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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1125	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Triaenops auritus	Paratriaenops auritus	Paratriaenops auritus	Paratriaenops auritus	Paratriaenops auritus	Paratriaenops auritus	Paratriaenops auritus	Paratriaenops auritus	Paratriaenops auritus	Paratriaenops auritus		[MSW3] Known only from the holotype. Often included in furculus (e.g., Hayman and Hill, 1971; Koopman, 1993, 1994), but see Peterson et al. (1995). Originally spelled aurita but emended to auritus by Peterson et al. (1995), presumably to agree in gender with the generic epithet.; [HMW] Triaenops aurita [sic] G. Grandidier, 1912 , “environs de Diegosuarez, dans Fextrême Nord de Madagascar [Diana Region, Antsiranana , northern Madagascar ]. Paratriamops aurituswas formerly included in R. fuırula; however, it was demonsuated to represent a distinct species based on morphological and genetic comparisons. Monotvpic.; [batnames2022] Known only from the holotype. Often included in furculus (e.g., Hayman and Hill, 1971; Koopman, 1993, 1994), but see Peterson et al. (1995). Originally spelled aurita but emended to auritus by Peterson et al. (1995), presumably to agree in gender with the generic epithet.; [MDD2022] moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae; [IUCN] Paratriaenops auritus was recognized as a full species by Peterson et al. (1995), which has been supported by more recent morphological and genetic analyses (Ranivo and Goodman 2006, Russell et al . 2007).; [batnames2023] Known only from the holotype. Often included in furculus (e.g., Hayman and Hill, 1971; Koopman, 1993, 1994), but see Peterson et al. (1995). Originally spelled aurita but emended to auritus by Peterson et al. (1995), presumably to agree in gender with the generic epithet.; [MDD2023] moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae; [MDD2025_2.0] moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae; [batnames2025_1.7] Known only from the holotype. Often included in furculus (e.g., Hayman and Hill, 1971; Koopman, 1993, 1994), but see Peterson et al. (1995). Originally spelled aurita but emended to auritus by Peterson et al. (1995), presumably to agree in gender with the generic epithet.; [MDD2025_2.2] moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae														auritus	Paratriaenops auritus was recognized as a full species by Peterson et al. (1995), which has been supported by more recent morphological and genetic analyses (Ranivo and Goodman 2006, Russell et al . 2007).			auritus	auritus			auritus (G. Grandidier, 1912)						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Triaenops auritus	Triaenops		auritus	Grandidier		1912		Bull. Mus. Natl. Hist. Nat. Paris	18		8		Grandidier's Trident Bat	Madagascar, near Diégo-Suarez (=Antsiranana).	Known only from the type locality.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Data Deficient.		Known only from the holotype. Often included in furculus (e.g., Hayman and Hill, 1971; Koopman, 1993, 1994), but see Peterson et al. (1995). Originally spelled aurita but emended to auritus by Peterson et al. (1995), presumably to agree in gender with the generic epithet.	0383245F2225977B8BD5F631FD78FE56	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinonycteridae_194.pdf.imf	hash://md5/ffba5c272223977c8e73f509ffb1ffd0	208	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/83/24/0383245F2225977B8BD5F631FD78FE56.xml	Paratriaenops auritus	Rhinonycteridae	Paratriaenops	auritus		1912	Tr aen ps de Grand d er German @fr | Golden Trident Bat @en	Triaenops aurita [sic] G. Grandidier, 1912 , “environs de Diegosuarez, dans Fextrême Nord de Madagascar [Diana Region, Antsiranana , northern Madagascar ]. Paratriamops aurituswas formerly included in R. fuırula; however, it was demonsuated to represent a distinct species based on morphological and genetic comparisons. Monotvpic.	Endemic to Madagascar , where it occurs at elevations of 50-200 m in the most exueme N; S distributional limits lie at c. 14 S between Ambilobe and Antsohihy.	