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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1114	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Mormopterus loriae cobourgiana	Ozimops cobourgianus	Ozimops cobourgianus	Ozimops cobourgianus	Ozimops cobourgianus	Ozimops cobourgianus	Ozimops cobourgianus	Ozimops cobourgianus	Ozimops cobourgianus	Ozimops cobourgianus		[HMW] 7 Tadarida loriae cobourgiana D. H. Johnson, 1959 , “ Black Rock Point ( on north shore of Van Diemen Gulf, 15 miles [= 24 km ] southeast of Cape Don lighthouse ), Cobourg Peninsula , Northern Tewritory , Australia : (Jai. 11° 26'S , long. 131° 56’ E ).” Ozimops cobourgianus has variously been treated within O. loriae or O. planiceps , as either a subspecies or a synonym. Species status was resolved in 2014 by T. B. Reardon and colleagues, who placed it in the Mormopterus subgenus Ozimops , which was then elevated to genus level by S. M. Jackson and C. P. Groves in 2015. Monotypic.; [batnames2022] Distinct from loriae; see Reardon et al. (2014).; [MDD2022] split from O. loriae; moved from Mormopterus to Ozimops; [IUCN] <span class="datalabel1">Johnson (1959) originally described cobourgiana as a subspecies of the small Papua New Guinean species Tadarida loriae . ;The recent taxonomic revision of the genus Mormopterus in Australia (Reardon et al. 2014) elevated M. cobourgianus to a full species. Mormopterus cobourgianus is included in the subgenus Ozimops (Reardon, McKenzie and Adams 2014).; [batnames2023] Distinct from loriae; see Reardon et al. (2014).; [MDD2023] split from O. loriae; moved from Mormopterus to Ozimops; [MDD2025_2.0] split from O. loriae; moved from Mormopterus to Ozimops; [batnames2025_1.7] Distinct from loriae; see Reardon et al. (2014).; [MDD2025_2.2] split from O. loriae; moved from Mormopterus to Ozimops														cobourgianus	<span class="datalabel1">Johnson (1959) originally described cobourgiana as a subspecies of the small Papua New Guinean species Tadarida loriae . ;The recent taxonomic revision of the genus Mormopterus in Australia (Reardon et al. 2014) elevated M. cobourgianus to a full species. Mormopterus cobourgianus is included in the subgenus Ozimops (Reardon, McKenzie and Adams 2014).			cobourgianus	cobourgianus			cobourgianus (D. H. Johnson, 1959)|coburgiana (Koopman, 1994) [incorrect subsequent spelling]						N/A																																								NA																											194287C9FF8EBA23B180F052B6C4FC4D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Molossidae_598.pdf.imf	hash://md5/e57bffb1ffbcba10b412f760b226ffce	669	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/19/42/87/194287C9FF8EBA23B180F052B6C4FC4D.xml	Ozimops cobourgianus	Molossidae	Ozimops	cobourgianus		1959	Tadaride de Cobourg @fr | Nordkisten-Bulldogfledermaus @de | Murciélago rabudo de Cobourg @es | Mangrove Free-tailed Bat @en | North-western Freetail Bat @en | Western Little Free-tailed Bat @en	7 Tadarida loriae cobourgiana D. H. Johnson, 1959 , “ Black Rock Point ( on north shore of Van Diemen Gulf, 15 miles [= 24 km ] southeast of Cape Don lighthouse ), Cobourg Peninsula , Northern Tewritory , Australia : (Jai. 11° 26'S , long. 131° 56’ E ).” Ozimops cobourgianus has variously been treated within O. loriae or O. planiceps , as either a subspecies or a synonym. Species status was resolved in 2014 by T. B. Reardon and colleagues, who placed it in the Mormopterus subgenus Ozimops , which was then elevated to genus level by S. M. Jackson and C. P. Groves in 2015. Monotypic.	Coastal N Australia , in Western Australia (Exmouth E to Broome) and Northern Territory up to the Queensland border, on the Gulf of Carpentaria (including Melville I).	Head—body 45-55 mm ,tail 28-34 mm , forearm 32-35- 1 mm ; weight 6-10- 5 g . Fur on head and back is light orange brown, sometimes with gray-brown frosting; underparts are cream, sometimes yellowy cream, and fur extends well onto wing and knee. Ears are triangular, with straight leading edges and rounded tips. Skin on face and wings is dark brown. Males are distinguishable from all other Ozimops species by their glans penis, which has large epithelial spines covering glans body, from base of glans to bacular mound. Skull is small and slender with no sagittal crest. P* is taller than cingulum of C' and readily visible; width of P, is ¢.80% the size of P,. The species can be distinguished from other Ozimops by a unique combination of allozyme alleles with number of fixed differences ranging from four to eleven.	