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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L113	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea	Carollia castanea		[MSW3] This complex probably includes more than one species; see Hoffman and Baker (2003).; [HMW] Carollia castanea H. Allen, 1890 , “ Costa Rica .” Restricted by R. H. Pine in 1972 to Angostura, San José, Costa Rica . S. Solari and R.J. Baker in 2006 identified C. castanea as a species complex based on molecular and morphological analyses. This account deals with the species as treated by L. J. McLellan and K. F. Koopman in 2008 that included C. castanea (sensu stricto) plus an unnamed species. Monotypic.; [batnames2022] Does not include colombiana , which is a synonym of brevicauda ; see Zurc and Velazco (2010). Identified as a species complex by Hoffman and Baker (2003), with specimens from much of the southern part of the range described as a new species, benkeithi , by Solari and Baker (2006). Two clades of castanea have been consistently recovered (e.g., castanea a and castanea b of Velazco, 2013) and the species as we recognize it here should be split. However, the distributional limits of the two clades within castanea overlap and it is not clear which clade should bear the name castanea ; see Velazco (2013). The type locality ("Costa Rica") was restricted by Pine (1972). See also McLellan and Koopman (2008), Jarrin-V et al. (2010), and López-Aguirre et al. (2015).; [IUCN] The species previously included populations with distinctive morphologies and karyotypes, that required description of a new species, C. benkeithi , to better recognize that evolutionary diversity (Solari and Baker 2006).; [batnames2023] Does not include colombiana , which is a synonym of brevicauda ; see Zurc and Velazco (2010). Identified as a species complex by Hoffman and Baker (2003), with specimens from much of the southern part of the range described as a new species, benkeithi , by Solari and Baker (2006). Two clades of castanea have been consistently recovered (e.g., castanea a and castanea b of Velazco, 2013) and the species as we recognize it here should be split. However, the distributional limits of the two clades within castanea overlap and it is not clear which clade should bear the name castanea ; see Velazco (2013). The type locality ("Costa Rica") was restricted by Pine (1972). See also McLellan and Koopman (2008), Jarrin-V et al. (2010), and López-Aguirre et al. (2015).; [batnames2025_1.7] Does not include colombiana, which is a synonym of brevicauda; see Zurc and Velazco (2010). Identified as a species complex by Hoffman and Baker (2003), with specimens from much of the southern part of the range described as a new species, benkeithi, by Solari and Baker (2006). Two clades of castanea have been consistently recovered (e.g., castanea a and castanea b of Velazco, 2013) and the species as we recognize it here should be split. However, the distributional limits of the two clades within castanea overlap and it is not clear which clade should bear the name castanea; see Velazco (2013). The type locality ("Costa Rica") was restricted by Pine (1972). See also McLellan and Koopman (2008), Jarrin-V et al. (2010), and López-Aguirre et al. (2015).														castanea	The species previously included populations with distinctive morphologies and karyotypes, that required description of a new species, C. benkeithi , to better recognize that evolutionary diversity (Solari and Baker 2006).			castanea	castanea			castanea H. Allen, 1890|castanet Uieda, 1980 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Allen's short-tailed bat	Honduras – E Peru, Bolivia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Carollia castanea	Costa Rica, Angostura.	H. Allen	1890	Proc. Am. Philos. Soc., 28:19.	Distribution: Ranging from Honduras through Central and tropical South America to French Guiana and Bolivia but west of the Andes not south of Ecuador.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Allen's short-tailed bat	Honduras – E Peru, Bolivia, Venezuela, Guianas	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	H. Allen	1890	Proc. Am. Philos. Soc., 28:19.		Honduras to Peru, Bolivia, W Brazil and Venezuela.	Costa Rica, Angostura.		H. ALLEN	1890	Size relatively small (forearm length, 33-39 mm; condylobasal length, 16-19 mm). Labial margin of maxillary toothrow with a distinct notch. Crown of anterior lower molar extremely low. Outer lower incisors somewhat reduced but not concealed dorsally by cingula of canines. Mandible somewhat Ushaped.	Distribution: Ranging from Honduras through Central and tropical South America to French Guiana and Bolivia but west of the Andes not south of Ecuador.	No subspecies.		