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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1084	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis	Nyctophilus arnhemensis		[HMW] Nyctophilus arnhemensis D. H. Johnson, 1959 , “Rocky Bay, south of Yirkala, Cape Arnhem Peninsula, Northern Territory , Australia (lat. 12° 16’ S , long. 136° 47' E ).” Nyctophilus arnhemensis is in the bifax species group. It is extremely similar to N. bfax in most morphological characters (although it is smaller) and could potentially be a subspecies of N. bifax , although some morphological data support recognition of the two species as separate. More studies with genetic data are needed to resolve its taxonomy. Monotypic.; [batnames2022]  bifax species group.; [IUCN] The genus Lamingtonia is a synonym of Nyctophilus .; [batnames2023]  bifax species group.; [batnames2025_1.7] bifaxspecies group.														arnhemensis	The genus Lamingtonia is a synonym of Nyctophilus .			arnhemensis 	arnhemensis 			arnhemensis D. H. Johnson, 1959		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Arnhem Land long-eared bat	N Northern Terr., Australia; ref. 4.112	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nyctophilus arnhemensis	Australia, Northern Territory, Cape Arnhem Peninsula, S. of Yirkala, Rocky Bay. (12° 13' S. and 136° 47' E.).	Johnson	1959	Proc. Biol. Soc. Wash., 72:184.	Distribution: Confined to extreme northern Australia from northeastern Western Australia to northwestern Queensland.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Arnhem Land long-eared	NW Australia bat	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Johnson	1959	Proc. Biol. Soc. Washington, 72:184.		N Australia.	Australia, Northern Territory, Cape Arnhem Peninsula, S of Yirkala, Rocky Bay. (12°13'S and 136°47'E).		JOHNSON	1959	Size fairly small (forearm length, 34-41 mm; condylobasal length, 13-15 mm). Noseleaves poorly developed. Band connecting ears (which are relatively long) relatively low. Teeth relatively large. Auditory bullae relatively large.	Distribution: Confined to extreme northern Australia from northeastern Western Australia to northwestern Queensland.	No subspecies.		131	species	N. arnhemensis	JOHNSON	1959	Nyctophilus	genus	Nyctophilus arnhemensis				Size fairly small (forearm length, 34-41 mm; condylobasal length, 13-15 mm). Noseleaves poorly developed. Band connecting ears (which are relatively long) relatively low. Teeth relatively large. Auditory bullae relatively large.	No subspecies.		3. N. arnhemensis JOHNSON 1959.	3	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nyctophilini	Nyctophilus arnhemensis	Nyctophilus		arnhemensis	Johnson		1959		Proc. Biol. Soc. Wash.	72		184		Northern Long-eared Bat	Australia, Northern Territory, Cape Arnhem Peninsula, S of Yirkala, Rocky Bay. (12°13'S, 36°47'E).	N Australia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).			4C3D87E8FFD46A74FA849F3F1FE3BD91	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	805	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFD46A74FA849F3F1FE3BD91.xml	Nyctophilus arnhemensis	Vespertilionidae	Nyctophilus	arnhemensis	Johnson	1959	Nyctophile d Arnhem @fr | Arnhem-Langohrfledermaus @de | Nictofila de Arnhem @es	Nyctophilus arnhemensis D. H. Johnson, 1959 , “Rocky Bay, south of Yirkala, Cape Arnhem Peninsula, Northern Territory , Australia (lat. 12° 16’ S , long. 136° 47' E ).” Nyctophilus arnhemensis is in the bifax species group. It is extremely similar to N. bfax in most morphological characters (although it is smaller) and could potentially be a subspecies of N. bifax , although some morphological data support recognition of the two species as separate. More studies with genetic data are needed to resolve its taxonomy. Monotypic.	N coast of Western Australia and Kimberley region (including Bonaparte Archipelago), N Northern Territory (including Tiwi Is, Groote Eylandt, and Sir Edward Pellew Group), and NW Queensland ; a specimen was recently recorded from NE Queensland in Cape York, but it needs to be verified.	