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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L110	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Carollia benkeithi	Carollia benkeithi	Carollia benkeithi	Carollia benkeithi	Carollia benkeithi	Carollia benkeithi	Carollia benkeithi	Carollia benkeithi	Carollia benkeithi		[HMW] Carollia benkeithi Solari & R. J. Baker, 2006 , “ 2 km S of Tingo Maria, Province of Leoncio Prado, Department of Huanuco , Peru , at approximately 9° 18’ S , 75º 59' W .” Carollia benkeithi was split from C. castanea , which was shown to be a species complex based on molecular and morphological analyses. Monotypic.; [batnames2022] Brazilian record from McKellan and Koopman (2008), and see Lemos et al. (2020). See discussion in Lemos et al. (2020) for morphological variation and characters useful for distinguishing among perspicillata , brevicauuda , and benkeithi in Brazil.; [MDD2022] recently described; [IUCN] This species was <span>previously included under Carollia castanea , it is now considered a distinct species (Solari and Baker 2006).; [batnames2023] Brazilian record from McKellan and Koopman (2008), and see Lemos et al. (2020). See discussion in Lemos et al. (2020) for morphological variation and characters useful for distinguishing among perspicillata , brevicauuda , and benkeithi in Brazil.; [MDD2023] recently described; [MDD2025_2.0] recently described; [batnames2025_1.7] Brazilian record from McKellan and Koopman (2008), and see Lemos et al. (2020). See discussion in Lemos et al. (2020) for morphological variation and characters useful for distinguishing among perspicillata, brevicauuda, and benkeithi in Brazil.; [MDD2025_2.2] recently described														benkeithi	This species was <span>previously included under Carollia castanea , it is now considered a distinct species (Solari and Baker 2006).			benkeithi	benkeithi			benkeithi Solari & R. J. Baker, 2006						N/A																																								NA																											03A687BCFF82FF8213B4FE4FF8A5F3C5	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	535	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF82FF8213B4FE4FF8A5F3C5.xml	Carollia benkeithi	Phyllostomidae	Carollia	benkeithi	Solari & R. J. Baker	2006	Carollia de Ben Keith @fr | Keith-Kurzschwanzblattnase @de | Carolia de Ben Keith @es | Benkeith's Short-tailed Bat @en	Carollia benkeithi Solari & R. J. Baker, 2006 , “ 2 km S of Tingo Maria, Province of Leoncio Prado, Department of Huanuco , Peru , at approximately 9° 18’ S , 75º 59' W .” Carollia benkeithi was split from C. castanea , which was shown to be a species complex based on molecular and morphological analyses. Monotypic.	Lowland forests of W Brazil , E Peru , and N Bolivia , S of the Amazon River. Because of abundance of individuals in the C. castanea complex, hampering precise species identification in the field and in collections, a detailed assessment ofits distribution is still needed.	Head-body 52-58 mm, tail 5-14 mm, ear 11-20 mm, hindfoot 8-14 mm, forearm 33-7-37-2 mm; weight 11-16 g. Ben Keith’s Short-tailed Batis small. Dorsal hairs lack sharply defined banding; they have broad buffy chestnut band at bases, followed by brown yellowish band, and then narrow chestnut to dull gray-brown tips. Ventral fur has short bicolored, brown-tipped hairs. Forearms are short and naked, with tuft of hairs at bases of thumbs. Uropatagium is wide, with shallow distal notch. Tail is short, almost one-third the length of uropatagium. Wings are attached to ankles. Muzzle is conspicuous but short; lower lip has central papillae surrounded by smaller warts in a V-shape. Noseleafis short, with elongated tip. Ears are moderately large and triangular, and tips are pointed. Rostrum is short, braincase is globular, and zygomatic arches are incomplete. Anterior upper tooth row (to P?) is almost parallel, but posterior one-half is more divergent. Upper and lower molars are not broad. Cusps of M| are reduced and inconspicuous in side view. Dental formula for all species of Carollinis 12/2, C 1/1, P 2/2 M 3/3 (22) = 32. Chromosomal complement has 2n = 22 and FN = 38, with a single pair of mediumsmall acrocentric autosomes.	Typically tropical evergreen forests, but also secondary forests and much less commonly clearings or plantations, at elevations mostly below 1000 m . Locally, Ben Keith’s Short-tailed Bat can be the most abundant species of bats in some habitats, but it can be variably abundant at a regional scale. Some specimens from eastern Peru were more commonly mist-netted in brushy vegetation containing Piper sp. ( Piperaceae ) along edges of a small airstrip in the middle ofa village.	