Head-body 47-64 mm, tail 19-28 mm, ear 15-20 mm, forearm 43-51 mm; weight 5-8 g. No specific data are available for hindfoot length. Grandidier`s Trident Bat small, but on average, the largest species of Paratriamops. Pelage has slightly darker tinge dorsally than ventrally, being reddish golden, orange, or bright brown. Wing membranes are dark brown, and ears and noseleaf are pale grayish. Noseleaf is large, narrow, and rounded, with strap-like, longitudinal, and anteriorly deeply bifurcated projection across anterior part of anterior leaf, above deep anterior medial emargination. Three tall pointed projections are present on posterior part of posterior noseleaf, its anterior part has in medial position short, laterally flattened, and slightly upward directed projection. Total width of three posterior projections is almost entire width of posterior leaf. Posterior leaf is scalloped by numerous cells, small and more or less equal in size, ananged in two transversal rows, separated from each other by fleshy septa, except central pair of cells of posterior row, which are separated by an incomplete septum. Tall posterior projections are not emarginated at their bases. There are no lateral supplementary leaflets, but thickened ridge beneath margin of anterior noseleaf. Ears are large and pointed, with anterior margin evenly rounded. Skull has very prominent nasal swellings laterallv and dorsally. Braincase s slightly higher than rostnrm, and there depression between rostrum and braincase, with low but long sagittal crest. Wide plates are present on dorsal sides of zygomatic bones. Greatest skull lengths are 16-8-18-8 mm, condylo-canine lengths are 14-8-16-2 mm, zygomatic widths are 8 -4-9-3 mm, and upper tooth row (C'-M") lengths are 5 -8-6-5 mm. is bilobed; C' slender, with slight cingulum; posterior secondary cusp represented rather as ridge extending onethird the crown height; P’ is small and extruded; C' and P‘ are in or nearlv in contact; and M‘ is only little reduced, with metacone. is bicuspid, I, is tricuspid and larger than C, is slender, P? has about two-thirds the crown area and one-half the height of P‘, and M‘ is unreduced. Baculum is simple, narrow, and c. l-8 mm long, with pointed distal tip, and broad and rounded proximal epiphysis; baculum width is c.0-5 mm (i. e. c.28 % of baculum length).	Lowland dry deciduous forests combined with limestone karst.	No infonnation.	No information.	Grandidiefs Trident Bat roosts at great depths in underground spaces; prefers narrow corridors c. 200 m from roost entrances. most probably does not enter torpor or hibernate. Echolocation consists of multiharrnonic QCF calls, with pulse composed of long CF component, complemented at start and end by two short FM components (FM-CF-FM). lnitial FM component has small upward sweep (not always detectable); terminating FM component is steep sweep of 5-7 kHz. Peak frequencies of females are lower than males. Calls last 4 -6-22 -3 milliseconds; interpulse intervals are 17 -8-101 milliseconds; CF component maximum frequencies are 106-5-110-2 kHz (mean 108 -8 kHz) in males and 95 -6-97-2 kHz (mean 96 -4 kHz) in females; CF component minimum frequencies are 104-8-108 -2 kHz (mean 106 -5 kHz) in males and 93 -8-94-1 kHz (mean 93 -9 kHz) in females; and end frequencies are 80-1-107-6 kHz (mean 82 -9 kHz).	Grandidier's Trident Bats roost in large colonies; largest numbers recorded were c. 2000 individuals in a natural cave and c. 1000 individuals in a mine.	Classified as Vulnerable on The IUCN Red List. Grandidier`s Trident Bat has small, fragmented, and declining distribution. 1t is regarded as threatened particularly from habitat destruction and fragmentation and disturbance at cave roosts. 1 t occurs in Ankarana, Analarnerana, and Daraina reserves in northern Madagascar , which cover a significant part of its distribution.	