savanna and spiny bush), dry vegetation on sand (original, transitional, disturbed, and degraded dry deciduous forests and original and disturbed spiny bush), and other local types of forests (e.g. gallery forests).	Considering its diet composition, Trouessart`s Trident Bat is most probably an aerial hawker, preying on flying insects of various sizes among vegetation. lndividuals were documented foraging in various forest habitats, most frequently in riparian forests and forest interiors and only occasionally at forest edges or forest clearings. Generally, dietary spectrum is narrow, with moths reportedly the most frequent prey item. Besides moths (on average 58-7%), beetles (21 -4%), bugs (7-5 %), cockroaches (4-2%), and flies (1 -7%) were also present in feces. Diets varied between seasons: dry season diets had c.64% moths, 12% beetles, 6% bugs, 1 % cockroaches, and 1 % flies, and wet season diets had c.22%, 52% beetles, 7% bugs, 13% cockroaches, and % flies. .	No information.	Trouessart's Trident Bats probably do not enter torpor or hibernate. They forage among vegetation at various heights aboveground. They leave day roosts around sunset and remain active for the entire night. Three hours after sunset, they begin to return to roosts and around midnight, flight activity increases and continues until almost dawn. Males return to roosts much earlier than females, many of which return just before dawn. Trouessart`s Trident Bat roosts in natural caves, common in its preferred karst regions. Besides caves and perhaps other underground spaces, hollows in trees and tree branches (e.g. in baobabs, Adansonía , Malvaceae ) probably serve as day roosts. Echolocation consists of multiharrnonic QCF calls where pulse is composed of long CF component, complemented at start and end by two short FM components (FM-CF-FM). lnitial FM component has very small upward sweep; terminating FM component has steep sweep of c. 18 kHz. Peak frequencies are lower in females than males. Calls last 8 -1 -22-5 milliseconds; interpulse intervals are 13-9-92-7 milliseconds; CF component maximum frequencies are 11-9-116 -8 kHz (mean 114 -8 kHz) in males and 98 -1 -102 -3 kHz (mean 99 -5 kHz) in females; CF component minimum frequencies are 106-8-113 kHz (mean 110 -5 kHz) in males and 91 -4-99 -4 kHz (mean 95 -7 kHz) in females; and end frequencies are 68-97 kHz (mean 81 -5 kHz). Remains of this species were found in small amounts of the diet of the bat hawk (Mruheiramphus alcinus).	Trouessarfs Trident Bats roost in rather large colonies; largest colony recorded, composed of both sexes, was estimated at c. lo,ooo individuals.	Classified as Least Concern on The IUCN Rai List. Trouessan`s Trident Bars could be threatened locally by disturbance at cave sites and from deforestation and forest fragmentation.	Goodman (2011) | Goodman à Ramvo (2008) | Goodman, Andrialidison et al (2005) | Goodman, Razakaratrimo à de Roland (2016) | Hill (1982a) | Kofoky er al (2009) | Olsson er al (2006) | Peterson et al (1995) | Rakotoarivelo et al (2007) | Rakotondramanana à Goodman (2017) | Ramasindrazana et a! (2013) | Fianivo à Goodman (2006) | Trouessart (1906)	https://zenodo.org/record/6611843/files/figure.png	8. Trouessart’s Trident Bat Paratriaenops furcula French: Ti aenops deTrouessartl German: Trouessan Dieizahnblartnase I Spanish: Rinonicteno deTrouessart Taxonomy. Tfiaenops fiımıla Trouessart, 1906 , “Grotte de Sarondrana (baie Saint-Augustin), cote Ouest de Madagascar [St. Augustine Bay, east coast of Madagascar ]. Widely used specific epithet ƒumdus has been replaced by the original epithet jinrula because it is a noun in apposition and does not need to be changed. Monotypic. Distribution. Endemic to Madagascar , where occurs in a narrow belt along most of W coast. Descriptive notes. Head-body 50-60 mm, tail 19-27 mm, forearm 4249 mm , ear 15-20 mm; weight 3-9 g. No specific data are available for hindfoot length. Trouessarfs Trident Bat is small. Pelage has slightly darker tinge dorsally than ventrally, it is gray, grayish brown. dull brown, or occasionally pale yellowish gray. \Ming membranes are dark brown, and ears and noseleaf are pale gray or grayish brown. Noseleaf is large (width 6-5-8-1 mm), narrow, and rounded, with straplike, longitudinal, and anteriorly deeply bifurcated projection across anterior part of anterior leaf, above deep anterior medial emargination. Three tall pointed