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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1094	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon	Nyctophilus microdon		[MSW3] See Flannery (1995a) and Bonaccorso (1998).; [HMW] Nyctophilus microdon Laurie & Hill, 1954 , “Welya, west of Hagen Range, central highlands, North-East New Guinea [= Papua New Guinea ], 7,000 ft [= 2134 m ].” Nyctophilus microdon is the only representative of the highly distinctive microdon species group. There is apparently an unnamed species of Nyctophilus in Papua New Guinea that has similar glans penis morphology to N. microdon according to H. E. Parnaby in 2009. Monotypic.; [batnames2022]  microdon species group. See Flannery (1995 a ) and Bonaccorso (1998).; [IUCN] This species is limited to the highlands of Papua New Guinea, so there is no suggestion of taxonomic issues apparent from the pattern of its distribution. However, a recent capture from Hela Province was genetically divergent from samples from Chimbu Province (Armstrong et al. 2019, K.N. Armstrong unpublished data). Further taxonomic work is required.; [batnames2023]  microdon species group. See Flannery (1995 a ) and Bonaccorso (1998).; [batnames2025_1.7] microdonspecies group. See Flannery (1995a) and Bonaccorso (1998).														microdon	This species is limited to the highlands of Papua New Guinea, so there is no suggestion of taxonomic issues apparent from the pattern of its distribution. However, a recent capture from Hela Province was genetically divergent from samples from Chimbu Province (Armstrong et al. 2019, K.N. Armstrong unpublished data). Further taxonomic work is required.			microdon 	microdon 			microdon Laurie & J. Edwards Hill, 1954		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Small-toothed long-eared	SE New Guinea bat	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nyctophilus microdon	Papua New Guinea, Western Highlands (?) Prov., Welya (W. of Hagen Range, 7000 ft. (2134 m)).	Laurie and Hill	1954	:78.	Distribution: Known only from the highlands of eastern New Guinea.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Small-toothed long-eared	SE New Guinea bat	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Laurie and Hill	1954	List of Land Mammals of New Guinea, Celebes, and adjacent Islands, p. 78.		EC New Guinea.	Papua New Guinea, Western Highlands (?) Prov., Welya (W of Hagen Range, 7,000 ft. (2,134 m)).		LAURIE & HILL	1954	Size medium (forearm length, 37-41 mm; condylobasal length, 13-15 mm). Posterior noseleaf fairly well developed. Band connecting ears relatively high. Teeth relatively small.	Distribution: Known only from the highlands of eastern New Guinea.	No subspecies.		131	species	N. microdon	LAURIE & HILL	1954	Nyctophilus	genus	Nyctophilus microdon				Size medium (forearm length, 37-41 mm; condylobasal length, 13-15 mm). Posterior noseleaf fairly well developed. Band connecting ears relatively high. Teeth relatively small.	No subspecies.		6. N. microdon LAURIE & HILL 1954.	6	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nyctophilini	Nyctophilus microdon	Nyctophilus		microdon	Laurie and Hill		1954		List of Land Mammals of New Guinea, Celebes, and adjacent Islands			78		Small-toothed Long-eared Bat	Papua New Guinea, Western Highlands (?) Prov., Welya (W of Hagen Range, 7,000 ft. (2,134 m)).	EC Papua New Guinea.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		See Flannery (1995a) and Bonaccorso (1998).	4C3D87E8FFCB6A74FA5B97891929B20F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	806	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFCB6A74FA5B97891929B20F.xml	Nyctophilus microdon	Vespertilionidae	Nyctophilus	microdon	Laurie & Hill	1954	Nyctophile a petites dents @fr | Kleinzahnige Langohrfledermaus @de | Nictofilade dientes pequenos @es | Small-toothed Bat @en	Nyctophilus microdon Laurie & Hill, 1954 , “Welya, west of Hagen Range, central highlands, North-East New Guinea [= Papua New Guinea ], 7,000 ft [= 2134 m ].” Nyctophilus microdon is the only representative of the highly distinctive microdon species group. There is apparently an unnamed species of Nyctophilus in Papua New Guinea that has similar glans penis morphology to N. microdon according to H. E. Parnaby in 2009. Monotypic.	