Head-body 40-59 mm, tail 35-641 mm, ear 16-21-5 mm, forearm 33-2-39-9 mm; weight 5-2-9-6 g. The Northern Long-eared Bat has very large ears and unique simple noseleaf consisting of two ridges, one further on muzzle and another immediately above nostrils, with vertical groove in middle and furred trough between them. Dorsal pelage is generally light russet-brown butis occasionally rich dark brown (hairs have dark bases), which shades on side toslightly lighter venter thatis not distinctly different. Face, ears, and wing membranes are medium brown. Rostrum is short and blunt, with ridge across muzzle over nostrils that is moderately developed, consisting of two rounded mounds separated medially by thin vertical groove. Ears are very large and broad, with bluntly rounded tips, horizontal ribbing on inner surfaces, inward curved anterior edges, and smooth posterior edges (ears can fold back at top of thick part of anterior edge); large and furred interauricular band crosses forehead between ears; tragus is small and bluntly rounded attip, being convex on anterior margin. Glans penis is a square-ended cylinder, with flat circular urethral opening on underside near tip. Baculum has moderately thin shaft in dorsal view; tip is deeply bifurcated, and base is strongly bifurcated; in lateral view, baculum is curved downward at base but shaft is straight to pointed tip. Skull is robust; tympanic bullae are small; skull is similar to that of the Eastern Long-eared Bat ( N. bifax ) in general morphology; and M? and lower molars are only moderately reduced.	Typically wet habitats such as mangrove forests, monsoon forests, rainforest patches, open savanna woodlands, tall open forests, and Melaleuca ( Myrtaceae ) and Pandanus ( Pandanaceae ) lined streams, waterholes, and swamps. Northern Longeared Bats are generally found in areas that receive more than 500 mm of rain/year.	Northern Long-eared Bats primarily forage by gleaning off the ground and foliage and might also catch prey aerially. They are relatively slow fliers ( 5 km /h most of the time) but can reach top speeds of 16 km /h. Foraging usually occurs close to and among dense vegetation, and they are highly maneuverable in cluttered areas. Diet in the Top End region consisted largely of termites, water beetles, cockroaches, crickets, and true bugs and occasionally moths and spiders. They reportedly feed on geckos and gecko eggs, but this requires additional verification.	Male Northern Long-eared Bats with scrotal testes have been reported in May-November and pregnant females in September—November. Females apparently give birth to twins from late October to February.	Northern Long-eared Bats roost during the day under bark of large paperbark trees ( Melaleuca ), within foliage, and among Pandanus leaves. They have also been reported roosting in houses. They spend the night foraging. Call shape 1s very steep FM sweep, with peak frequencies of 50-52 kHz (mean 50-8 kHz).	The Northern Long-eared Bat probably roosts in small groups.	Classified as Least Concern on The IUCN Red List. The Northern Long-eared Bat is widespread and does not seem to face any major threats.	Bullen & McKenzie (2002a) | Churchill (2008) | Churchill et al. (1984) | Flannery (1995b) | McKenzie & Churchill (2008) | McKenzie, Fontanini et al. (1995) | McKenzie, Reardon & Parnaby (2008) | Milne (2002) | Milne et al. (2016) | Parnaby (2009)	https://zenodo.org/record/6397954/files/figure.png	88. Northern Long-eared Bat Nyctophilus arnhemensis French: Nyctophile dArnhem / German: Arnhem-Langohrfledermaus / Spanish: Nictofila de Arnhem Taxonomy. Nyctophilus arnhemensis D. H. Johnson, 1959 , “Rocky Bay, south of Yirkala, Cape Arnhem Peninsula, Northern Territory , Australia (lat. 12° 16’ S , long. 136° 47' E ).” Nyctophilus arnhemensis is in the bifax species group. It is extremely similar to N. bfax in most morphological characters (although it is smaller) and could potentially be a subspecies of N. bifax , although some morphological data support recognition of the two species as separate. More studies with genetic data are needed to resolve its taxonomy. Monotypic. Distribution. N coast of Western Australia and Kimberley region (including Bonaparte Archipelago), N Northern Territory (including Tiwi Is, Groote Eylandt, and Sir Edward Pellew Group), and NW Queensland ; a specimen was recently recorded from NE Queensland in Cape York, but it needs to be verified. Descriptive notes. Head-body 40-59 mm, tail 35-641 mm, ear 16-21-5 mm, forearm 33-2-39-9 mm; weight 5-2-9-6 g. The Northern Long-eared Bat has very large ears and unique simple