Similar to other species of Carollia , fruits are the primary food of Ben Keith's Short-tailed Bat. It depends on Piper spp. in dry seasons, butit eats other fruit species during wet seasons, with preferences for fruits produced by understory plants such as Solanum ( Solanaceae ) and Vismia ( Hypericaceae ). Insects are occasionally eaten. A study in south-eastern Peru found insect remains in ¢.20% of fecal samples, and more than 65% of feces included only fruit (including seeds of 48 plant species), and only one (2%) included pollen. Species of Carollia reacted to potential food (artificial fruit and real ripe Piper ) only when it had the typical odor of ripe fruit; they used odor as a primary clue to find ripe fruit. Shape and position offruits play important roles in foraging behavior. Echolocation is used for orientation in space and obstacle avoidance; echolocation calls are important at close range for final location of a fruit so it can be taken in flight.	Female Ben Keith's Short-tailed Bats have been reproductively inactive in May-July and pregnant in February-March, September, and November, which is similar to what is known for other species of Carollia . No specific data exist for Ben Keith's Short-tailed Bat, but the closely related Chestnut Short-tailed Bat ( C. castanea ) in Peru and Bolivia seems to be predominantly polyestrous.	Ben Keith's Short-tailed Bat is active during the first hours after sunset. Field data suggest that it is more active in the first one-half of the night, with only occasional captures after midnight. Scarcity of roost records probably indicates the difficulty of properly identifying it inside a roost, or it might use roosts opportunistically. Known roosts include shallow caves, rock crevices, tree holes, and human structures such as tunnels and bridges.	Ben Keith’s Short-tailed Bat seems to have small home ranges. It will share hollow trees with congeners such as the Silky Short-tailed Bat (C. brevicaudum), Seba’s Short-tailed Bat ( C. perspicillata ), and the Proboscis Bat ( Rhynchonycteris naso ).	Not assessed on The IUCN Red List. Ben Keith's Short-tailed Bat can be fairly abundant locally, and its wide distribution might suggestit is not currently threatened.	Ascorra, Solari & Wilson (1996) | Ascorra, Wilson & Romo (1991) | Bonaccorso et al. (2007) | Fleming (1991) | Gorchov et al. (1995) | Hoffmann & Baker (2003) | McLellan & Koopman (2008) | Pine (1972) | Solari & Baker (2006) | Tuttle (1970) | Wilson (1979)	https://zenodo.org/record/6458817/files/figure.png	103. Ben Keith's Short-tailed Bat Carollia benkeithi French: Carollia de Ben Keith / German: Keith-Kurzschwanzblattnase / Spanish: Carolia de Ben Keith Other common names: Benkeith's Short-tailed Bat Taxonomy. Carollia benkeithi Solari & R. J. Baker, 2006 , “ 2 km S of Tingo Maria, Province of Leoncio Prado, Department of Huanuco , Peru , at approximately 9° 18’ S , 75º 59' W .” Carollia benkeithi was split from C. castanea , which was shown to be a species complex based on molecular and morphological analyses. Monotypic. Distribution. Lowland forests of W Brazil , E Peru , and N Bolivia , S of the Amazon River. Because of abundance of individuals in the C. castanea complex, hampering precise species identification in the field and in collections, a detailed assessment ofits distribution is still needed. Descriptive notes. Head-body 52-58 mm, tail 5-14 mm, ear 11-20 mm, hindfoot 8-14 mm, forearm 33-7-37-2 mm; weight 11-16 g. Ben Keith’s Short-tailed Batis small. Dorsal hairs lack sharply defined banding; they have broad buffy chestnut band at bases, followed by brown yellowish band, and then narrow chestnut to dull gray-brown tips. Ventral fur has short bicolored, brown-tipped hairs. Forearms are short and naked, with tuft of hairs at bases of thumbs. Uropatagium is wide, with shallow distal notch. Tail is short, almost one-third the length of uropatagium. Wings are attached to ankles. Muzzle is conspicuous but short; lower lip has central papillae surrounded by smaller warts in a V-shape. Noseleafis short, with elongated tip. Ears are moderately large and triangular, and tips are pointed. Rostrum is short, braincase is globular, and zygomatic arches are incomplete. Anterior upper tooth row (to P?) is almost parallel, but posterior one-half is more divergent. Upper and lower molars are not broad. Cusps of M| are reduced and inconspicuous in side view. Dental formula for all species of Carollinis 12/2, C 1/1, P 2/2 M 3/3 (22) = 32. Chromosomal complement has 2n = 22 and FN = 38, with a single pair of mediumsmall acrocentric autosomes. Habitat. Typically tropical evergreen forests, but also secondary forests and much less commonly clearings or plantations, at elevations mostly below 1000 m . Locally, Ben Keith’s Short-tailed Bat can be the most abundant species of bats in some habitats, but it can be variably abundant at a regional scale. Some specimens from eastern Peru were more commonly mist-netted in brushy vegetation containing Piper sp. ( Piperaceae ) along edges of a small airstrip in the middle ofa village. Food and Feeding. Similar to other species of Carollia , fruits are the primary food of Ben Keith's Short-tailed Bat. It depends on Piper spp. in dry seasons, butit eats other fruit species during wet seasons, with preferences for fruits produced by understory plants such as Solanum ( Solanaceae ) and Vismia ( Hypericaceae ). Insects are occasionally eaten. A study in south-eastern Peru found insect remains in ¢.20% of fecal samples, and more than 65% of feces included only fruit (including seeds of 48 plant species), and only one (2%) included pollen. Species of Carollia reacted to potential food (artificial fruit and real ripe Piper ) only when it had the typical odor of ripe fruit; they used odor as a primary clue to find ripe fruit. Shape and position offruits play important roles in foraging behavior. Echolocation is used for orientation in space and obstacle avoidance; echolocation calls are important at close range for final location of a fruit so it can be taken in flight. Breeding. Female Ben Keith's Short-tailed Bats have been reproductively inactive in May-July and pregnant in February-March, September, and November, which is similar to what is known for other species of Carollia . No specific data exist for Ben Keith's Short-tailed Bat, but the closely related Chestnut Short-tailed Bat ( C. castanea ) in Peru and Bolivia seems to be predominantly polyestrous. Activity patterns. Ben Keith's Short-tailed Bat is active during the first hours after sunset. Field data suggest that it is more active in the first one-half of the night, with only occasional captures after midnight. Scarcity of roost records probably indicates the difficulty of properly identifying it inside a roost, or it might use roosts opportunistically. Known roosts include shallow caves, rock crevices, tree holes, and human structures such as tunnels and bridges. Movements, Home range and Social organization. Ben Keith’s Short-tailed Bat seems to have small home ranges. It will share hollow trees with congeners such as the Silky Short-tailed Bat (C. brevicaudum), Seba’s Short-tailed Bat ( C. perspicillata ), and the Proboscis Bat ( Rhynchonycteris naso ). Status and Conservation. Not assessed on The IUCN Red List. Ben Keith's Short-tailed Bat can be fairly abundant locally, and its wide distribution might suggestit is not currently threatened. Bibliography. Ascorra, Solari & Wilson (1996), Ascorra, Wilson & Romo (1991), Bonaccorso et al. (2007), Fleming (1991), Gorchov et al. (1995), Hoffmann & Baker (2003), McLellan & Koopman (2008), Pine (1972), Solari & Baker (2006), Tuttle (1970), Wilson (1979).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Carollia benkeithi	Carollia		benkeithi	Solari & Baker	2006	0	Occas. Pap. Mus. Texas Tech Univ.	######	Ben Keith's Short-tailed Bat	None.		Peru, Brazil	Not listed.	Least Concern	Brazilian record from McKellan and Koopman (2008), and see Lemos et al. (2020). See discussion in Lemos et al. (2020) for morphological variation and characters useful for distinguishing among perspicillata , brevicauuda , and benkeithi in Brazil.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Carollia benkeithi	23	Ben Keith's Short-tailed Bat	Benkeith's Short-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	CAROLLIINAE	NA	Carollia	NA	benkeithi	Solari & R. J. Baker	2006	0	Carollia_benkeithi	Solari, S., & Baker, R. J. (2006). Mitochondrial DNA Sequence, Karyotypic, and Morphological Variation in the Carollia castanea Species Complex (Chiroptera, Phyllostomidae) with Description of a New Species. Occasional Papers, Museum of Texas Tech University, 254, 5.	https://www.biodiversitylibrary.org/page/57663646#page/6/mode/1up	TTU 46817		"2 km S of Tingo María, Province of Leoncio Prado, Department of Huánuco, Peru, at approximately 9° 18' S, 75° 59' W."	-9.3	-75.98	benkeithi Solari & R. J. Baker, 2006	recently described	Solari, S. and Baker, R.J. 2006. Mitochondrial DNA Sequence, Karyotypic, and Morphological Variation in the Carollia castanea Species Complex (Chiroptera, Phyllostomidae) with Description of a New Species. Occasional Papers of the Museum of Texas Tech University, 254, 1-16.	Brazil|Peru|Bolivia	South America	Neotropic	LC	0	0	0	Carollia_benkeithi	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Carollia benkeithi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Carollia	benkeithi	Solari &; Baker, 2006	This species was <span>previously included under Carollia castanea , it is now considered a distinct species (Solari and Baker 2006).	90000000	Carollia benkeithi	Least Concern		2019	2016-07-11 00:00:00 UTC	3.1	English	This species is listed as Least Concern because of ;its wide geographic distribution, including large sections of continuous native forests, and ;natural abundance at local scale. There is no major threats to these habitats or the species itself.	