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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L12	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	Pipistrellus helios	N/A	Pipistrellus africanus helios	Neoromicia helios	Neoromicia helios	Afronycteris helios	Afronycteris helios	Neoromicia helios	Afronycteris helios	Afronycteris helios	Afronycteris helios	Afronycteris helios	Afronycteris helios		[MSW3] Considered to be a subspecies of nanus by some authors, but differences in bacular morphology (Hill and Harrison, 1987), roosting and social behavior (O'Shea, 1980; Happold and Happold, 1996), habitat and pelage coloration, and the presence of a pair of glands on the interfemoral membrane in helios (O'Shea, 1980) that are rarely found in nanus indicate that helios is a distinct species (M. Happold, pers. comm.).; [HMW] Pipistrellus helios Heller, 1912 , Merelle Water, 30 miles (= 48 km ) south Mount Marsabit , Kenya . Traditionally treated as a race of N. nanus , but split as a separate species based on the shape of the baculum. Monotypic.; [batnames2022] Previously included in Neoromicia ; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Considered to be a subspecies of nanus by some authors, but differences in bacular morphology (Hill and Harrison, 1987), roosting andsocial behavior (O'Shea, 1980; Happold and Happold, 1996), habitat and pelage coloration, and the presence of a pair of glands on theinterfemoral membrane in helios (O'Shea, 1980) that are rarely found in nanus indicate that helios is a distinct species (M. Happold, pers. comm.).; [MDD2022] moved from Neoromicia to the recently described Afronycteris, although further genetic studies will be needed to cement this arrangement; [IUCN] This is considered to be a subspecies of Pipistrellus nanus ;(currently Neoromicia nana ) by some authors (Simmons 2005).; [batnames2023] Previously included in Neoromicia ; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Considered to be a subspecies of nanus by some authors, but differences in bacular morphology (Hill and Harrison, 1987), roosting andsocial behavior (O'Shea, 1980; Happold and Happold, 1996), habitat and pelage coloration, and the presence of a pair of glands on theinterfemoral membrane in helios (O'Shea, 1980) that are rarely found in nanus indicate that helios is a distinct species (M. Happold, pers. comm.).; [MDD2023] moved from Neoromicia to the recently described Afronycteris, although further genetic studies will be needed to cement this arrangement; [MDD2025_2.0] moved from Neoromicia to the recently described Afronycteris, although further genetic studies will be needed to cement this arrangement; [batnames2025_1.7] Previously included in Neoromicia; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Considered to be a subspecies of nanus by some authors, but differences in bacular morphology (Hill and Harrison, 1987), roosting andsocial behavior (O'Shea, 1980; Happold and Happold, 1996), habitat and pelage coloration, and the presence of a pair of glands on theinterfemoral membrane in helios (O'Shea, 1980) that are rarely found in nanus indicate that helios is a distinct species (M. Happold, pers. comm.).; [MDD2025_2.2] moved from Neoromicia to the recently described Afronycteris, although further genetic studies will be needed to cement this arrangement														helios	This is considered to be a subspecies of Pipistrellus nanus ;(currently Neoromicia nana ) by some authors (Simmons 2005).			helios	helios			helios (E. Heller, 1912)						N/A							Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Kenya, Sudan; ref. 4.123																															NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Neoromicia helios	Neoromicia		helios	Heller	y	1912		Smiths. Misc. Coll.	60	12	3		Samburu Pipistrelle	Kenya, 30 mi S Mt. Marsabit, Merelle Water.	Kenya, Somalia, Djibouti, NE Uganda, extreme S Sudan, N Tanzania. Maybe more widespread (Peterson, 1987).	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).		Considered to be a subspecies of nanus by some authors, but differences in bacular morphology (Hill and Harrison, 1987), roosting and social behavior (O'Shea, 1980; Happold and Happold, 1996), habitat and pelage coloration, and the presence of a pair of glands on the interfemoral membrane in helios (O'Shea, 1980) that are rarely found in nanus indicate that helios is a distinct species (M. Happold, pers. comm.).	