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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1101	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri	Nyctophilus walkeri		[HMW] Nyctophilus walker Thomas, 1892 , “Adelaide River, Northern Territory , Australia .” Nyctophilus walkeri is tentatively in the macrotis species group, but its exact relationships to other species of Nyctophilus are unresolved and require additional analysis. Monotypic.; [batnames2022]  microtis species group.; [batnames2023]  microtis species group.; [batnames2025_1.7] microtis species group.														walkeri				walkeri 	walkeri 			walkeri O. Thomas, 1892		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		N Northern Terr., Australia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nyctophilus walkeri	Australia, Northern Territory, Adelaide River.	Thomas	1892	Ann. Mag. Nat. Hist., ser. 6, 9:405.	Distribution: Confined to extreme northern Australia from northeastern Western Australia to northwestern Queensland.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Pygmy long-eared bat	NW Australia	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1892	Ann. Mag. Nat. Hist., ser. 6, 9:405.		Northern Territory and N Western Australia (Australia).	Australia, Northern Territory, Adelaide River.		THOMAS	1892	Size relatively small (forearm length, 31-35mm; condylobasal length, 11-13 mm). Noseleaves poorly developed. Band connecting ears fairly high. Teeth relatively large.	Distribution: Confined to extreme northern Australia from northeastern Western Australia to northwestern Queensland.	No subspecies.		130	species	N. walkeri	THOMAS	1892	Nyctophilus	genus	Nyctophilus walkeri				Size relatively small (forearm length, 31-35mm; condylobasal length, 11-13 mm). Noseleaves poorly developed. Band connecting ears fairly high. Teeth relatively large.	No subspecies.		1. N. walkeri THOMAS 1892.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nyctophilini	Nyctophilus walkeri	Nyctophilus		walkeri	Thomas		1892		Ann. Mag. Nat. Hist., ser. 6	9		405		Pygmy Long-eared Bat	Australia, Northern Territory, Adelaide River.	Northern Territory and N Western Australia (Australia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).			4C3D87E8FFD56A6BFA579FAE1D41BD19	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	804	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFD56A6BFA579FAE1D41BD19.xml	Nyctophilus walkeri	Vespertilionidae	Nyctophilus	walkeri	Thomas	1892	Nyctophile de Walker @fr | Walker Langohrfledermaus @de | Nictofila de Walker @es	Nyctophilus walker Thomas, 1892 , “Adelaide River, Northern Territory , Australia .” Nyctophilus walkeri is tentatively in the macrotis species group, but its exact relationships to other species of Nyctophilus are unresolved and require additional analysis. Monotypic.	NE Western Australia , N Northern Territory , and NW Queensland , N Australia .	Head-body 38-44 mm, tail 26-36 mm, ear 10-7-14-1 mm, forearm 30-1-36 mm; weight 3-7 g. The Pygmy Longeared Bat is the smallest species of Nyctophilus , with relatively large ears and unique simple noseleaf consisting of two ridges, one further on muzzle and another immediately above nostrils, with vertical groove in middle and furred trough between them. Dorsal pelage is pale orangish brown to fawn; venter is cream or buff. Wing membranes are blackish brown, and face, ears, and wing bones are pinkish brown. Rostrum is short and blunt, with ridge across muzzle over nostrils that is well developed with deep but thin medial notch. Ears are large and broad, with bluntly rounded tips but are much smaller, narrower, and more tapered at tip compared with other species of Nyctophilus , having horizontal ribbing on inner surfaces, inward curved anterior edges, and smooth posterior edges (ears can fold back at top of thick part of anterior edge); interauricular membrane typical of the genus is either missing or barely present and covered by fur; tragusis almost rectangular, being straight or very slightly convex on anterior margin. Baculum has distinct distal notch, curves downward at base,is bifurcated basally, and is straight for the rest of shaft. Skull is robust, tympanic bullae are small, and M? and lower molars are not reduced.	Riverine Melaleuca ( Myrtaceae ) and Pandanus ( Pandanaceae ) forests, monsoon rainforest patches, open savanna woodlands, tall open forests, and mixed shrublands, generally associated with permanent water sources.	Pygmy Long-eared Bats forage by aerial-hawking and gleaning off the ground and foliage. They usually fly within 1-2 m of the ground or water surface and are highly maneuverable in cluttered vegetation. Their flight is jerky and irregular, with 2-3 wing flaps before a glide, which might be an adaptation to avoid predation. Diet consists mostly of moths and beetles in the Top End region, but cockroaches, termites, mosquitoes, leafhoppers, lacewings, and spiders were also eaten.	