The natural history of Carollia benkeithi most likely closely resembles that of C. castanea . Based on records from several sources, Solari and Baker (2006) stated that this species was typically found in tropical evergreen forests at lower elevations, mostly below 1,000 m. ;Specimens from Balta in eastern Peru were most commonly mistnetted in the brushy vegetation containing ;Piper ; sp. along the edges ;of the small air strip in the middle of the village (Gardner pers. comm).	Threats to this species are not known.	Species of the genus Carollia are often common to abundant, and C. benkeithi is not an exception (Solari and Baker 2006). However, this species is not as abundant as C. brevicauda or C. perspicillata , and is typically associated to humid lowland forests. The species appears to be uncommon in the central Amazon basin of Brazil.	Stable	Carollia benkeithi is known from eastern Peru, northeastern Bolivia and western Brazil (Solari and Baker 2006).		Terrestrial	Most of the distribution of this species, in southeastern Peru and northwestern Bolivia, include large national parks and reserves that protect extensive areas of lowland and premontane forests. Populations of this species should be well protected within this area.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Carollia		benkeithi	Solari & Baker	2006	0	Occas. Pap. Mus. Texas Tech Univ.	254:05:00	Ben Keith's Short-tailed Bat	None.		Peru, Brazil	Not listed.	Least Concern	Brazilian record from McKellan and Koopman (2008), and see Lemos et al. (2020). See discussion in Lemos et al. (2020) for morphological variation and characters useful for distinguishing among perspicillata , brevicauuda , and benkeithi in Brazil.	Carollia benkeithi	1004872	23	Ben Keith's Short-tailed Bat	Benkeith's Short-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	CAROLLIINAE	NA	Carollia	NA	benkeithi	Solari & R. J. Baker	2006	0	Carollia_benkeithi	Solari, S., & Baker, R. J. (2006). Mitochondrial DNA Sequence, Karyotypic, and Morphological Variation in the Carollia castanea Species Complex (Chiroptera, Phyllostomidae) with Description of a New Species. Occasional Papers, Museum of Texas Tech University, 254, 5.	https://www.biodiversitylibrary.org/page/57663646#page/6/mode/1up	TTU 46817		"2 km S of Tingo María, Province of Leoncio Prado, Department of Huánuco, Peru, at approximately 9° 18' S, 75° 59' W."	-9.3	-75.9833	benkeithi Solari & R. J. Baker, 2006	recently described	Solari, S. and Baker, R.J. 2006. Mitochondrial DNA Sequence, Karyotypic, and Morphological Variation in the Carollia castanea Species Complex (Chiroptera, Phyllostomidae) with Description of a New Species. Occasional Papers of the Museum of Texas Tech University, 254, 1-16.				Brazil|Peru|Bolivia	South America	Neotropic	LC	0	0	0	Carollia_benkeithi	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Carollia_benkeithi	1004872	23	Ben Keith's Short-tailed Bat	Benkeith's Short-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Carolliinae	NA	Carollia	NA	benkeithi	Solari & R. J. Baker	0	Carollia benkeithi	Solari, S. and Baker, R.J. 2006. Mitochondrial DNA sequence, karyotypic, and morphological variation in the _Carollia castanea_ species complex (Chiroptera: Phyllostomidae), with description of a new species. Occasional Papers, Museum of Texas Tech University 254:1-16.	https://www.biodiversitylibrary.org/page/57663650	TTU-M46187	holotype		"2 km S of Tingo María, Province of Leoncio Prado, Department of Huánuco, Peru, at approximately 9° 18' S, 75° 59' W."	-9.3	-75.9833	recently described	Solari, S. and Baker, R.J. 2006. Mitochondrial DNA Sequence, Karyotypic, and Morphological Variation in the Carollia castanea Species Complex (Chiroptera, Phyllostomidae) with Description of a New Species. Occasional Papers of the Museum of Texas Tech University, 254, 1-16.				Brazil|Peru|Bolivia	South America	Neotropic	LC	0	0	0	Carollia_benkeithi	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Carollia		benkeithi	Solari & Baker	2006	0	Occas. Pap. Mus. Texas Tech Univ.	254:05:00	Ben Keith's Short-tailed Bat	None.		Peru, Brazil	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/88110352/88110355/' target='_blank'>Least Concern</a>	Brazilian record from McKellan and Koopman (2008), and see Lemos et al. (2020). See discussion in Lemos et al. (2020) for morphological variation and characters useful for distinguishing among perspicillata, brevicauuda, and benkeithi in Brazil.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Carollia benkeithi; Carollia benkeithi; Carollia benkeithi; Carollia benkeithi; Carollia benkeithi; benkeithi; Carollia de Ben Keith; Keith-Kurzschwanzblattnase; Carolia de Ben Keith; Benkeith's Short-tailed Bat; Ben Keith's Short-tailed Bat; Benkeith's Short-tailed Bat; Ben Keith's Short-tailed Bat; C. benkeithi