4C3D87E8FFB96A06FF499FC51787BCF0	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	824	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFB96A06FF499FC51787BCF0.xml	Neoromicia helios	Vespertilionidae	Neoromicia	helios		1912	Vespeére soleil @fr | Heller Zwergfledermaus @de | Neoromicia de Heller @es | Helios Pipistrelle Bat @en | Heller's Pipistrelle @en | Heller's Pipistrelle Bat @en | Samburu Pipistrelle @en | Samburu Pipistrelle Bat @en	Pipistrellus helios Heller, 1912 , Merelle Water, 30 miles (= 48 km ) south Mount Marsabit , Kenya . Traditionally treated as a race of N. nanus , but split as a separate species based on the shape of the baculum. Monotypic.	Extreme S South Sudan , S Somalia , NE & E Uganda , Kenya , and N Tanzania ; isolated record from Djibouti needs confirmation.	Head-body ¢.34-41 mm, tail 26-34 mm, ear 5-11 mm, forearm 26-30 mm; weight 2-6 g. Males average larger than females. Pelage of Heller’s Serotine is dense and soft; dorsally pale brown to pale yellowish brown (hairs tricolored, with basal one-quarter blackish brown, central onehalf pale creamy fawn, and terminal one-quarter pale brown; mid-dorsal hairs 5-7 mm long); ventrally paler (hairs blackish brown for basal three-fifths, with pale brown tips). Wings are dark brown to blackish, with very narrow to moderate white hind border; interfemoral membrane is semi-translucent and paler than wings. Ears are pale brown, with rounded tips; tragus is about one-half ear length, broadest above midheight, hatched-shaped, with anterior margin slightly to distinctly concave, posterior margin with abrupt, obtuse angle above mid-height, and rounded tip. Most adults of both sexes have a pair of glands (1-2-3-8 mm in diameter), one on each side oftail, in proximal region of interfemoral membrane; function of these glands unknown, and in males there is no correlation with testis size. Skull is small for an African pipistrellelike bat (greatest skull lengths 10-4-11-4 mm); profile of forehead is strongly concave; there is no occipital helmet. I? is bicuspid and broad; I’ reaches posterior cusp of I*; P? is present and well developed, displaced lingually, but visible in labial view; lower molars are myotodont. Dental formulais12/3,C1/1,P 2/2, M 3/3 (x2) = 34.	Widely reported from dry habitats, but also from East African coastal forest mosaic, Isoberlinia ( Fabaceae ) woodland, undifferentiated montane vegetation, and mosaics of lowland rainforest and secondary grassland, at 400-1333 m.	Heller's Serotine feeds on small insects, foraging by slow hawking 2-5 m aboveground.	In Masalani, eastern Kenya , Heller’s Serotine shows restricted seasonal monoestry, with births in November, and lactation until late December or early January. There is no evidence of testicular activity until March, when testes descend, remaining in scrotal position from April through September. Based on this, copulation may occur from May to August, when males are more vocalat their roosts, and ovulation and fertilization in late August. In Masalani, the overall ratio of males to females was 2:1. A seasonal phase of dispersal was observed in late May to mid-June, when some adults left and other joined the population. Litter size is two.	Heller's Serotine roosts by day, and intermittently during the night, between overlapping, vertically hanging fronds of wild date palms ( Phoenix reclinata , Arecaceae ), in small crevices around edges of roofs thatched with bundles of palm fronds, and in crevices in vertical sections of thatch used to cover walls. Enters torpor during the day. Echolocation parameters recorded as: search-phase call shape steep FM/QCEF, intensity high,start frequency c.115 kHz, and end frequency 65 kHz. Predators include Heart-nosed False-vampires (Cardioderma cor) and possibly tree snakes.	Heller’s Serotines roost singly or in groups of 2-12; during most of the year, most adult males roosted singly or with one female or, less frequently, with two or more females. Adult females roosted with one male, but sometimes with two other females and without a male. Adult males almost never roosted with other males. Females showed roost fidelity, competing aggressively for prime roosts (those large enough to accommodate one male and one or two females), fighting intruders and chasing them in flight. Bats were active in and near roosts throughout the night. While some bats rested, territorial males patrolled outside their roosts, chasing other males away. Males vocalized at their roosts and females were attracted to the most vocal males. Females roosted with several males sequentially, showing preference for certain individuals. Prior to parturition in November, all females left the study area, presumably to establish a maternity colony. These data suggest a mating system based on resource-defense polygyny.	