A female Pygmy LLong-eared Bat captured in September in the Northern Territory was in early pregnancy with twins, and three females were captured in late pregnancy in mid-October. A pregnant female with a fetus in each uterine horn was captured in October in the Kimberley region. Young are likely born around early November.	Pygmy LLong-eared Bats spend the day roosting in foliage and forage throughout the night. They have been reported roosting in dead fronds of large Livistona ( Arecaceae ) palms and have been seen emerging from Pandanus thickets at dusk. Call shape is steep FM sweep, with peak frequencies of40-7-50-8 kHz (mean 45-8 kHz) in the Top End region. In Western Australia , peak frequencies were 54-58 kHz (mean 56 kHz) and were noteasily distinguished from other species of long-eared bat.	Pygmy [Long-eared Bats reportedly roost in large enough colonies that they can be heard squabbling with each other during the day, but no specific numbers are available. They appearto be long-lived because captured individuals commonly have teeth worn down to nubs that are virtually flush with gums.	Classified as Least Concern on The IUCN Red List, as it is locally common in much ofits distribution and does not seem to face any major threats.	Bullen & McKenzie (2002a) | Churchill (2008) | Churchill et al. (1984) | McKenzie, Lumsden & Parnaby (2008) | Milne (2002) | Milne etal. (2016) | Parnaby (2009)	https://zenodo.org/record/6397950/files/figure.png	86. Pygmy Long-eared Bat Nyctophilus walkeri French: Nyctophile de Walker / German: WalkerLangohrfledermaus / Spanish: Nictofila de Walker Taxonomy. Nyctophilus walker Thomas, 1892 , “Adelaide River, Northern Territory , Australia .” Nyctophilus walkeri is tentatively in the macrotis species group, but its exact relationships to other species of Nyctophilus are unresolved and require additional analysis. Monotypic. Distribution. NE Western Australia , N Northern Territory , and NW Queensland , N Australia . Descriptive notes. Head-body 38-44 mm, tail 26-36 mm, ear 10-7-14-1 mm, forearm 30-1-36 mm; weight 3-7 g. The Pygmy Longeared Bat is the smallest species of Nyctophilus , with relatively large ears and unique simple noseleaf consisting of two ridges, one further on muzzle and another immediately above nostrils, with vertical groove in middle and furred trough between them. Dorsal pelage is pale orangish brown to fawn; venter is cream or buff. Wing membranes are blackish brown, and face, ears, and wing bones are pinkish brown. Rostrum is short and blunt, with ridge across muzzle over nostrils that is well developed with deep but thin medial notch. Ears are large and broad, with bluntly rounded tips but are much smaller, narrower, and more tapered at tip compared with other species of Nyctophilus , having horizontal ribbing on inner surfaces, inward curved anterior edges, and smooth posterior edges (ears can fold back at top of thick part of anterior edge); interauricular membrane typical of the genus is either missing or barely present and covered by fur; tragusis almost rectangular, being straight or very slightly convex on anterior margin. Baculum has distinct distal notch, curves downward at base,is bifurcated basally, and is straight for the rest of shaft. Skull is robust, tympanic bullae are small, and M? and lower molars are not reduced. Habitat. Riverine Melaleuca ( Myrtaceae ) and Pandanus ( Pandanaceae ) forests, monsoon rainforest patches, open savanna woodlands, tall open forests, and mixed shrublands, generally associated with permanent water sources. Food and Feeding. Pygmy Long-eared Bats forage by aerial-hawking and gleaning off the ground and foliage. They usually fly within 1-2 m of the ground or water surface and are highly maneuverable in cluttered vegetation. Their flight is jerky and irregular, with 2-3 wing flaps before a glide, which might be an adaptation to avoid predation. Diet consists mostly of moths and beetles in the Top End region, but cockroaches, termites, mosquitoes, leafhoppers, lacewings, and spiders were also eaten. Breeding. A female Pygmy LLong-eared Bat captured in September in the Northern Territory was in early pregnancy with twins, and three females were captured in late pregnancy in mid-October. A pregnant female with a fetus in each uterine horn was captured in October in the Kimberley region. Young are likely born around early November. Activity patterns. Pygmy LLong-eared Bats spend the day roosting in foliage and forage throughout the night. They have been reported roosting in dead fronds of large Livistona ( Arecaceae ) palms and have been seen emerging from Pandanus thickets at dusk. Call shape is steep FM sweep, with peak frequencies of40-7-50-8 kHz (mean 45-8 kHz) in the Top End region. In Western Australia , peak frequencies were 54-58 kHz (mean 56 kHz) and were noteasily distinguished from other species