Largely restricted to mangrove forests and adjacent monsoon floodplain forests and woodlands.	Like most molossids, the Northern Coastal Free-tailed Bat forages in open areas. In the Northern Territory ,it has been found to eat mostly beetles, ants, and leathoppers.	Northern Coastal Free-tailed Bats give birth to single young early in the wet season (December), and young are weaned from March to May.	The Northern Coastal Free-tailed Bat is nocturnal, and may emerge relatively early from roosts, shortly after sunset. It roosts in small spouts and crevices in the dead upper branches of gray mangrove trees (Avicennia marina, Acanthaceae ).	The Northern Coastal Free-tailed Bat has been observed roosting in groups of up to 100 individuals; however, after emergence it forages alone or in pairs.	Classified as Least Concern on The IUCN Red List (as Mormopterus cobourgianus ), because it has a large range (estimated at ¢. 120,000 km ?) with an area of occupancy of ¢. 4700 km * (c. 1200 km * in Western Australia and ¢. 3500 km ? in the Northern Territory ). Population size is difficult to estimate; although the species is not common,it is reliably encountered in mangrove habitat, and, given its extensive area of occupancy, population size is likely to be above 10,000 mature individuals. Much of its habitat is in remote areas, and the population is not subjected to large-scale direct anthropogenic threats other than climate change. In May 2016, a massive area of coastal mangrove dieback was reported on the Gulf of Carpentaria , and although this represents less than 2% of the area of occupancy ofthis species,if further events occur they could lead to declines. Other localized threats come from coastal development, oil spillage and silting from rivers, caused by upstream grazing and industry.	Churchill (2008) | Jackson & Groves (2015) | Milne et al. (2016) | Reardon et al. (2014)		120. Northern Coastal Free-tailed Bat Ozimops cobourgianus French: Tadaride de Cobourg / German: Nordkisten-Bulldogfledermaus / Spanish: Murciélago rabudo de Cobourg Other common names: Mangrove Free-tailed Bat , North-western Freetail Bat , Western Little Free-tailed Bat Taxonomy. 7 Tadarida loriae cobourgiana D. H. Johnson, 1959 , “ Black Rock Point ( on north shore of Van Diemen Gulf, 15 miles [= 24 km ] southeast of Cape Don lighthouse ), Cobourg Peninsula , Northern Tewritory , Australia : (Jai. 11° 26'S , long. 131° 56’ E ).” Ozimops cobourgianus has variously been treated within O. loriae or O. planiceps , as either a subspecies or a synonym. Species status was resolved in 2014 by T. B. Reardon and colleagues, who placed it in the Mormopterus subgenus Ozimops , which was then elevated to genus level by S. M. Jackson and C. P. Groves in 2015. Monotypic. Distribution. Coastal N Australia , in Western Australia (Exmouth E to Broome) and Northern Territory up to the Queensland border, on the Gulf of Carpentaria (including Melville I). Descriptive notes. Head—body 45-55 mm ,tail 28-34 mm , forearm 32-35- 1 mm ; weight 6-10- 5 g . Fur on head and back is light orange brown, sometimes with gray-brown frosting; underparts are cream, sometimes yellowy cream, and fur extends well onto wing and knee. Ears are triangular, with straight leading edges and rounded tips. Skin on face and wings is dark brown. Males are distinguishable from all other Ozimops species by their glans penis, which has large epithelial spines covering glans body, from base of glans to bacular mound. Skull is small and slender with no sagittal crest. P* is taller than cingulum of C' and readily visible; width of P, is ¢.80% the size of P,. The species can be distinguished from other Ozimops by a unique combination of allozyme alleles with number of fixed differences ranging from four to eleven. Habitat. Largely restricted to mangrove forests and adjacent monsoon floodplain forests and woodlands. Food and Feeding. Like most molossids, the Northern Coastal Free-tailed Bat forages in open areas. In the Northern Territory ,it has been found to eat mostly beetles, ants, and leathoppers. Breeding. Northern Coastal Free-tailed Bats give birth to single young early in the wet season (December), and young are weaned from March to May. Activity patterns. The Northern Coastal Free-tailed Bat is nocturnal, and may emerge relatively early from roosts, shortly after sunset. It roosts in small spouts and crevices in the dead upper branches of gray mangrove trees (Avicennia marina, Acanthaceae ). Movements, Home range and Social organization. The Northern Coastal Free-tailed Bat has been observed roosting in groups of up to 100 individuals; however, after emergence it forages alone or in pairs. Status and Conservation. Classified as Least Concern on The IUCN Red List (as Mormopterus cobourgianus ), because it has a large range (estimated at ¢. 