84	species	C. castanea	H. ALLEN	1890	Carollia	genus	Carollia castanea				Size relatively small (forearm length, 33-39 mm; condylobasal length, 16-19 mm). Labial margin of maxillary toothrow with a distinct notch. Crown of anterior lower molar extremely low. Outer lower incisors somewhat reduced but not concealed dorsally by cingula of canines. Mandible somewhat U- shaped.	No subspecies.		4. C. castanea H. ALLEN 1890.	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Carolliinae		Carollia castanea	Carollia		castanea	H. Allen		1890		Proc. Am. Philos. Soc.	28		19		Chestnut Short-tailed Bat	Costa Rica, Angostura.	Honduras to Peru, Bolivia, W Brazil and Venezuela.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		This complex probably includes more than one species; see Hoffman and Baker (2003).	03A687BCFF82FF8216B8FE74F83AF91B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	535	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF82FF8216B8FE74F83AF91B.xml	Carollia castanea	Phyllostomidae	Carollia	castanea	H. Allen	1890	Carollia d Allen @fr | Kastanienbraune Kurzschwanzblattnase @de | Carolia de Allen @es	Carollia castanea H. Allen, 1890 , “ Costa Rica .” Restricted by R. H. Pine in 1972 to Angostura, San José, Costa Rica . S. Solari and R.J. Baker in 2006 identified C. castanea as a species complex based on molecular and morphological analyses. This account deals with the species as treated by L. J. McLellan and K. F. Koopman in 2008 that included C. castanea (sensu stricto) plus an unnamed species. Monotypic.	Central America from W Honduras to NW South America, including Colombia , Venezuela , Guyana , Ecuador , and N Peru , N of the Amazon River. Because of abundance of individuals of the C. castanea complex, hampering precise species identification in the field and in collections, a detailed assessment of its distribution is still needed.	Head-body 48-60 mm, tail 7-14 mm, ear 15-19 mm, hindfoot 9-14 mm, forearm 34-38 mm; weight 11-16 g. The Chestnut Short-tailed Bat is small. Dorsal color varies from dull, dark gray-brown or chestnut to pale tan, rarely grayish; tricolored banding of hairs is not well defined; bases are dull brown; and fur is ¢. 6 mm long. Ventral hairs are short and bicolored, with brown tips. Forearms are short and naked, with tuft of hair at bases of thumbs. Uropatagium is wide, with shallow distal notch. Tail is short, almost one-third the length of uropatagium. Wings are attached to ankles. Muzzle is conspicuous but short; lower lip has central papillae surrounded by smaller warts in a Vor U-shape. Noseleaf is short with elongated tip, and horseshoe is fused below nostrils and free on sides. Ears are moderately large and triangular, with pointed tips. Rostrum is short, braincase is globular, and zygomatic arches are incomplete. Anterior upper tooth row (to P?) is almost parallel, but posterior one-half is more divergent. Lower premolars are graded, with P, being smaller rather than subequal; there is a gap between them. Upper and lower molars are broad. Cusps of M, are reduced and not visible in lateral view. Chromosomal complement has 2n = 20 (females) and 21 (males) and FN = 38, with an autosometranslocated to the subtelocentric X-chromosome.	Common in secondary woodland, clearings, and plantations and less common in mature forests, clearings, and fruit groves, mostly below elevations of 1000 m .	The Chestnut Short-tailed Bat is most specialized on fruits of Piper ( Piperaceae ), mostly in dry seasons; it eats fruits form various trees like Markea panamensis ( Solanaceae ) and Dipteryx panamensis ( Fabaceae ) in wet seasons. Insects are occasionally eaten. The Chestnut Short-tailed Bat forages near moist areas and is caught most frequently in tropical evergreen forests. It uses understory vegetation, where it concentratesits feeding on fruits of shrubs and small trees, especially slender green, candle-like fruits of Piper plants. Itis one of the most important seed dispersers for Piper and many other plants with small fruits.	Central American populations of Chestnut Short-tailed Bats in Panama have at least two birth peaks in April-May and August-November, confirming a polyestrous breeding pattern. Nevertheless, these records were taken from widely separate localities, and a fixed breeding pattern might characterize populations in a given area.	The Chestnut Short-tailed Bat is most active 1-3 hours shortly after sunset. Peak activity seems to occur for 3-5 hours of the night. Field data also confirm that it 1s more active during the first one-half of the night, with occasional captures after midnight. Scarcity of records might indicate difficulty of properly identifying it in roosts, or an opportunistic use of roosts where available. Chestnut Short-tailed Bats roost in caves, tunnels, hollow trees, and mines and under overhanging roots.	