Much of C highlands of Papua New Guinea in Eastern Highlands , Chimbu , Western Highlands , Southern Highlands , and Enga provinces.	Head—body 42-5-63 mm, tail 36-49 mm, ear 14-19-4 mm, hindfoot 7-10 mm, forearm 37-40-9 mm; weight 5-:8-9-5 g. The Small-toothed Long-eared Bat has very large ears and unique simple noseleaf consisting of two ridges, one further on muzzle and another immediately above nostrils, with vertical groove in middle and furred trough between them. Dorsal pelage is dark chocolate-brown (rich reddish brown according to Parnaby in 2009); venteris paler (hairs have pale gray-brown tips and chocolate-brown bases). Face, ears, and wing membranes are medium brown. Rostrum is short and blunt, with ridge across muzzle over nostrils that is moderately developed, consisting of two well-developed mounds joined in midline by obvious elastic membrane. Ears are very large and broad, with bluntly rounded tips, horizontal ribbing on inner surfaces, inward curved anterior edges, and smooth posterior edges (ears can fold back at top ofthick part of anterior edge); large and furred interauricular band crosses forehead between ears; tragus is enlarged, with roughly contoured lateral margin and truncated tip, being fairly straight on anterior margin. Glans penis is very small, with elongated urethral lappets and entirely unique distal structure. Baculum has moderately thin shaft in dorsal view; tip is bifurcated; and other details of baculum are not published but distinctive according to Parnaby in 2009. Skull is robust and broad; tympanic bullae are of moderate size; there is no sagittal crest, which is presentin all other species of Nyctophilus ; and M? and lower molars are completely unreduced.	Mid-montane and montane forests at elevations of 1600-3315 m.	The Small-toothed Long-eared Bat is probably an insect gleaner.	No information.	The Small-toothed Long-eared Bat roosts in trees and caves.	Small-toothed Long-eared Bats roost alone or in small groups.	Classified as Data Deficient on The IUCN Red List. The Smalltoothed Long-eared Bat appears to be more common than expected, and new specimens are turning up that extend its currently known distribution. Additional research is still needed, but a classification as Least Concern might be appropriate.	Bonaccorso (1998) | Bonaccorso, Hamilton & Parnaby (2008b) | Flannery (1995b) | Helgen & Opiang (2009) | Parnaby (2002a, 2009)	https://zenodo.org/record/6397958/files/figure.png	90. Small-toothed Long-eared Bat Nyctophilus microdon French: Nyctophile a petites dents / German: Kleinzahnige Langohrfledermaus / Spanish: Nictofila de dientes pequenos Other common names: Small-toothed Bat Taxonomy. Nyctophilus microdon Laurie & Hill, 1954 , “Welya, west of Hagen Range, central highlands, North-East New Guinea [= Papua New Guinea ], 7,000 ft [= 2134 m ].” Nyctophilus microdon is the only representative of the highly distinctive microdon species group. There is apparently an unnamed species of Nyctophilus in Papua New Guinea that has similar glans penis morphology to N. microdon according to H. E. Parnaby in 2009. Monotypic. Distribution. Much of C highlands of Papua New Guinea in Eastern Highlands , Chimbu , Western Highlands , Southern Highlands , and Enga provinces. Descriptive notes. Head—body 42-5-63 mm, tail 36-49 mm, ear 14-19-4 mm, hindfoot 7-10 mm, forearm 37-40-9 mm; weight 5-:8-9-5 g. The Small-toothed Long-eared Bat has very large ears and unique simple noseleaf consisting of two ridges, one further on muzzle and another immediately above nostrils, with vertical groove in middle and furred trough between them. Dorsal pelage is dark chocolate-brown (rich reddish brown according to Parnaby in 2009); venteris paler (hairs have pale gray-brown tips and chocolate-brown bases). Face, ears, and wing membranes are medium brown. Rostrum is short and blunt, with ridge across muzzle over nostrils that is moderately developed, consisting of two well-developed mounds joined in midline by obvious elastic membrane. Ears are very large and broad, with bluntly rounded tips, horizontal ribbing on inner surfaces, inward curved anterior edges, and smooth posterior edges (ears can fold back at top ofthick part of anterior edge); large and furred interauricular band crosses forehead between ears; tragus is enlarged, with roughly contoured lateral margin and truncated tip, being fairly straight on anterior margin. Glans penis is very small, with elongated urethral lappets and entirely unique distal structure. Baculum has moderately thin shaft in dorsal view; tip is bifurcated; and other details of baculum are not published but distinctive according to Parnaby in 2009. Skull is robust and broad; tympanic bullae are of moderate size; there is no sagittal crest, which is presentin all other species of Nyctophilus ; and M? and lower molars are completely unreduced. Habitat. Mid-montane and montane forests at elevations of 1600-3315 m. Food and Feeding. The Small-toothed Long-eared Bat is probably an insect gleaner. Breeding. No information. Activity patterns. The Small-toothed Long-eared Bat roosts in trees and caves. Movements, Home range and Social organization. Small-toothed Long-eared Bats roost alone or in small groups. Status and Conservation. Classified as Data Deficient on The IUCN Red List. The Smalltoothed Long-eared Bat appears to be more common than expected, and new specimens are turning up that extend its currently known distribution. Additional research is still needed, but a classification as Least Concern might be appropriate. Bibliography. Bonaccorso (1998), Bonaccorso, Hamilton & Parnaby (2008b), Flannery (1995b), Helgen & Opiang (2009), Parnaby (2002a, 2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Nyctophilus microdon	Nyctophilus		microdon	Laurie & Hill	1954	0	List of Land Mammals of New Guinea, Celebes, and adjacent Islands	p. 78	Small-toothed Long-eared Bat	None.	Papua New Guinea, Western Highlands (?) Prov., Welya (W of Hagen Range, 7,000 ft. (2,134 m)).	EC Papua New Guinea.	Not listed.	Least Concern	 microdon species group. See Flannery (1995 a ) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctophilus microdon	23	Small-toothed Long-eared Bat	Small-toothed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Nyctophilus	NA	microdon	Laurie & J. Edwards Hill	1954	0						"Welya, west of Hagen Range, central highlands, North-East New Guinea [= Papua New Guinea], 7,000 ft [= 2134 m]."			microdon Laurie & J. Edwards Hill, 1954	NA	NA	Papua New Guinea	Oceania	Australasia/Oceania	DD	0	0	0	Nyctophilus_microdon	0	sciname match	Nyctophilus_microdon	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	15007	Nyctophilus microdon	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Nyctophilus	microdon	Laurie &; Hill, 1954	This species is limited to the highlands of Papua New Guinea, so there is no suggestion of taxonomic issues apparent from the pattern of its distribution. However, a recent capture from Hela Province was genetically divergent from samples from Chimbu Province (Armstrong et al. 2019, K.N. Armstrong unpublished data). Further taxonomic work is required.	20000000	Nyctophilus microdon	Least Concern		2021	2020-12-10 00:00:00 UTC	3.1	English	This species is assessed as Least Concern because while there is a lack of information about its population size and extent of occurrence, its habitat is mostly intact and no major threats that could cause widespread or imminent decline have been identified. It might be distributed more widely at mid-montane elevations on the island of New Guinea.	