120,000 km ?) with an area of occupancy of ¢. 4700 km * (c. 1200 km * in Western Australia and ¢. 3500 km ? in the Northern Territory ). Population size is difficult to estimate; although the species is not common,it is reliably encountered in mangrove habitat, and, given its extensive area of occupancy, population size is likely to be above 10,000 mature individuals. Much of its habitat is in remote areas, and the population is not subjected to large-scale direct anthropogenic threats other than climate change. In May 2016, a massive area of coastal mangrove dieback was reported on the Gulf of Carpentaria , and although this represents less than 2% of the area of occupancy ofthis species,if further events occur they could lead to declines. Other localized threats come from coastal development, oil spillage and silting from rivers, caused by upstream grazing and industry. Bibliography. Churchill (2008), Jackson & Groves (2015), Milne et al. (2016), Reardon et al. (2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Molossidae	Ozimops cobourgianus	Ozimops		cobourgianus	Johnson	1959	1	Proc. Biol. Soc. Wash.	75:05:00	Northwestern Free-tailed Bat	None.	Australia, Northern Territory, Coubourg Peninsula, 15 mi SE of Cape Don Lighthouse, on north shore of Van Dieman Gulf, Black Rock Point	N Australia	Not listed.	Least Concern	Distinct from loriae; see Reardon et al. (2014).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Ozimops cobourgianus	23	Northern Coastal Free-tailed Bat	Mangrove Free-tailed Bat|North-western Freetail Bat|Western Little Free-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MOLOSSIDAE	MOLOSSINAE	NA	Ozimops	NA	cobourgianus	D. H. Johnson	1959	1	Tadarida_loriae_cobourgiana	Johnson, D. H. (1959). Four New Mammals from the Northern Territory of Australia. Proceedings of the Biological Society of Washington, 72, 185.	https://www.biodiversitylibrary.org/page/34641962#page/199/mode/1up	USNM 284243		"Black Rock Point (on north shore of Van Diemen Gulf, 15 miles [= 24 km] southeast of Cape Don lighthouse), Cobourg Peninsula, Northern Territory, Australia (lat. 11Â° 26' S, long. 131Â° 56' E)."	-11.43	131.93	cobourgianus (D. H. Johnson, 1959)	split from O. loriae; moved from Mormopterus to Ozimops	Reardon, T. B., McKenzie, N. L., Cooper, S. J. B., Appleton, B., Carthew, S., & Adams, M. (2014). A molecular and morphological investigation of species boundaries and phylogenetic relationships in Australian free-tailed bats Mormopterus (Chiroptera: Molossidae). Australian journal of zoology, 62(2), 109-136.|Jackson, S. M., & Groves, C. (2015). Taxonomy of Australian mammals. CSIRO Publishing, Austalia, Clayton.	Australia	Oceania	Australasia/Oceania	LC	0	0	0	Ozimops_cobourgianus	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	70000000	Ozimops cobourgianus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MOLOSSIDAE	Ozimops	cobourgianus	(Johnson, 1959)	<span class="datalabel1">Johnson (1959) originally described cobourgiana as a subspecies of the small Papua New Guinean species Tadarida loriae . ;The recent taxonomic revision of the genus Mormopterus in Australia (Reardon et al. 2014) elevated M. cobourgianus to a full species. Mormopterus cobourgianus is included in the subgenus Ozimops (Reardon, McKenzie and Adams 2014).	