Goodman (2011) | Goodman à Ranıvo (2008) | Goodman, Andrıafıdıson er al (2005) | Grandıdıer (1912) | H (1982a) | Kofoky er al (2009) | Peterson et al (1995) | Rakotondremanana & Goodman (2017) | Ramasındrazana er al (2013) | Ranıvo à Goodman (2006)	https://zenodo.org/record/6611841/files/figure.png	7. Grandidiefs Trident Bat Paratriaenops auritus French: Tr aen ps de Grand d er German: Grand drer-Dre zahnblannase Spanish: R non terio de Grındıdıer Other common names: Golden Trident Bat Taxonomy. Triaenops aurita [sic] G. Grandidier, 1912 , “environs de Diegosuarez, dans Fextrême Nord de Madagascar [Diana Region, Antsiranana , northern Madagascar ]. Paratriamops aurituswas formerly included in R. fuırula; however, it was demonsuated to represent a distinct species based on morphological and genetic comparisons. Monotvpic. Distribution . Endemic to Madagascar , where it occurs at elevations of 50-200 m in the most exueme N; S distributional limits lie at c. 14 S between Ambilobe and Antsohihy. Descriptive notes. Head-body 47-64 mm, tail 19-28 mm, ear 15-20 mm, forearm 43-51 mm; weight 5-8 g. No specific data are available for hindfoot length. Grandidier`s Trident Bat small, but on average, the largest species of Paratriamops. Pelage has slightly darker tinge dorsally than ventrally, being reddish golden, orange, or bright brown. Wing membranes are dark brown, and ears and noseleaf are pale grayish. Noseleaf is large, narrow, and rounded, with strap-like, longitudinal, and anteriorly deeply bifurcated projection across anterior part of anterior leaf, above deep anterior medial emargination. Three tall pointed projections are present on posterior part of posterior noseleaf, its anterior part has in medial position short, laterally flattened, and slightly upward directed projection. Total width of three posterior projections is almost entire width of posterior leaf. Posterior leaf is scalloped by numerous cells, small and more or less equal in size, ananged in two transversal rows, separated from each other by fleshy septa, except central pair of cells of posterior row, which are separated by an incomplete septum. Tall posterior projections are not emarginated at their bases. There are no lateral supplementary leaflets, but thickened ridge beneath margin of anterior noseleaf. Ears are large and pointed, with anterior margin evenly rounded. Skull has very prominent nasal swellings laterallv and dorsally. Braincase s slightly higher than rostnrm, and there depression between rostrum and braincase, with low but long sagittal crest. Wide plates are present on dorsal sides of zygomatic bones. Greatest skull lengths are 16-8-18-8 mm, condylo-canine lengths are 14-8-16-2 mm, zygomatic widths are 8 -4-9-3 mm, and upper tooth row (C'-M") lengths are 5 -8-6-5 mm. is bilobed; C' slender, with slight cingulum; posterior secondary cusp represented rather as ridge extending onethird the crown height; P’ is small and extruded; C' and P‘ are in or nearlv in contact; and M‘ is only little reduced, with metacone. is bicuspid, I, is tricuspid and larger than C, is slender, P? has about two-thirds the crown area and one-half the height of P‘, and M‘ is unreduced. Baculum is simple, narrow, and c. l-8 mm long, with pointed distal tip, and broad and rounded proximal epiphysis; baculum width is c.0-5 mm (i. e. c.28 % of baculum length). Habitat. Lowland dry deciduous forests combined with limestone karst. Food and Feeding. No infonnation. Breeding. No information. Activity patterns. Grandidiefs Trident Bat roosts at great depths in underground spaces; prefers narrow corridors c. 200 m from roost entrances. most probably does not enter torpor or hibernate. Echolocation consists of multiharrnonic QCF calls, with pulse composed of long CF component, complemented at start and end by two short FM components (FM-CF-FM). lnitial FM component has small upward sweep (not always detectable); terminating FM component is steep sweep of 5-7 kHz. Peak frequencies of females are lower than males. Calls last 4 -6-22 -3 milliseconds; interpulse intervals are 17 -8-101 milliseconds; CF component maximum frequencies are 106-5-110-2 kHz (mean 108 -8 kHz) in males and 95 -6-97-2 kHz (mean 96 -4 kHz) in females; CF component minimum frequencies are 104-8-108 -2 kHz (mean 106 -5 kHz) in males and 93 -8-94-1 kHz (mean 93 -9 kHz) in females; and end frequencies are 80-1-107-6 kHz (mean 82 -9 kHz). Movements, Home range and Social organization. Grandidier's Trident Bats roost in large colonies; largest numbers recorded were c. 2000 individuals in a natural cave and c. 1000 individuals in a mine. Status and Conservation. Classified as Vulnerable on The IUCN Red List. Grandidier`s Trident Bat has small, fragmented, and declining distribution. 