of long-eared bat. Movements, Home range and Social organization. Pygmy [Long-eared Bats reportedly roost in large enough colonies that they can be heard squabbling with each other during the day, but no specific numbers are available. They appearto be long-lived because captured individuals commonly have teeth worn down to nubs that are virtually flush with gums. Status and Conservation. Classified as Least Concern on The IUCN Red List, as it is locally common in much ofits distribution and does not seem to face any major threats. Bibliography. Bullen & McKenzie (2002a), Churchill (2008), Churchill et al. (1984), McKenzie, Lumsden & Parnaby (2008), Milne (2002), Milne etal. (2016), Parnaby (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Nyctophilus walkeri	Nyctophilus		walkeri	Thomas	1892	0	Ann. Mag. Nat. Hist.	ser. 6, 9: 405	Pygmy Long-eared Bat	None.	Australia, Northern Territory, Adelaide River.	Northern Territory and N Western Australia (Australia).	Not listed.	Least Concern	 microtis species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctophilus walkeri	23	Pygmy Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Nyctophilus	NA	walkeri	O. Thomas	1892	0	Nyctophilus_Walkeri	Thomas, O. (1892). Description of a third species of the genus Nyctophilus. Annals and Magazine of Natural History, ser. 6, 9, 406.	https://www.biodiversitylibrary.org/item/63496#page/452/mode/1up	BM 1892.4.4.1		"Adelaide River, Northern Territory, Australia."			walkeri O. Thomas, 1892	NA	NA	Australia	Oceania	Australasia/Oceania	LC	0	0	0	Nyctophilus_walkeri	0	sciname match	Nyctophilus_walkeri	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	15011	Nyctophilus walkeri	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Nyctophilus	walkeri	Thomas, 1892		20000000	Nyctophilus walkeri	Least Concern		2020	2019-07-09 00:00:00 UTC	3.1	English	This species is listed as Least Concern because it is relatively wide-ranging, occurs in a number of protected areas, and it is not thought to be declining.	This species is associated with watercourses ; ; surrounded by melaleuca and pandanus, gallery forest or Livistona palms (Churchill et al. 2008) generally in rocky escarpment areas.	There appear to be no major threats to this species although physical disturbance of riparian vegetation though clearing or livestock may impact the habitat of this species.	Though previously considered rare, this species is now known to be locally common within its range.	Stable	This species is endemic to northern Australia, where it is found in northern Western Australia, northern Northern Territory, and at Lawn Hill Gorge, Queensland.		Terrestrial	The species has been recorded from a number of protected areas.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Nyctophilus		walkeri	Thomas	1892	0	Ann. Mag. Nat. Hist.	ser. 6, 9: 406	Pygmy Long-eared Bat	None.	Australia, Northern Territory, Adelaide River	Northern Territory and N Western Australia (Australia)	Not listed.	Least Concern	 microtis species group.	Nyctophilus walkeri	1005767	23	Pygmy Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Nyctophilus	NA	walkeri	O. Thomas	1892	0	Nyctophilus_Walkeri	Thomas, O. (1892). Description of a third species of the genus Nyctophilus. Annals and Magazine of Natural History, ser. 6, 9, 406.	https://www.biodiversitylibrary.org/item/63496#page/452/mode/1up	BM 1892.4.4.1		"Adelaide River, Northern Territory, Australia."			walkeri O. Thomas, 1892	NA	NA				Australia	Oceania	Australasia/Oceania	LC	0	0	0	Nyctophilus_walkeri	0	sciname match	Nyctophilus_walkeri	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctophilus_walkeri	1005767	23	Pygmy Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Nyctophilus	NA	walkeri	O. Thomas	0	Nyctophilus Walkeri	Thomas, O. 1892-05-01. Description of a third species of the genus _Nyctophilus_. Annals and Magazine of Natural History (6)9(53):405-406.	https://www.biodiversitylibrary.org/page/19268646	BMNH:Mamm:1892.4.4.1	holotype	https://data.nhm.ac.uk/object/b8f2b562-3540-4233-8e62-0d26ca7fe19d	"Adelaide River, Northern Territory, Australia."			NA	NA				Australia	Oceania (Continent)	Australasia	LC	0	0	0	Nyctophilus_walkeri	0	sciname match	Nyctophilus_walkeri	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Nyctophilus		walkeri	Thomas	1892	0	Ann. Mag. Nat. Hist.	ser. 6, 9: 406	Pygmy Long-eared Bat	None.	Australia, Northern Territory, Adelaide River	Northern Territory and N Western Australia (Australia)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/15011/22003303/' target='_blank'>Least Concern</a>	microtis species group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctophilus walkeri; Nyctophilus walkeri; Nyctophilus walkeri; Nyctophilus walkeri; Nyctophilus walkeri; Nyctophilus walkeri; walkeri; Nyctophile de Walker; Walker Langohrfledermaus; Nictofila de Walker; Pygmy Long-eared Bat; Pygmy Long-eared Bat; Pygmy Long-eared Bat; N. walkeri