200000000	Ozimops cobourgianus	Least Concern		2017	2016-07-31 00:00:00 UTC	3.1	English	<p><span lang="EN-AU">Ozimops cobourgianus <span lang="EN-AU"> has both a large extent of occurrence <span lang="EN-AU">(estimated to be around 120,000 kmÂ²) <span lang="EN-AU">and an area of occupancy of approximately 4,700 kmÂ² (1,200 kmÂ² in Western Australia and 3,500 kmÂ² in the Northern Territory). The species is not especially common, although they are reliably encountered in Western Australian mangal habitat. The population size is therefore difficult to estimate, but given that it occupies extensive areas of suitable coastal habitat, population size is likely to exceed a threshold of 10,000 mature individuals. </span></span></p> <p><span lang="EN-AU">There is no empirical data on population trends for this species. Much of its habitat is in remote areas, including some large protected areas in the Northern Territory, and the population is not subjected to large-scale direct anthropogenic threats. However, localised threats exist from coastal development, oil spillage and silting from rivers (from upstream grazing and industry). In May 2016, a massive area of coastal mangrove dieback was reported in the Gulf of Carpentaria, but this represents &lt;2 % of the area of occupancy of this species. </p> <p><span lang="EN-AU">The species is listed as Least Concern, but tentatively so because of the uncertainty about the direction, rate and magnitude of population trend.</span></p>	<p>The Western Little Free-tailed Bat occurs mostly in mangrove forests and woodlands (McKenzie and Bullen 2012), but also in near-coastal Melaleuca forests, rainforests, eucalypt forests, woodlands, open floodplains and saline coastal flats (Milne et al. 2008). The only recorded roost information comes from Western Australia where roosts were found in hollows of mangrove trees.</p><p>Females give birth to a single young between December and February (Milne et al . 2008). The generation length is taken to be 4-5 years, recognising that for insectivorous bats, reproductive maturity is about 10 months, and longevity about 8-10 years; generation length is taken as the midpoint of these values.</p>	<p><span lang="EN-US">At present, no significant threats have been identified for O. cobourgianus . ;To some extent, the relative remoteness of the mangrove communities that this species depends on, affords some protection from large-scale human mediated threats.</p> <p><span lang="EN-US">However, threats have been recognised at a local level. ;Some areas of mangroves have been cleared for development associated with mining transport in the Pilbara region (Armstrong 2011). Surface run-off from grazing and mining may cause silting in some mangrove areas. ;Extensive mangrove dieback has been reported recently for an extensive area in the Gulf of Carpentaria over about 300 km of the coastline that forms the eastern extent of its distribution (N. Duke pers. comm. 2016). The cause of this dieback has yet to be determined. ;It is also not known if the death of these trees has or will affect local population of O. cobourgianus .</span></p> <p><span lang="EN-US">The potential impact of sea-level rise from climate change on the mangal habitat in northern Australia has not been considered in the context of the requirements of this bat species, but it should be monitored. In addition, changes in rainfall patterns and humidity could affect breeding seasonal breeding aggregations.</p>	<p>The population size is unknown. Milne et al . (2008) considered it â€œcommonâ€, and noted that â€œit occurs sparsely across its rangeâ€. They also reported â€œspectacular, but transient, aggregationsâ€ with â€œswarms of up to one hundredâ€ in some areas, but this is highly seasonal and may represent the aggregations of a large proportion of the breeding population, at least in Western Australia (N.L. McKenzie pers. comm. 2016). At other times of the year the species may be difficult to encounter in these areas. There is a large distribution gap along the Kimberley coast in Western Australia and the western part of the Top End of the Northern Territory, suggesting that at least two isolated populations exist. </p>	Unknown	<span lang="EN-AU">Ozimops cobourgianus <span lang="EN-AU"> occurs in coastal and near coastal areas (within 100 km of the coastline; Milne and Pavey 2011) between Shark Bay in Western Australia and the Northern Territory-Queensland border, although there is a major gap in this distribution in the higher rainfall northern and eastern Kimberley and western Top End of the Northern Territory (Milne et al . 2008, McKenzie and Bullen 2012). It has been recorded from Koolan, Lachlan and Sunday Islands in the south-west Kimberley, but from no other islands in that state (McKenzie and Bullen 2012). In Western Australia the distribution is associated with the coastal mangrove fringe, while in the Northern Territory the distribution extends further inlandâ€”up to 100 km from the coast.