projections are present on posterior part of posterior leaf, and anterior part has in medial position short, laterally flattened, slightly upward directed projection. Total width of three posterior projections is almost entire width of posterior leaf. Posterior noseleaf is scalloped by numerous cells, small and more or les equal in size, arranged in two transversal rows, separated from each other by fleshy septa, except central pair of cells of posterior row that are separated by incomplete septum. Tall posterior projections are not emarginated at their bases. There are no lateral supplementary leaflets, but thickened ridge is beneath margin of anterior noseleaf. Ears are large and pointed, with anterior margin evenly rounded. Skull has laterally and dorsally very prominent nasal swellings. Braincase is slightly higher than rostrum, and there is depression between rostrum and braincase, with low but long sagittal crest. Wide plates are present on dorsal sides of zygomatic bones. Greatest skull lengths are 15-9-18 mm, condylo-canine lengths are 14-1-15-5 mm, zygomatic widths are 8-9 mm, and upper tooth row (C'-M‘) lengths are 5 -6-6-3 mm. I’ is bilobed; C' is slender, with slight cingulum; posterior secondary cusp is represented lather as a ridge extending one-third the crown height; P’ is small and extruded; C‘ and P‘ are in or nearly in contact; and M’ is only little reduced, with metacone. is bicuspid, 12 is tricuspid and larger than Il, C, is slender, P? has about twothirds crown area and one-half height of P‘, and M‘ is unreduced. Baculum is simple, very narrow, and 1 -5-1 -9 mm long, with pointed distal tip and broad and triangular proximal epiphysis; baculum widths are 0-2-0-5 mm (c.l 4—27% of baculum length). Habitat. Coastal lowland dry forests from sea level to elevations of c. l 40 m . Trouessart`s Trident Bats most frequently occupy dry deciduous forest limestone karst; however, they are found in numerous other dry vegetation fonnations on limestone karst (dry open palm savanna and spiny bush), dry vegetation on sand (original, transitional, disturbed, and degraded dry deciduous forests and original and disturbed spiny bush), and other local types of forests (e.g. gallery forests). Food and Feeding. Considering its diet composition, Trouessart`s Trident Bat is most probably an aerial hawker, preying on flying insects of various sizes among vegetation. lndividuals were documented foraging in various forest habitats, most frequently in riparian forests and forest interiors and only occasionally at forest edges or forest clearings. Generally, dietary spectrum is narrow, with moths reportedly the most frequent prey item. Besides moths (on average 58-7%), beetles (21 -4%), bugs (7-5 %), cockroaches (4-2%), and flies (1 -7%) were also present in feces. Diets varied between seasons: dry season diets had c.64% moths, 12% beetles, 6% bugs, 1 % cockroaches, and 1 % flies, and wet season diets had c.22%, 52% beetles, 7% bugs, 13% cockroaches, and % flies. . s Breeding. No information. Activity patterns. Trouessart's Trident Bats probably do not enter torpor or hibernate. They forage among vegetation at various heights aboveground. They leave day roosts around sunset and remain active for the entire night. Three hours after sunset, they begin to return to roosts and around midnight, flight activity increases and continues until almost dawn. Males return to roosts much earlier than females, many of which return just before dawn. Trouessart`s Trident Bat roosts in natural caves, common in its preferred karst regions. Besides caves and perhaps other underground spaces, hollows in trees and tree branches (e.g. in baobabs, Adansonía , Malvaceae ) probably serve as day roosts. Echolocation consists of multiharrnonic QCF calls where pulse is composed of long CF component, complemented at start and end by two short FM components (FM-CF-FM). lnitial FM component has very small upward sweep; terminating FM component has steep sweep of c. 18 kHz. Peak frequencies are lower in females than males. Calls last 8 -1 -22-5 milliseconds; interpulse intervals are 13-9-92-7 milliseconds; CF component maximum frequencies are 11-9-116 -8 kHz (mean 114 -8 kHz) in males and 98 -1 -102 -3 kHz (mean 99 -5 kHz) in females; CF component minimum frequencies are 106-8-113 kHz (mean 110 -5 kHz) in males and 91 -4-99 -4 kHz (mean 95 -7 kHz) in females; and end frequencies are 