Classified as Data Deficient on The IUCN Red List.	Happold & Van Cakenberghe (2013) | Hayman & Hill (1971) | Hill & Harrison (1987) | Koopman (1975, 1993) | O'Shea (1980) | O'Shea & Vaughan (1980) | Peterson (1987) | Simmons (2005)	https://zenodo.org/record/6398053/files/figure.png	133. Heller’s Serotine Neoromicia helios French: Vespeére soleil / German: HellerZwergfledermaus / Spanish: Neoromicia de Heller Other common names: Helios Pipistrelle Bat , Heller's Pipistrelle , Heller's Pipistrelle Bat , Samburu Pipistrelle , Samburu Pipistrelle Bat Taxonomy. Pipistrellus helios Heller, 1912 , Merelle Water, 30 miles (= 48 km ) south Mount Marsabit , Kenya . Traditionally treated as a race of N. nanus , but split as a separate species based on the shape of the baculum. Monotypic. Distribution. Extreme S South Sudan , S Somalia , NE & E Uganda , Kenya , and N Tanzania ; isolated record from Djibouti needs confirmation. Descriptive notes. Head-body ¢.34-41 mm, tail 26-34 mm, ear 5-11 mm, forearm 26-30 mm; weight 2-6 g. Males average larger than females. Pelage of Heller’s Serotine is dense and soft; dorsally pale brown to pale yellowish brown (hairs tricolored, with basal one-quarter blackish brown, central onehalf pale creamy fawn, and terminal one-quarter pale brown; mid-dorsal hairs 5-7 mm long); ventrally paler (hairs blackish brown for basal three-fifths, with pale brown tips). Wings are dark brown to blackish, with very narrow to moderate white hind border; interfemoral membrane is semi-translucent and paler than wings. Ears are pale brown, with rounded tips; tragus is about one-half ear length, broadest above midheight, hatched-shaped, with anterior margin slightly to distinctly concave, posterior margin with abrupt, obtuse angle above mid-height, and rounded tip. Most adults of both sexes have a pair of glands (1-2-3-8 mm in diameter), one on each side oftail, in proximal region of interfemoral membrane; function of these glands unknown, and in males there is no correlation with testis size. Skull is small for an African pipistrellelike bat (greatest skull lengths 10-4-11-4 mm); profile of forehead is strongly concave; there is no occipital helmet. I? is bicuspid and broad; I’ reaches posterior cusp of I*; P? is present and well developed, displaced lingually, but visible in labial view; lower molars are myotodont. Dental formulais12/3,C1/1,P 2/2, M 3/3 (x2) = 34. Habitat. Widely reported from dry habitats, but also from East African coastal forest mosaic, Isoberlinia ( Fabaceae ) woodland, undifferentiated montane vegetation, and mosaics of lowland rainforest and secondary grassland, at 400-1333 m. Food and Feeding. Heller's Serotine feeds on small insects, foraging by slow hawking 2-5 m aboveground. Breeding. In Masalani, eastern Kenya , Heller’s Serotine shows restricted seasonal monoestry, with births in November, and lactation until late December or early January. There is no evidence of testicular activity until March, when testes descend, remaining in scrotal position from April through September. Based on this, copulation may occur from May to August, when males are more vocalat their roosts, and ovulation and fertilization in late August. In Masalani, the overall ratio of males to females was 2:1. A seasonal phase of dispersal was observed in late May to mid-June, when some adults left and other joined the population. Litter size is two. Activity patterns. Heller's Serotine roosts by day, and intermittently during the night, between overlapping, vertically hanging fronds of wild date palms ( Phoenix reclinata , Arecaceae ), in small crevices around edges of roofs thatched with bundles of palm fronds, and in crevices in vertical sections of thatch used to cover walls. Enters torpor during the day. Echolocation parameters recorded as: search-phase call shape steep FM/QCEF, intensity high,start frequency c.115 kHz, and end frequency 65 kHz. Predators include Heart-nosed False-vampires (Cardioderma cor) and possibly tree snakes. Movements, Home range and Social organization. Heller’s