1t is regarded as threatened particularly from habitat destruction and fragmentation and disturbance at cave roosts. 1 t occurs in Ankarana, Analarnerana, and Daraina reserves in northern Madagascar , which cover a significant part of its distribution. Bibliography . Goodman (2011), Goodman à Ranıvo (2008). Goodman, Andrıafıdıson er al (2005). Grandıdıer (1912) H (1982a). Kofoky er al (2009), Peterson et al (1995), Rakotondremanana & Goodman (2017). Ramasındrazana er al (2013). Ranıvo à Goodman (2006)	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinonycteridae	Paratriaenops auritus	Paratriaenops		auritus	Grandidier	1912	1	Bull. Mus. Natl. Hist. Nat. Paris	18:08	Grandidier's Trident Bat	None.	Madagascar, near Diégo-Suarez (=Antsiranana).	Known only from the type locality.	Not listed.	Vulnerable	Known only from the holotype. Often included in furculus (e.g., Hayman and Hill, 1971; Koopman, 1993, 1994), but see Peterson et al. (1995). Originally spelled aurita but emended to auritus by Peterson et al. (1995), presumably to agree in gender with the generic epithet.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Paratriaenops auritus	23	Grandidier's Trident Bat	Golden Trident Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINONYCTERIDAE	NA	NA	Paratriaenops	NA	auritus	G. Grandidier	1912	1	Triaenops_aurita	Grandidier, G. (1912). Une nouvelle Chauve-Souris de Madagascar, le Triaenops aurita G. G. Bulletin du Muséeum national d'histoire naturelle, 18, 8.	https://www.biodiversitylibrary.org/item/106493#page/22/mode/1up	MCZ 45080		"environs de Diego-Suarez, dans l'extrême Nord de Madagascar [= Diana Region, Antsiranana, northern Madagascar]."			auritus (G. Grandidier, 1912)	moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333.	Madagascar	Africa	Afrotropic	VU	0	0	0	Paratriaenops_auritus	0	oldname match	Triaenops_auritus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	40025	Paratriaenops auritus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Paratriaenops	auritus	(Grandidier, 1912)	Paratriaenops auritus was recognized as a full species by Peterson et al. (1995), which has been supported by more recent morphological and genetic analyses (Ranivo and Goodman 2006, Russell et al . 2007).	20000000	Paratriaenops auritus	Vulnerable	B1ab(iii)	2017	2016-04-24 00:00:00 UTC	3.1	English	This species is listed as Vulnerable as it has what appears to be a restricted geographic range of less than 20,000 km² in northwestern Madagascar. The habitat is severely fragmented, and there is a continuing decline in the area and quality of habitat mainly due to agricultural activities. Further survey work is needed to help better ascertain the limits of the range of this species.	This is an obligate cave-dwelling species that can be locally abundant, but has also been noted as rare within certain parts of its range (Robinson et al. 2006). Its patchy local distribution and density may be related to the presence of suitable caves for roosting (Goodman et al . 2005) and this species seems to prefer relatively narrow caves (Cardiff 2006). This species is thought to be dependent on relatively intact forest for foraging (Goodman et al . 2005).	As an apparent forest dependent species any decline in extent or quality of the remaining forest poses a threat to this species. In northern Madagascar, forest is lost because of expanding agriculture. There is also the threat of disturbance to cave roosting sites.	