The Chestnut Short-tailed Bat has small home ranges. It was found sharing a shallow limestone cave with Seba’s Shorttailed Bats ( C. perspicillata ).	Classified as Least Concern on The IUCN Red List. The Chest nut Short-tailed Bat has a wide distribution and presumably large and stable population.	Allen (1890c) | Bonaccorso et al. (2007) | Fleming (1991) | Fleming et al. (1972) | Gorchov etal. (1995) | Hoffmann & Baker (2003) | McLellan & Koopman (2008) | Pine (1972) | Reid (2009) | Solari & Baker (2006) | Thies & Kalko (2004) | Thies etal. (2006) | Wilson (1979)	https://zenodo.org/record/6458819/files/figure.png	104. Chestnut Short-tailed Bat Carollia castanea French: Carollia dAllen / German: Kastanienbraune Kurzschwanzblattnase / Spanish: Carolia de Allen Taxonomy. Carollia castanea H. Allen, 1890 , “ Costa Rica .” Restricted by R. H. Pine in 1972 to Angostura, San José, Costa Rica . S. Solari and R.J. Baker in 2006 identified C. castanea as a species complex based on molecular and morphological analyses. This account deals with the species as treated by L. J. McLellan and K. F. Koopman in 2008 that included C. castanea (sensu stricto) plus an unnamed species. Monotypic. Distribution. Central America from W Honduras to NW South America, including Colombia , Venezuela , Guyana , Ecuador , and N Peru , N of the Amazon River. Because of abundance of individuals of the C. castanea complex, hampering precise species identification in the field and in collections, a detailed assessment of its distribution is still needed. Descriptive notes. Head-body 48-60 mm, tail 7-14 mm, ear 15-19 mm, hindfoot 9-14 mm, forearm 34-38 mm; weight 11-16 g. The Chestnut Short-tailed Bat is small. Dorsal color varies from dull, dark gray-brown or chestnut to pale tan, rarely grayish; tricolored banding of hairs is not well defined; bases are dull brown; and fur is ¢. 6 mm long. Ventral hairs are short and bicolored, with brown tips. Forearms are short and naked, with tuft of hair at bases of thumbs. Uropatagium is wide, with shallow distal notch. Tail is short, almost one-third the length of uropatagium. Wings are attached to ankles. Muzzle is conspicuous but short; lower lip has central papillae surrounded by smaller warts in a Vor U-shape. Noseleaf is short with elongated tip, and horseshoe is fused below nostrils and free on sides. Ears are moderately large and triangular, with pointed tips. Rostrum is short, braincase is globular, and zygomatic arches are incomplete. Anterior upper tooth row (to P?) is almost parallel, but posterior one-half is more divergent. Lower premolars are graded, with P, being smaller rather than subequal; there is a gap between them. Upper and lower molars are broad. Cusps of M, are reduced and not visible in lateral view. Chromosomal complement has 2n = 20 (females) and 21 (males) and FN = 38, with an autosometranslocated to the subtelocentric X-chromosome. Habitat. Common in secondary woodland, clearings, and plantations and less common in mature forests, clearings, and fruit groves, mostly below elevations of 1000 m . Food and Feeding. The Chestnut Short-tailed Bat is most specialized on fruits of Piper ( Piperaceae ), mostly in dry seasons; it eats fruits form various trees like Markea panamensis ( Solanaceae ) and Dipteryx panamensis ( Fabaceae ) in wet seasons. Insects are occasionally eaten. The Chestnut Short-tailed Bat forages near moist areas and is caught most frequently in tropical evergreen forests. It uses understory vegetation, where it concentratesits feeding on fruits of shrubs and small trees, especially slender green, candle-like fruits of Piper plants. Itis one of the most important seed dispersers for Piper and many other plants with small fruits. Breeding. Central American populations of Chestnut Short-tailed Bats in Panama have at least two birth peaks in April-May and August-November, confirming a polyestrous breeding pattern. Nevertheless, these records were taken from widely separate localities, and a fixed breeding pattern might characterize populations in a given area. Activity patterns. The Chestnut Short-tailed Bat is most active 1-3 hours shortly after sunset. Peak activity seems to occur for 3-5 hours of the night. Field data also confirm that it 1s more active during the first one-half of the night, with occasional captures after midnight. Scarcity of records might indicate difficulty of properly identifying it in roosts, or an opportunistic use of roosts where available. Chestnut Short-tailed Bats roost in caves, tunnels, hollow trees, and mines and under overhanging roots. Movements, Home range and Social organization. The Chestnut Short-tailed Bat has small home ranges. It was found sharing a shallow limestone cave with Seba’s Shorttailed Bats ( C. perspicillata ). Status and Conservation. Classified as Least Concern on The IUCN Red List. The Chest nut Short-tailed Bat has a wide distribution and presumably large and stable population. Bibliography. Allen (1890c), Bonaccorso et al. (2007), Fleming (1991), Fleming et al. (1972), Gorchov etal. (1995), Hoffmann & Baker (2003), McLellan & Koopman (2008), Pine (1972), Reid (2009), Solari & Baker (2006), Thies & Kalko (2004), Thies etal. (2006), Wilson (1979).