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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1071	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	Nyctimene minutus varius	Nyctimene minutus	Nyctimene varius	Nyctimene varius	Nyctimene varius	Nyctimene minutus	Nyctimene varia	Nyctimene varia	Nyctimene varia	Nyctimene varia	Nyctimene varia		[MSW3] albiventer species group. The status of minutus and varius is unclear, and is currently under review by K. Helgen (pers. comm.).; [HMW] Nyctimene varius K. Andersen, 1910 , “Mt. Mada, Buru [ Island , Moluccas , Indonesia ].” Nyctimene varius is currently in the albiventer species group along with N. albiventer , N. draconilla , N. masalai , N. malaitensis , and N. vizcaccia (including bougainuville). Type specimen of minutus from Manado in North Sulawesi seems to be the only specimen from Sulawesi and probably represents geographically mislabeled N. albiventer , it is considered a synonym of N. a. albiventer here, which restricts it to only Moluccas populations under the name N. varius , although validity of this nomenclatural change is still in need of additional research. Monotypic.; [batnames2022]  albiventer species group. Previously conidered a subspecies of minutus , which appears to be a misidentified albiventer (see that species account). The status of varius remains unclear, but we provisionally recognize it at the specific level pending additional research.; [MDD2022] previously known under the name minutus; next oldest name for the species previously known as N. minutus (name is now a synonym of N. albiventer since the type specimen likely represents a misidentified specimen of that species); [IUCN] The holotype from Manado in north Sulawesi of Nyctimene minutus is probably a geographically mislabelled "N. albiventer ". We follow Simmons (2005) here in including N. varius as a subspecies of N. minutus as the taxonomic status of N. minutus and N. m. varius is unclear.; [batnames2023]  albiventer species group. Previously conidered a subspecies of minutus , which appears to be a misidentified albiventer (see that species account). The status of varius remains unclear, but we provisionally recognize it at the specific level pending additional research.; [MDD2023] previously known under the name minutus; next oldest name for the species previously known as N. minutus (name is now a synonym of N. albiventer since the type specimen likely represents a misidentified specimen of that species); [MDD2025_2.0] previously known under the name minutus; next oldest name for the species previously known as N. minutus (name is now a synonym of N. albiventer since the type specimen likely represents a misidentified specimen of that species); [batnames2025_1.7] albiventerspecies group. Previously conidered a subspecies of minutus, which appears to be a misidentified albiventer (see that species account). The status of varia remains unclear, but we provisionally recognize it at the specific level pending additional research. We have changed the ending of the species epithet to agree with the feminine genus name Nyctimene; see Zijlstra (2023).; [MDD2025_2.2] previously known under the name minutus; varia is the next oldest name for the species previously known as N. minutus (name is now a synonym of N. albiventer since the type specimen likely represents a misidentified specimen of that species)								minutus, varius			minutus?			varius	The holotype from Manado in north Sulawesi of Nyctimene minutus is probably a geographically mislabelled "N. albiventer ". We follow Simmons (2005) here in including N. varius as a subspecies of N. minutus as the taxonomic status of N. minutus and N. m. varius is unclear.			varia	varia			varia Andersen, 1910						N/A					Distribu tion: Known only from Celebes and Buru (Moluc cas).																			36	species	N. minutus	ANDERSEN	1910	Nyctimene	genus	Nyctimene minutus				Dorsal stripe narrow (much less than one third breadth of furred area of back). Premaxillae not proclivous. Size relatively small (forearm length, 51-55 mm; maxillary tooth row length, 8.8- 10.0 mm). Upper molar subequal in size to last upper premolar. Ears not unusually broad and somewhat pointed. Inner cusp of middle upper premolar completely fused with outer.	Two subspecies:		1. N. minutus ANDERSEN 1910 [albiventer group].	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Nyctimene minutus	Nyctimene		minutus	K. Andersen		1910		Ann. Mag. Nat. Hist., ser. 7	6		622		Lesser Tube-nosed Fruit Bat	Indonesia, N Sulawesi, Minahassa, Tondano.	Sulawesi, C Moluccas.	IUCN/SSC Action Plan (1992) – Not Threatened. IUCN 2003 – Vulnerable.	varius K. Andersen, 1910.	