This species is thought to be relatively abundant within a localized area in the northern part of Madagascar. The largest roost recorded was 2,000 individuals in Parc National d’Ankarana (S.G. Cardiff pers. obs.) and up to 1,000 individuals were recorded from a mine tunnel approximately 5 km from Andavokoera (J. Ranivo pers. comm).	Unknown	This species is endemic to the island of Madagascar where it has a restricted range in the north-western tip of the island, ranging between near sea-level and 600 m above sea-level and the southern limit is the Andrafiamena mountains (Goodman and Ramasindrazana 2013).		Terrestrial	This species is known to occur in three protected areas: Parc National d'Ankarana, Réserve Spéciale d’Analamerana and Parc National de Loky-Manambato (Ranivo and Goodman 2006, Russell et al. 2007). Conservation should be focused at the known cave roosts to secure the existing populations and closely monitor seasonal and annual population changes.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinonycteridae	Paratriaenops		auritus	Grandidier	1912	1	Bull. Mus. Natl. Hist. Nat. Paris	18:08	Grandidier's Trident Bat	None.	Madagascar, near Diégo-Suarez (=Antsiranana).	Known only from the type locality.	Not listed.	Vulnerable	Known only from the holotype. Often included in furculus (e.g., Hayman and Hill, 1971; Koopman, 1993, 1994), but see Peterson et al. (1995). Originally spelled aurita but emended to auritus by Peterson et al. (1995), presumably to agree in gender with the generic epithet.	Paratriaenops auritus	1004762	23	Grandidier's Trident Bat	Golden Trident Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinonycteridae	NA	NA	Paratriaenops	NA	auritus	G. Grandidier	1912	1	Triaenops_aurita	Grandidier, G. (1912). Une nouvelle Chauve-Souris de Madagascar, le Triaenops aurita G. G. Bulletin du Muséeum national d'histoire naturelle, 18, 8.	https://www.biodiversitylibrary.org/item/106493#page/22/mode/1up	MCZ 45080		"environs de Diego-Suarez, dans l'extrême Nord de Madagascar [= Diana Region, Antsiranana, northern Madagascar]."			auritus (G. Grandidier, 1912)	moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333.				Madagascar	Africa	Afrotropic	VU	0	0	0	Paratriaenops_auritus	0	oldname match	Triaenops_auritus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Paratriaenops_auritus	1004762	23	Grandidier's Trident Bat	Golden Trident Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinonycteridae	NA	NA	Paratriaenops	NA	auritus	G. Grandidier	1	Triaenops aurita	Grandidier, G. 1912. Un nouvelle Chauve-Souris de Madagascar, le _Triaenops aurita_ G.G. Bulletin du Muséum national d'histoire naturelle (1)18:8-9.	https://www.biodiversitylibrary.org/page/34142095	MCZ:Mamm:45080	holotype	https://mczbase.mcz.harvard.edu/guid/MCZ:Mamm:45080	"environs de Diego-Suarez, dans l'extrême Nord de Madagascar [= Diana Region, Antsiranana, northern Madagascar]."			moved from Triaenops to Paratriaenops; moved from Hipposideridae to Rhinonycteridae	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333.				Madagascar	Africa	Afrotropic	VU	0	0	0	Paratriaenops_auritus	0	oldname match	Triaenops_auritus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinonycteridae	Paratriaenops		auritus	Grandidier	1912	1	Bull. Mus. Natl. Hist. Nat. Paris	18:08	Grandidier's Trident Bat	None.	Madagascar, near Diégo-Suarez (=Antsiranana).	Known only from the type locality.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/40025/22064746/' target='_blank'>Vulnerable</a>	Known only from the holotype. Often included in furculus (e.g., Hayman and Hill, 1971; Koopman, 1993, 1994), but see Peterson et al. (1995). Originally spelled aurita but emended to auritus by Peterson et al. (1995), presumably to agree in gender with the generic epithet.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Triaenops auritus; Paratriaenops auritus; Paratriaenops auritus; Paratriaenops auritus; Paratriaenops auritus; Paratriaenops auritus; auritus; Tr aen ps de Grand d er German; Golden Trident Bat; Grandidier's Trident Bat; Golden Trident Bat; Grandidier's Trident Bat; Grandidier's Trident Bat; P. auritus