This species roosts in trees and caves, as solitary animals or in small groups. It gleans insects from foliage within the interior of its mid-montane forest habitat (Bonaccorso 1998).	There appear to be no major threats to this species that could cause widespread or imminent decline. Its higher elevation habitats are generally intact.	There is no information on population size, but its habitat is mostly intact. It is only known from 15 specimens in three (3) provinces of Papua New Guinea (records in the Australian National Wildlife Collection, Australian Museum, South Australian Museum, Armstrong et al. 2019). Its population is inferred to be stable.	Unknown	This species is endemic to Papua New Guinea where it has been recorded from Sandaun Province, Hela Province, and Chimbu Province, between 1,900 and 2,800 m asl (Bonaccorso 1998, Armstrong et al. 2019). Its projected range in mid-montane elevations and extends from Sundan Province in the west to Chimbu Province in the east. Its extent of occurrence and area of occupancy could be larger than what has been documented to date.	The species is not known to be hunted, used, or traded.	Terrestrial	The species is presumed to be present in protected areas within its elevational range. Broadscale protection of forests will help maintain population size in this species. Future surveys should target it with harp traps. Additional research is needed to determine its range, population status and trends, ecology, and threats.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Nyctophilus		microdon	Laurie & Hill	1954	0	List of Land Mammals of New Guinea, Celebes, and adjacent Islands	p. 78	Small-toothed Long-eared Bat	None.	Papua New Guinea, Western Highlands (?) Prov., Welya (W of Hagen Range, 7,000 ft. (2,134 m)).	EC Papua New Guinea.	Not listed.	Least Concern	 microdon species group. See Flannery (1995 a ) and Bonaccorso (1998).	Nyctophilus microdon	1005762	23	Small-toothed Long-eared Bat	Small-toothed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Nyctophilus	NA	microdon	Laurie & J. Edwards Hill	1954	0						"Welya, west of Hagen Range, central highlands, North-East New Guinea [= Papua New Guinea], 7,000 ft [= 2134 m]."			microdon Laurie & J. Edwards Hill, 1954	NA	NA				Papua New Guinea	Oceania	Australasia/Oceania	DD	0	0	0	Nyctophilus_microdon	0	sciname match	Nyctophilus_microdon	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctophilus_microdon	1005762	23	Small-toothed Long-eared Bat	Small-toothed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Nyctophilus	NA	microdon	Laurie & J. Edwards Hill	0	Nyctophilus microdon	Laurie, E.M.O. and Hill, J.E. 1954-06-30. List of land mammals of New Guinea, Celebes and adjacent islands 1758–1952. British Museum, London, 175 pp.	https://www.biodiversitylibrary.org/page/49635618	BMNH:Mamm:1953.218	holotype	https://data.nhm.ac.uk/object/4dd9efa7-363f-4b2f-a945-0744f1cf5398	"Welya, west of Hagen Range, central highlands, North-East New Guinea [= Papua New Guinea], 7,000 ft [= 2134 m]."			NA	NA				Papua New Guinea	Oceania (Continent)	Australasia	LC	0	0	0	Nyctophilus_microdon	0	sciname match	Nyctophilus_microdon	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Nyctophilus		microdon	Laurie & Hill	1954	0	List of Land Mammals of New Guinea, Celebes, and adjacent Islands	p. 78	Small-toothed Long-eared Bat	None.	Papua New Guinea, Western Highlands (?) Prov., Welya (W of Hagen Range, 7,000 ft. (2,134 m)).	EC Papua New Guinea.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/15007/22009794/' target='_blank'>Least Concern</a>	microdonspecies group. See Flannery (1995a) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctophilus microdon; Nyctophilus microdon; Nyctophilus microdon; Nyctophilus microdon; Nyctophilus microdon; Nyctophilus microdon; microdon; Nyctophile a petites dents; Kleinzahnige Langohrfledermaus; Nictofilade dientes pequenos; Small-toothed Bat; Small-toothed Long-eared Bat; Small-toothed Bat; Small-toothed Long-eared Bat; Small-toothed Long-eared Bat; N. microdon