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Carollia castanea	Carollia		castanea	H. Allen	1890	0	Proc. Am. Philos. Soc.	######	Chestnut Short-tailed Bat	None.	Costa Rica, Angostura	E Guatemala to Peru, Venezuela, and Guyana	Not listed.	Least Concern	Does not include colombiana , which is a synonym of brevicauda ; see Zurc and Velazco (2010). Identified as a species complex by Hoffman and Baker (2003), with specimens from much of the southern part of the range described as a new species, benkeithi , by Solari and Baker (2006). Two clades of castanea have been consistently recovered (e.g., castanea a and castanea b of Velazco, 2013) and the species as we recognize it here should be split. However, the distributional limits of the two clades within castanea overlap and it is not clear which clade should bear the name castanea ; see Velazco (2013). The type locality ("Costa Rica") was restricted by Pine (1972). See also McLellan and Koopman (2008), Jarrin-V et al. (2010), and López-Aguirre et al. (2015).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Carollia castanea	23	Chestnut Short-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	CAROLLIINAE	NA	Carollia	NA	castanea	H. Allen	1890	0	Carollia_castanea	Allen, H. (1890). Description of a New Species of Carollia and Remarks on Carollia brevicauda. Proceedings of the American Philosophical Society, 28, 19.	https://books.google.com/books?id=1aWKj8xDMXoC&pg=RA1-PA19&lpg=RA1-PA19&dq=Description+of+a+New+Species+of+Carollia+and+Remarks+on+Carollia+brevicauda.+Proceedings+of+the+American+Philosophical+Society&source=bl&ots=0KGaqYaWsC&sig=ACfU3U0xLZwqaFoD2QmWxvVSoisTsEOdag&hl=en&sa=X&ved=2ahUKEwjyiIOr8P_sAhUFx1kKHZ5cA-4Q6AEwAHoECAgQAg#v=onepage&q=Description%20of%20a%20New%20Species%20of%20Carollia%20and%20Remarks%20on%20Carollia%20brevicauda.%20Proceedings%20of%20the%20American%20Philosophical%20Society&f=false	USNM 12914/36834		"Costa Rica." Restricted by R. H. Pine in 1972 to Angostura, San José, Costa Rica.			castanea H. Allen, 1890	NA	NA	Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Ecuador|Peru	North America|South America	Neotropic	LC	0	0	0	Carollia_castanea	0	sciname match	Carollia_castanea	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Carollia castanea	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Carollia	castanea	H. Allen, 1890	The species previously included populations with distinctive morphologies and karyotypes, that required description of a new species, C. benkeithi , to better recognize that evolutionary diversity (Solari and Baker 2006).	90000000	Carollia castanea	Least Concern		2016	2016-03-14 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, presumed large population, and because it is likely to have stable populations.	Within the genus Carollia , this species is the most specialized in fruits of Piper (Thies and Kalko 2004). It forages near moist areas, being taken most frequently in tropical evergreen forests. It uses the understory vegetation levels, where it concentrates its feeding on the fruits of shrubs and treelets, especially the slender green, candle like fruits of plants of the genus Piper . Due to its high numbers, this bat is one of the most important seed dispersers for Piper and many other plants with small fruits. In Panama, birth peaks occur in March to April and July to August (Wilson 1979, ;Eisenberg 1989, Emmons and Feer 1997). Thies et al.  (2006) report that the activity of this species is not affected by moonlight conditions, and that only strong rain reduces the time it spends flying. If the expected home range is similar to that of other species in the C. castanea complex, it would be close to 6.8 +/- 2.2, as reported by Bonaccorso ;et al.  (2006) for Carollia sp. (listed as C. castanea but most probably the unnamed species indicated by Solari and Baker 2006) in eastern Ecuador.	No major threats are known. The species seems very tolerant to changes in forest cover, and shows no preference for specific habitats.	It is abundant to common in second growth woodland, clearings and plantations, and less common in mature forest (Emmons and Feer 1997, Reid 2009). It is easily caught during sampling in appropriate areas (Davalos pers. comm.), but at other areas it can be uncommon.	