68-97 kHz (mean 81 -5 kHz). Remains of this species were found in small amounts of the diet of the bat hawk (Mruheiramphus alcinus). Movements, Home range and Social organization. Trouessarfs Trident Bats roost in rather large colonies; largest colony recorded, composed of both sexes, was estimated at c. lo,ooo individuals. Status and Conservation. Classified as Least Concern on The IUCN Rai List. Trouessan`s Trident Bars could be threatened locally by disturbance at cave sites and from deforestation and forest fragmentation. Bibliography. Goodman (2011), Goodman à Ramvo (2008). Goodman, Andrialidison et al (2005), Goodman, Razakaratrimo à de Roland (2016). Hill (1982a). Kofoky er al (2009), Olsson er al (2006). Peterson et al (1995), Rakotoarivelo et al (2007). Rakotondramanana à Goodman (2017). Ramasindrazana et a! (2013), Fianivo à Goodman (2006). Trouessart (1906)	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinonycteridae	Paratriaenops furcula	Paratriaenops		furcula	Trouessart	1907	1	Bull. Mus. Natn. Hist. Nat. Paris	1906(7): 446	Trouessart's Trident Bat	None.	Madagascar, near Tulear (= Toliara), St. Augustine Bay, Grotte de Sarondrano.	N and W Madagascar, Aldabra and Cosmoledo Isls (Seychelles).	Not listed.	Least Concern as Paratriaenops furculus 	Does not include auritus ; see Peterson et al. (1995), but also see Hayman and Hill (1971). Originally spelled furcula but emended to furculus by Hill (1982), presumably to agree in gender with the generic epithet. However, HMW pointed out that this correction was uneccessary as furcula is a noun in apposition and therefore does not change endings to agree with the genus name. May include furinea Tate, 1941, possibly a lapsus for furcula (see discussion in Hill, 1982).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Paratriaenops furcula	23	Trouessart's Trident Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINONYCTERIDAE	NA	NA	Paratriaenops	NA	furcula	Trouessart	1906	1	Triaenops_furcula	Trouessart, M. E. L. (1906). Description de mammifères nouveaux d'Afrique er de Madagascar. Bulletin du Muséum d'histoire naturelle, 12, 446.	https://www.biodiversitylibrary.org/item/137042#page/456/mode/1up	MNHN 1912-40		"Grotte de Sarondrana (baie Saint-Augustin), côte Ouest de Madagascar [= St. Augustine Bay, east coast of Madagascar]."			furcula (Trouessart, 1907)	generally spelt furculus, but the name has been changed to furcula to match the generic gender; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333.	Madagascar	Africa	Afrotropic	LC	0	0	0	Paratriaenops_furcula	0	oldname match	Triaenops_furculus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Paratriaenops furculus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Paratriaenops	furculus	Trouessart, 1906		20000000	Paratriaenops furculus	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution. There is no evidence that this species is declining fast enough to place it in a category of higher threat although its roosting colonies should be monitored especially in areas undergoing habitat change.	In Madagascar, this species is restricted to dry deciduous and spiny forest in the south and west (Goodman et al. 2005) and appears to be an obligate cave dweller (Olsson et al . 2006). Undercut sea cliffs may provide suitable roosting habitats on small atolls (Hutson 2004). This species in Madagascar feeds predominantly on Lepidoptera (Rakotoarivelo et al . 2007) and appears to require relatively intact forest for foraging (Kofoky et al . 2007).	There may be possible threats from disturbance at the cave sties. This species is tolerant to some degree of forest degradation, but appears to require forested areas to survive (Goodman et al. 2005) and may therefore be one of the few Malagasy bat species really susceptible to deforestation and forest fragmentation.	There are few data available on P. furculus , but it appears to be a relatively rare member of the chiropteran assemblage in western Madagascar as determined by mist netting and acoustic sampling (Kofoky et al. 2007, Rakotoarivelo and Randrianandriananina 2007). However, it can occur in large colonies and a roost of over 10,000 individuals was reported from the Sept Lacs in southern Madagascar (Olsson et al. 2006). There is no information on population size of P. furculus from Aldabra but it appears to be rather rare with only a few specimen records (Hutson 2004).	