Serotines roost singly or in groups of 2-12; during most of the year, most adult males roosted singly or with one female or, less frequently, with two or more females. Adult females roosted with one male, but sometimes with two other females and without a male. Adult males almost never roosted with other males. Females showed roost fidelity, competing aggressively for prime roosts (those large enough to accommodate one male and one or two females), fighting intruders and chasing them in flight. Bats were active in and near roosts throughout the night. While some bats rested, territorial males patrolled outside their roosts, chasing other males away. Males vocalized at their roosts and females were attracted to the most vocal males. Females roosted with several males sequentially, showing preference for certain individuals. Prior to parturition in November, all females left the study area, presumably to establish a maternity colony. These data suggest a mating system based on resource-defense polygyny. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Bibliography. Happold & Van Cakenberghe (2013), Hayman & Hill (1971), Hill & Harrison (1987), Koopman (1975, 1993), O'Shea (1980), O'Shea & Vaughan (1980), Peterson (1987), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Afronycteris helios	Afronycteris		helios	Heller	1912	1	Smiths. Misc. Coll.	60(12): 3	Samburu Serotine	None.	Kenya, 30 mi S Mt. Marsabit, Merelle Water.	Kenya, Somalia, Djibouti, NE Uganda, extreme S Sudan, N Tanzania. Maybe more widespread (Peterson, 1987).	Not listed.	Data Deficient as Neoromicia helios 	Previously included in Neoromicia ; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Considered to be a subspecies of nanus by some authors, but differences in bacular morphology (Hill and Harrison, 1987), roosting andsocial behavior (O'Shea, 1980; Happold and Happold, 1996), habitat and pelage coloration, and the presence of a pair of glands on theinterfemoral membrane in helios (O'Shea, 1980) that are rarely found in nanus indicate that helios is a distinct species (M. Happold, pers. comm.).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Afronycteris helios	23	Heller's Serotine	Helios Pipistrelle Bat|Heller's Pipistrelle|Samburu Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Afronycteris	NA	helios	E. Heller	1912	1	Pipistrellus_helios	Heller, E. (1912). New Races of Insectivores, Bats, and Lemurs from British East Africa. Smithsonian Miscellaneous Collections, 60(12), 3	https://repository.si.edu/bitstream/handle/10088/23473/SMC_60_Heller_1912_12_1-13.pdf?sequence=1&isAllowed=y	USNM 181813		Merelle Water, 30 miles (= 48 km) south Mount Marsabit, Kenya.			helios (E. Heller, 1912)	moved from Neoromicia to the recently described Afronycteris, although further genetic studies will be needed to cement this arrangement	Monadjem, A., Demos, T. C., Dalton, D. L., Webala, P. W., Musila, S., Kerbis Peterhans, J. C., & Patterson, B. D. (2020). A revision of the pipistrelle-like bats (Mammali: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society, zlaa087.	South Sudan|Somalia|Uganda|Kenya|Tanzania|Djibouti?	Africa	Afrotropic	DD	0	0	0	Neoromicia_helios	1	oldname match	Neoromicia_helios	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	44921	Neoromicia helios	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Neoromicia	helios	(Heller, 1912)	This is considered to be a subspecies of Pipistrellus nanus ;(currently Neoromicia nana ) by some authors (Simmons 2005).	20000000	Neoromicia helios	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Data Deficient in view of the absence of sufficient information on its extent of occurrence, natural history, threats and conservation status.	There is little information about the natural history of this bat, in part because it has often been confused with Neoromicia nana .	The threats to this species are not known.	Little information is available on the population abundance or size of this species.	Unknown	This East African species has been recorded from Djibouti, southern Somalia, southern Sudan, northeastern Uganda, Kenya and northern Tanzania. It may be more widely distributed (Peterson 1987, Simmons 2005).		Terrestrial	It is not known if the species is present in any protected areas. There is a need for studies into the distribution, abundance, natural history and possible threats to this species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Afronycteris		helios	Heller	1912	1	Smiths. Misc. Coll.	60(12): 3	Samburu Serotine	None.	