</span>		Terrestrial	<p><span lang="EN-US">No specific conservation programmes include this species, but it occurs in several national parks and conservation reserves.</p>	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Molossidae	Ozimops		cobourgianus	Johnson	1959	1	Proc. Biol. Soc. Wash.	75:05:00	Northwestern Free-tailed Bat	None.	Australia, Northern Territory, Coubourg Peninsula, 15 mi SE of Cape Don Lighthouse, on north shore of Van Dieman Gulf, Black Rock Point	N Australia	Not listed.	Least Concern	Distinct from loriae; see Reardon et al. (2014).	Ozimops cobourgianus	1005247	23	Northern Coastal Free-tailed Bat	Mangrove Free-tailed Bat|North-western Freetail Bat|Western Little Free-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Molossidae	MOLOSSINAE	NA	Ozimops	NA	cobourgianus	D. H. Johnson	1959	1	Tadarida_loriae_cobourgiana	Johnson, D. H. (1959). Four New Mammals from the Northern Territory of Australia. Proceedings of the Biological Society of Washington, 72, 185.	https://www.biodiversitylibrary.org/page/34641962#page/199/mode/1up	USNM 284243		"Black Rock Point (on north shore of Van Diemen Gulf, 15 miles [= 24 km] southeast of Cape Don lighthouse), Cobourg Peninsula, Northern Territory, Australia (lat. 11Â° 26' S, long. 131Â° 56' E)."	-11.4333	131.9333	cobourgianus (D. H. Johnson, 1959)	split from O. loriae; moved from Mormopterus to Ozimops	Reardon, T. B., McKenzie, N. L., Cooper, S. J. B., Appleton, B., Carthew, S., & Adams, M. (2014). A molecular and morphological investigation of species boundaries and phylogenetic relationships in Australian free-tailed bats Mormopterus (Chiroptera: Molossidae). Australian journal of zoology, 62(2), 109-136.|Jackson, S. M., & Groves, C. (2015). Taxonomy of Australian mammals. CSIRO Publishing, Austalia, Clayton.				Australia	Oceania	Australasia/Oceania	LC	0	0	0	Ozimops_cobourgianus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Ozimops_cobourgianus	1005247	23	Northern Coastal Free-tailed Bat	Mangrove Free-tailed Bat|Northwestern Freetail Bat|Western Little Free-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Molossidae	Molossinae	NA	Ozimops	NA	cobourgianus	D. H. Johnson	1	Tadarida loriae cobourgiana	Johnson, D.H. 1959-11-04. Four new mammals from the Northern Territory of Australia. Proceedings of the Biological Society of Washington 72:183-188.	https://www.biodiversitylibrary.org/page/34641964	USNM:MAMM:284243	holotype	http://n2t.net/ark:/65665/321ed80a9-976a-4e20-aa1e-9dd8e61efa6b	"Black Rock Point (on north shore of Van Diemen Gulf, 15 miles [= 24 km] southeast of Cape Don lighthouse), Cobourg Peninsula, Northern Territory, Australia (lat. 11Â° 26' S, long. 131Â° 56' E)."	-11.4333	131.9333	split from O. loriae; moved from Mormopterus to Ozimops	Reardon, T. B., McKenzie, N. L., Cooper, S. J. B., Appleton, B., Carthew, S., & Adams, M. (2014). A molecular and morphological investigation of species boundaries and phylogenetic relationships in Australian free-tailed bats Mormopterus (Chiroptera: Molossidae). Australian journal of zoology, 62(2), 109-136.|Jackson, S. M., & Groves, C. (2015). Taxonomy of Australian mammals. CSIRO Publishing, Austalia, Clayton.				Australia	Oceania (Continent)	Australasia	LC	0	0	0	Ozimops_cobourgianus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Molossidae	Ozimops		cobourgianus	Johnson	1959	1	Proc. Biol. Soc. Wash.	75:05:00	Northwestern Free-tailed Bat	None.	Australia, Northern Territory, Coubourg Peninsula, 15 mi SE of Cape Don Lighthouse, on north shore of Van Dieman Gulf, Black Rock Point	N Australia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/71536513/209550699/' target='_blank'>Least Concern</a>	Distinct from loriae; see Reardon et al. (2014).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Ozimops cobourgianus; Ozimops cobourgianus; Ozimops cobourgianus; Ozimops cobourgianus; Ozimops cobourgianus; cobourgianus; Tadaride de Cobourg; Nordkisten-Bulldogfledermaus; Murciélago rabudo de Cobourg; Mangrove Free-tailed Bat; North-western Freetail Bat; Western Little Free-tailed Bat; Northern Coastal Free-tailed Bat; Mangrove Free-tailed Bat; North-western Freetail Bat; Western Little Free-tailed Bat; Northwestern Free-tailed Bat; O. cobourgianus