Unknown	This species is found on Madagascar where it is restricted to lowland (an elevation span of 30 m to 200 m above sea level) areas in the west and south-west (Goodman et al. 2005; Ranivo and Goodman 2006). It is also found on Cosmoledo and Aldabra Atolls in the outer Seychelles (Hutson 2004).		Terrestrial	This species is present in Parc National du Tsingy de Bemaraha, Parc National de Namoroka, Parc National Kirindy-Mite and Parc National Tsimanampetsotsa (Goodman et al. 2005). Roosting colonies need to be the focus of further conservation and research.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinonycteridae	Paratriaenops		furcula	Trouessart	1907	1	Bull. Mus. Natn. Hist. Nat. Paris	1906(7): 446	Trouessart's Trident Bat	None.	Madagascar, near Tulear (= Toliara), St. Augustine Bay, Grotte de Sarondrano.	N and W Madagascar, Aldabra and Cosmoledo Isls (Seychelles).	Not listed.	Least Concern as Paratriaenops furculus 	Does not include auritus ; see Peterson et al. (1995), but also see Hayman and Hill (1971). Originally spelled furcula but emended to furculus by Hill (1982), presumably to agree in gender with the generic epithet. However, HMW pointed out that this correction was uneccessary as furcula is a noun in apposition and therefore does not change endings to agree with the genus name. May include furinea Tate, 1941, possibly a lapsus for furcula (see discussion in Hill, 1982).	Paratriaenops furcula	1004763	23	Trouessart's Trident Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinonycteridae	NA	NA	Paratriaenops	NA	furcula	Trouessart	1906	1	Triaenops_furcula	Trouessart, M. E. L. (1906). Description de mammifères nouveaux d'Afrique er de Madagascar. Bulletin du Muséum d'histoire naturelle, 12, 446.	https://www.biodiversitylibrary.org/item/137042#page/456/mode/1up	MNHN 1912-40		"Grotte de Sarondrana (baie Saint-Augustin), côte Ouest de Madagascar [= St. Augustine Bay, east coast of Madagascar]."			furcula (Trouessart, 1907)	generally spelt furculus, but the name has been changed to furcula because it is a noun in apposition and does not change with generic gender; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333.				Madagascar	Africa	Afrotropic	LC	0	0	0	Paratriaenops_furcula	0	oldname match	Triaenops_furculus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Paratriaenops_furcula	1004763	23	Trouessart's Trident Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinonycteridae	NA	NA	Paratriaenops	NA	furcula	Trouessart	1	Triaenops furcula	Trouessart, É.L. 1906. Description de Mammifères Nouveaux d'Afrique et de Madagascar. Bulletin du Muséum d'histoire naturelle 12:443-446.	https://www.biodiversitylibrary.org/page/42892047	MNHN-ZM-MO-1912-40	holotype	http://coldb.mnhn.fr/catalognumber/mnhn/zm/mo-1912-40	"Grotte de Sarondrana (baie Saint-Augustin), côte Ouest de Madagascar [= St. Augustine Bay, east coast of Madagascar]."			generally spelt furculus, but the name has been changed to furcula because it is a noun in apposition and does not change with generic gender; moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333.				Madagascar	Africa	Afrotropic	LC	0	0	0	Paratriaenops_furcula	0	oldname match	Triaenops_furculus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinonycteridae	Paratriaenops		furcula	Trouessart	1907	1	Bull. Mus. Natn. Hist. Nat. Paris	1906(7): 446	Trouessart's Trident Bat	None.	Madagascar, near Tulear (= Toliara), St. Augustine Bay, Grotte de Sarondrano.	N and W Madagascar, Aldabra and Cosmoledo Isls (Seychelles).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/81060220/22040490/' target='_blank'>Least Concern</a>	Does not include auritus; see Peterson et al. (1995), but also see Hayman and Hill (1971). Originally spelled furcula but emended to furculus by Hill (1982), presumably to agree in gender with the generic epithet. However, HMW pointed out that this correction was uneccessary as furcula is a noun in apposition and therefore does not change endings to agree with the genus name. May include furinea Tate, 1941, possibly a lapsus for furcula (see discussion in Hill, 1982).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Triaenops furculus; Paratriaenops furcula; Paratriaenops furcula; Paratriaenops furcula; Paratriaenops furculus; Paratriaenops furcula; furcula; Ti aenops deTrouessartl German; Rinonictenode Trouessart; Trouessart's Trident Bat; Trouessart's Trident Bat; Trouessart's Trident Bat; P. furcula