Kenya, 30 mi S Mt. Marsabit, Merelle Water.	Kenya, Somalia, Djibouti, NE Uganda, extreme S Sudan, N Tanzania. Maybe more widespread (Peterson, 1987).	Not listed.	Data Deficient as Neoromicia helios 	Previously included in Neoromicia ; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Considered to be a subspecies of nanus by some authors, but differences in bacular morphology (Hill and Harrison, 1987), roosting andsocial behavior (O'Shea, 1980; Happold and Happold, 1996), habitat and pelage coloration, and the presence of a pair of glands on theinterfemoral membrane in helios (O'Shea, 1980) that are rarely found in nanus indicate that helios is a distinct species (M. Happold, pers. comm.).	Afronycteris helios	1005699	23	Heller's Serotine	Helios Pipistrelle Bat|Heller's Pipistrelle|Samburu Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Afronycteris	NA	helios	E. Heller	1912	1	Pipistrellus_helios	Heller, E. (1912). New Races of Insectivores, Bats, and Lemurs from British East Africa. Smithsonian Miscellaneous Collections, 60(12), 3	https://repository.si.edu/bitstream/handle/10088/23473/SMC_60_Heller_1912_12_1-13.pdf?sequence=1&isAllowed=y	USNM 181813		Merelle Water, 30 miles (= 48 km) south Mount Marsabit, Kenya.			helios (E. Heller, 1912)	moved from Neoromicia to the recently described Afronycteris, although further genetic studies will be needed to cement this arrangement	Monadjem, A., Demos, T. C., Dalton, D. L., Webala, P. W., Musila, S., Kerbis Peterhans, J. C., & Patterson, B. D. (2020). A revision of the pipistrelle-like bats (Mammali: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society, zlaa087.				South Sudan|Somalia|Uganda|Kenya|Tanzania|Djibouti?	Africa	Afrotropic	DD	0	0	0	Neoromicia_helios	1	oldname match	Neoromicia_helios	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Afronycteris_helios	1005699	23	Heller's Serotine	Helios Pipistrelle Bat|Heller's Pipistrelle|Samburu Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Afronycteris	NA	helios	E. Heller	1	Pipistrellus helios	Heller, E. 1912-11-04. New races of insectivores, bats, and lemurs from British East Africa. Smithsonian Miscellaneous Collections 60(12):1-13.	https://www.biodiversitylibrary.org/page/8910533	USNM:MAMM:181813	holotype	http://n2t.net/ark:/65665/3d66ec1da-1e7f-4644-b835-d213a3be056f	Merelle Water, 30 miles (= 48 km) south Mount Marsabit, Kenya.			moved from Neoromicia to the recently described Afronycteris, although further genetic studies will be needed to cement this arrangement	Monadjem, A., T. C. Demos, D. L. Dalton, P. W. Webala, S. Musila, J. C. K. Peterhans and B. D. Patterson (2021). A revision of pipistrelle-like bats (Mammalia: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society 191(4): 1114–1146.				South Sudan|Somalia|Uganda|Kenya|Tanzania|Djibouti?	Africa	Afrotropic	DD (as Neoromicia helios)	0	0	0	Neoromicia_helios	1	oldname match	Neoromicia_helios	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Afronycteris		helios	Heller	1912	1	Smiths. Misc. Coll.	60(12): 3	Samburu Serotine	None.	Kenya, 30 mi S Mt. Marsabit, Merelle Water.	Kenya, Somalia, Djibouti, NE Uganda, extreme S Sudan, N Tanzania. Maybe more widespread (Peterson, 1987).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/44921/22047381/' target='_blank'>Data Deficient as Neoromicia helios</a>	Previously included in Neoromicia; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Considered to be a subspecies of nanus by some authors, but differences in bacular morphology (Hill and Harrison, 1987), roosting andsocial behavior (O'Shea, 1980; Happold and Happold, 1996), habitat and pelage coloration, and the presence of a pair of glands on theinterfemoral membrane in helios (O'Shea, 1980) that are rarely found in nanus indicate that helios is a distinct species (M. Happold, pers. comm.).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Neoromicia helios; Neoromicia helios; Afronycteris helios; Afronycteris helios; Neoromicia helios; Afronycteris helios; helios; Vespeére soleil; Heller Zwergfledermaus; Neoromicia de Heller; Helios Pipistrelle Bat; Heller's Pipistrelle; Heller's Pipistrelle Bat; Samburu Pipistrelle; Samburu Pipistrelle Bat; Heller's Serotine; Helios Pipistrelle Bat; Heller's Pipistrelle; Samburu Pipistrelle; Samburu Pipistrelle; Samburu Serotine; Pipistrellus helios; A. helios