noseleaf consisting of two ridges, one further on muzzle and another immediately above nostrils, with vertical groove in middle and furred trough between them. Dorsal pelage is generally light russet-brown butis occasionally rich dark brown (hairs have dark bases), which shades on side toslightly lighter venter thatis not distinctly different. Face, ears, and wing membranes are medium brown. Rostrum is short and blunt, with ridge across muzzle over nostrils that is moderately developed, consisting of two rounded mounds separated medially by thin vertical groove. Ears are very large and broad, with bluntly rounded tips, horizontal ribbing on inner surfaces, inward curved anterior edges, and smooth posterior edges (ears can fold back at top of thick part of anterior edge); large and furred interauricular band crosses forehead between ears; tragus is small and bluntly rounded attip, being convex on anterior margin. Glans penis is a square-ended cylinder, with flat circular urethral opening on underside near tip. Baculum has moderately thin shaft in dorsal view; tip is deeply bifurcated, and base is strongly bifurcated; in lateral view, baculum is curved downward at base but shaft is straight to pointed tip. Skull is robust; tympanic bullae are small; skull is similar to that of the Eastern Long-eared Bat ( N. bifax ) in general morphology; and M? and lower molars are only moderately reduced. Habitat. Typically wet habitats such as mangrove forests, monsoon forests, rainforest patches, open savanna woodlands, tall open forests, and Melaleuca ( Myrtaceae ) and Pandanus ( Pandanaceae ) lined streams, waterholes, and swamps. Northern Longeared Bats are generally found in areas that receive more than 500 mm of rain/year. Food and Feeding. Northern Long-eared Bats primarily forage by gleaning off the ground and foliage and might also catch prey aerially. They are relatively slow fliers ( 5 km /h most of the time) but can reach top speeds of 16 km /h. Foraging usually occurs close to and among dense vegetation, and they are highly maneuverable in cluttered areas. Diet in the Top End region consisted largely of termites, water beetles, cockroaches, crickets, and true bugs and occasionally moths and spiders. They reportedly feed on geckos and gecko eggs, but this requires additional verification. Breeding. Male Northern Long-eared Bats with scrotal testes have been reported in May-November and pregnant females in September—November. Females apparently give birth to twins from late October to February. Activity patterns. Northern Long-eared Bats roost during the day under bark of large paperbark trees ( Melaleuca ), within foliage, and among Pandanus leaves. They have also been reported roosting in houses. They spend the night foraging. Call shape 1s very steep FM sweep, with peak frequencies of 50-52 kHz (mean 50-8 kHz). Movements, Home range and Social organization. The Northern Long-eared Bat probably roosts in small groups. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Northern Long-eared Bat is widespread and does not seem to face any major threats. Bibliography. Bullen & McKenzie (2002a), Churchill (2008), Churchill et al. (1984), Flannery (1995b), McKenzie & Churchill (2008), McKenzie, Fontanini et al. (1995), McKenzie, Reardon & Parnaby (2008), Milne (2002), Milne et al. (2016), Parnaby (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Nyctophilus arnhemensis	Nyctophilus		arnhemensis	Johnson	1959	0	Proc. Biol. Soc. Wash.	75:04:00	Northern Long-eared Bat	None.	Australia, Northern Territory, Cape Arnhem Peninsula, S of Yirkala, Rocky Bay. (12Â°13'S, 36Â°47'E).	N Australia.	Not listed.	Least Concern	 bifax species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctophilus arnhemensis	23	Northern Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Nyctophilus	NA	arnhemensis	D. H. Johnson	1959	0	Nyctophilus_arnhemensis	Johnson, D. H. (1959). Four New Mammals from the Northern Territory of Australia. Proceedings of the Biological Society of Washington, 72, 184.	https://www.biodiversitylibrary.org/page/34641962#page/198/mode/1up	USNM 284240		"Rocky Bay, south of Yirkala, Cape Arnhem Peninsula, Northern Territory, Australia (lat. 12Â° 16' S, long. 136Â° 47' E)."	