albiventer species group. The status of minutus and varius is unclear, and is currently under review by K. Helgen (pers. comm.).	03AD87FAFFF8F6168CB1323FF930F3CF	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	118	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFF8F6168CB1323FF930F3CF.xml	Nyctimene varius	Pteropodidae	Nyctimene	varius	K. Andersen	1910	Petit Nyctiméne @fr | Kleiner Rohrennasenflughund @de | Nyctimeno pequeno @es	Nyctimene varius K. Andersen, 1910 , “Mt. Mada, Buru [ Island , Moluccas , Indonesia ].” Nyctimene varius is currently in the albiventer species group along with N. albiventer , N. draconilla , N. masalai , N. malaitensis , and N. vizcaccia (including bougainuville). Type specimen of minutus from Manado in North Sulawesi seems to be the only specimen from Sulawesi and probably represents geographically mislabeled N. albiventer , it is considered a synonym of N. a. albiventer here, which restricts it to only Moluccas populations under the name N. varius , although validity of this nomenclatural change is still in need of additional research. Monotypic.	Restricted to C Moluccas (Buru and Seram Is).	Ear c. 11 mm , forearm 55 mm (holotype). Fur of the Lesser Tubenosed Fruit Bat is somewhat long and woolly. Dorsal pelage is mottled dark brownish, with dark, thin dorsal stripe on lower one-half of back. Ventral pelage is lighter grayish white. Ears are bluntly pointed. Second digit of wing has a claw, and wing attaches at second digit of foot. Tail is short, black, and wrinkled, and narrow uropatagium connects at base and stretches to calcar at ankles. Skull and mandible are robust, and rostrum is short. Single lower incisor is completely deciduous, falling out before adulthood; lower molars are broad and rounded in dorsal view; C, replaces incisors and is long and powerful; P, is elongated and longer than P| and P,; and teeth are robust.	Mid-montane forests at elevations of 700-900 m .	No information.	No information.	No information.	No information.	Classified as Vulnerable on The IUCN Red List (as N. minutus ). The Lesser Tube-nosed Fruit Bat might be threatened by mining and logging that occurs throughout its distribution. It occurs in Manusela National Park (Seram Island). Virtually nothing is known about its ecology, and there is considerable confusion about its taxonomy. Additional research is needed, including sampling efforts because it is known from only a few specimens.	Andersen (1910b, 1912b) | Bergmans (2001) | Heaney & Peterson (1984) | Hutson, Helgen, Schlitter & Suyanto (2008)		103. Lesser Tube-nosed Fruit Bat Nyctimene varius French: Petit Nyctiméne / German: Kleiner Rohrennasenflughund / Spanish: Nyctimeno pequeno Taxonomy. Nyctimene varius K. Andersen, 1910 , “Mt. Mada, Buru [ Island , Moluccas , Indonesia ].” Nyctimene varius is currently in the albiventer species group along with N. albiventer , N. draconilla , N. masalai , N. malaitensis , and N. vizcaccia (including bougainuville). Type specimen of minutus from Manado in North Sulawesi seems to be the only specimen from Sulawesi and probably represents geographically mislabeled N. albiventer , it is considered a synonym of N. a. albiventer here, which restricts it to only Moluccas populations under the name N. varius , although validity of this nomenclatural change is still in need of additional research. Monotypic. Distribution. Restricted to C Moluccas (Buru and Seram Is). Descriptive notes. Ear c. 11 mm , forearm 55 mm (holotype). Fur of the Lesser Tubenosed Fruit Bat is somewhat long and woolly. Dorsal pelage is mottled dark brownish, with dark, thin dorsal stripe on lower one-half of back. Ventral pelage is lighter grayish white. Ears are bluntly pointed. Second digit of wing has a claw, and wing attaches at second digit of foot. Tail is short, black, and wrinkled, and narrow uropatagium connects at base and stretches to calcar at ankles. Skull and mandible are robust, and rostrum is short. Single lower incisor is completely deciduous, falling out before adulthood; lower molars are broad and rounded in dorsal view; C, replaces incisors and is long and powerful; P, is elongated and longer than P| and P,; and teeth are robust. Habitat. Mid-montane forests at elevations of 700-900 m . Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Vulnerable on The IUCN Red List (as N. minutus ). The Lesser Tube-nosed Fruit Bat might be threatened by mining and logging that occurs throughout its distribution. It occurs in Manusela National Park (Seram Island). Virtually nothing is known about its ecology, and there is considerable confusion about its taxonomy. Additional research is needed, including sampling efforts because it is known from only a few specimens. Bibliography. Andersen (1910b, 1912b), Bergmans (2001), Heaney & Peterson (1984), Hutson, Helgen, Schlitter & Suyanto (2008).