Stable	This species occurs throughout Honduras to northwestern Colombia, east to Venezuela and Guyana and south to western Ecuador and northern Peru (Solari and Baker 2006, ;McLellan ;and ;Koopman 2008). This definition is more restricted that the one used by Simmons (2005), which included populations now recognized as a new species, ;C. benkeithi ;(Solari and Baker 2006).		Terrestrial	The species occurs in protected areas throughout its range.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Carollia		castanea	H. Allen	1890	0	Proc. Am. Philos. Soc.	28:19:00	Chestnut Short-tailed Bat	None.	Costa Rica, Angostura	E Guatemala to Peru, Venezuela, and Guyana	Not listed.	Least Concern	Does not include colombiana , which is a synonym of brevicauda ; see Zurc and Velazco (2010). Identified as a species complex by Hoffman and Baker (2003), with specimens from much of the southern part of the range described as a new species, benkeithi , by Solari and Baker (2006). Two clades of castanea have been consistently recovered (e.g., castanea a and castanea b of Velazco, 2013) and the species as we recognize it here should be split. However, the distributional limits of the two clades within castanea overlap and it is not clear which clade should bear the name castanea ; see Velazco (2013). The type locality ("Costa Rica") was restricted by Pine (1972). See also McLellan and Koopman (2008), Jarrin-V et al. (2010), and López-Aguirre et al. (2015).	Carollia castanea	1004874	23	Chestnut Short-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	CAROLLIINAE	NA	Carollia	NA	castanea	H. Allen	1890	0	Carollia_castanea	Allen, H. (1890). Description of a New Species of Carollia and Remarks on Carollia brevicauda. Proceedings of the American Philosophical Society, 28, 19.	https://books.google.com/books?id=1aWKj8xDMXoC&pg=RA1-PA19&lpg=RA1-PA19&dq=Description+of+a+New+Species+of+Carollia+and+Remarks+on+Carollia+brevicauda.+Proceedings+of+the+American+Philosophical+Society&source=bl&ots=0KGaqYaWsC&sig=ACfU3U0xLZwqaFoD2QmWxvVSoisTsEOdag&hl=en&sa=X&ved=2ahUKEwjyiIOr8P_sAhUFx1kKHZ5cA-4Q6AEwAHoECAgQAg#v=onepage&q=Description%20of%20a%20New%20Species%20of%20Carollia%20and%20Remarks%20on%20Carollia%20brevicauda.%20Proceedings%20of%20the%20American%20Philosophical%20Society&f=false	USNM 12914/36834		"Costa Rica." Restricted by R. H. Pine in 1972 to Angostura, San José, Costa Rica.			castanea H. Allen, 1890	NA	NA				Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Ecuador|Peru	North America|South America	Neotropic	LC	0	0	0	Carollia_castanea	0	sciname match	Carollia_castanea	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Carollia_castanea	1004874	23	Chestnut Short-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Carolliinae	NA	Carollia	NA	castanea	H. Allen	0	Carollia castanea	Allen, H. 1890-02-25. Description of a new species of _Carollia_ and remarks on _Carollia brevicauda_. Proceedings of the American Philosophical Society 28:19-26.	https://www.biodiversitylibrary.org/page/6517959	USNM:MAMM:12914 (= USNM:MAMM:A36834)	holotype	http://n2t.net/ark:/65665/31e688b99-b7f5-489e-a80f-df6ba7f776d3	"Costa Rica." Restricted by R. H. Pine in 1972 to Angostura, San José, Costa Rica.			NA	NA				Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Ecuador|Peru	North America|South America	Neotropic	LC	0	0	0	Carollia_castanea	0	sciname match	Carollia_castanea	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Carollia		castanea	H. Allen	1890	0	Proc. Am. Philos. Soc.	28:19:00	Chestnut Short-tailed Bat	None.	Costa Rica, Angostura	E Guatemala to Peru, Venezuela, and Guyana	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/88110411/88110432/' target='_blank'>Least Concern</a>	Does not include colombiana, which is a synonym of brevicauda; see Zurc and Velazco (2010). Identified as a species complex by Hoffman and Baker (2003), with specimens from much of the southern part of the range described as a new species, benkeithi, by Solari and Baker (2006). Two clades of castanea have been consistently recovered (e.g., castanea a and castanea b of Velazco, 2013) and the species as we recognize it here should be split. However, the distributional limits of the two clades within castanea overlap and it is not clear which clade should bear the name castanea; see Velazco (2013). The type locality ("Costa Rica") was restricted by Pine (1972). See also McLellan and Koopman (2008), Jarrin-V et al. (2010), and López-Aguirre et al. (2015).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Carollia castanea; Carollia castanea; Carollia castanea; Carollia castanea; Carollia castanea; castanea; Carollia d Allen; Kastanienbraune Kurzschwanzblattnase; Carolia de Allen; Chestnut Short-tailed Bat; Chestnut Short-tailed Bat;; C. castanea