-12.27	136.78	arnhemensis D. H. Johnson, 1959	NA	NA	Australia	Oceania	Australasia/Oceania	LC	0	0	0	Nyctophilus_arnhemensis	0	sciname match	Nyctophilus_arnhemensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	15000	Nyctophilus arnhemensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Nyctophilus	arnhemensis	Johnson, 1959	The genus Lamingtonia is a synonym of Nyctophilus .	20000000	Nyctophilus arnhemensis	Least Concern		2020	2019-07-09 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its relatively wide distribution, presumed large population, and because it is not believed to be declining.	This species has been recorded in rainforest patches, open Eucalyptus forests, in mangroves, and over freshwater lagoons and waterholes (McKenzie and Churchill 2008). At least west of, and including, the Dampier Peninsula, it has only been found in mangroves (N. McKenzie pers. comm.). It has been found roosting in foliage and under the roof of a house (McKenzie and Churchill 2008). Females may give birth to a litter of two young.	There appear to be no major threats to this species although inappropriate fire regimes may affect prey abundance.	It is common throughout its known range and is presumed to have a stable global population.	Stable	This species is endemic to northern Australia, where it is found in the tropical parts of the Gulf of Carpentaria, Queensland (H. Parnaby pers. comm.), Northern Territory, and Kimberley and along the coast of Western Australia. It is also present on offshore islands of the Melville, Bonaparte Archipelago, Groote Eylandt, and the Sir Edward Pellew Group (McKenzie and Churchill 2008). There is at least one record from the tip of the Cape York Peninsula in Queensland, so it may be more widely distributed (T. Reardon pers. comm.), but the taxonomy of this species requires clarification (H. Parnaby pers. comm.).		Terrestrial	This species is present in a number of protected areas.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Nyctophilus		arnhemensis	Johnson	1959	0	Proc. Biol. Soc. Wash.	75:04:00	Northern Long-eared Bat	None.	Australia, Northern Territory, Cape Arnhem Peninsula, S of Yirkala, Rocky Bay. (12Â°13'S, 36Â°47'E).	N Australia.	Not listed.	Least Concern	 bifax species group.	Nyctophilus arnhemensis	1005753	23	Northern Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Nyctophilus	NA	arnhemensis	D. H. Johnson	1959	0	Nyctophilus_arnhemensis	Johnson, D. H. (1959). Four New Mammals from the Northern Territory of Australia. Proceedings of the Biological Society of Washington, 72, 184.	https://www.biodiversitylibrary.org/page/34641962#page/198/mode/1up	USNM 284240		"Rocky Bay, south of Yirkala, Cape Arnhem Peninsula, Northern Territory, Australia (lat. 12Â° 16' S, long. 136Â° 47' E)."	-12.2667	136.7833	arnhemensis D. H. Johnson, 1959	NA	NA				Australia	Oceania	Australasia/Oceania	LC	0	0	0	Nyctophilus_arnhemensis	0	sciname match	Nyctophilus_arnhemensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctophilus_arnhemensis	1005753	23	Northern Long-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Nyctophilus	NA	arnhemensis	D. H. Johnson	0	Nyctophilus arnhemensis	Johnson, D.H. 1959-11-04. Four new mammals from the Northern Territory of Australia. Proceedings of the Biological Society of Washington 72:183-188.	https://www.biodiversitylibrary.org/page/34641963	USNM:MAMM:284240	holotype	http://n2t.net/ark:/65665/3ba913f94-6801-4793-88d0-120b8f406e3b	"Rocky Bay, south of Yirkala, Cape Arnhem Peninsula, Northern Territory, Australia (lat. 12Â° 16' S, long. 136Â° 47' E)."	-12.2667	136.7833	NA	NA				Australia	Oceania (Continent)	Australasia	LC	0	0	0	Nyctophilus_arnhemensis	0	sciname match	Nyctophilus_arnhemensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Nyctophilus		arnhemensis	Johnson	1959	0	Proc. Biol. Soc. Wash.	75:04:00	Northern Long-eared Bat	None.	Australia, Northern Territory, Cape Arnhem Peninsula, S of Yirkala, Rocky Bay. (12Â°13'S, 36Â°47'E).	N Australia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/15000/22010474/' target='_blank'>Least Concern</a>	bifaxspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctophilus arnhemensis; Nyctophilus arnhemensis; Nyctophilus arnhemensis; Nyctophilus arnhemensis; Nyctophilus arnhemensis; Nyctophilus arnhemensis; arnhemensis; Nyctophile d Arnhem; Arnhem-Langohrfledermaus; Nictofila de Arnhem; Northern Long-eared Bat; Northern Long-eared Bat; Northern Long-eared Bat; N. arnhemensis