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Nyctimene varius	Nyctimene		varius	K. Andersen	1910	0	Ann. Mag. Nat. Hist.	ser. 8, 6: 622	Lesser Tube-nosed Fruit Bat	None.	Indonesia, Buru, Mount Mada.	Sulawesi, C Moluccas.	Not listed.	Vulnerable under Nyctimene minutus 	 albiventer species group. Previously conidered a subspecies of minutus , which appears to be a misidentified albiventer (see that species account). The status of varius remains unclear, but we provisionally recognize it at the specific level pending additional research.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctimene varius	23	Lesser Tube-nosed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	NYCTIMENINAE	NA	Nyctimene	NA	varius	K. Andersen	1910	0						"Mt. Mada, Buru [Island, Moluccas, Indonesia]."			varius K. Andersen, 1910	previously known under the name minutus; next oldest name for the species previously known as N. minutus (name is now a synonym of N. albiventer since the type specimen likely represents a misidentified specimen of that species)	Wilson, D. E. & Mittermeier, R. A.Handbook of the Mammals of the World. 9. Bats. Lynx Edicions, Barcelona.	Indonesia	Oceania	Australasia/Oceania	VU	0	0	0	Nyctimene_varius	0	oldname match	Nyctimene_minutus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14960	Nyctimene minutus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Nyctimene	minutus	K. Andersen, 1910	The holotype from Manado in north Sulawesi of Nyctimene minutus is probably a geographically mislabelled "N. albiventer ". We follow Simmons (2005) here in including N. varius as a subspecies of N. minutus as the taxonomic status of N. minutus and N. m. varius is unclear.	20000000	Nyctimene minutus	Vulnerable	B1ab(ii,iii,v)	2021	2020-11-25 00:00:00 UTC	3.1	English	Nyctimene minutus is assessed as Vulnerable under criterion B1ab(ii,iii,v) as its extent of occurrence (EOO) is 13,403 km². Its range is severely fragmented over two islands, and it is limited to areas above 700 m. There is a continuing decline in the extent and quality of its habitat and area of occupancy (AOO) in Buru and Seram islands mainly due to clear-cut logging and mining of its forest habitat. The species population is inferred to be declining due to the loss of habitat.	Nyctimene minutus is a frugivorous species that has been recorded from mid-montane forest (700–900 m) on Buru and Seram islands. The species is assumed to roost singly or in small colonies in the foliage of trees based on similar species. Research is needed to determine the species ecology.	Habitat loss due to clear-cut logging and probably also mining are the main threats to this species (Dudun pers. comm.). This species is probably not hunted, as it is too small.	Data on the species population status are very limited as it is only known from a small number of specimens. The species is known from two (2) locations (Buru and Seram islands), each is assumed to represent a subpopulation. The overall population and number of mature individuals of Nyctimene minutus is inferred to be declining given the species restricted distribution and the past and ongoing forest loss and degradation which is resulting in a decline in its area of occupancy (AOO) on the islands of Buru and Seram. Additional research is needed to confirm the species population status and trends throughout the species range.	Decreasing	Nyctimene minutus is confined to mid to high elevation regions of the islands of Buru and Seram in Indonesia (700 to ca 900 m). The species extent of occurrence is 13,403 km². Surveys on the island of Ambon have not confirmed the presence of N. minutus . The geographically mislabelled specimen from Manado in north Sulawesi is not considered here as part of the range of the species. Additional research is warranted to further illuminate the species distribution.	The species is not known to be hunted or used.	Terrestrial	There are no species-specific conservation measures in place for the species, but. it is present in Manusela National Park on Seram Island. Protection, management, and restoration of its forest habitats is needed to secure the species. Taxonomic research is warranted to clarify the relationship between Nyctimene minutus and N. m. varius . Ecological research is needed to understand the species population status and trends, and determine the species life history, habitat requirements, and threats.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Nyctimene		varius	K. Andersen	1910	0	Ann. Mag. Nat. Hist.	ser. 8, 6: 622	Lesser Tube-nosed Fruit Bat	None.	Indonesia, Buru, Mount Mada.	Sulawesi, C Moluccas.	Not listed.	Vulnerable under Nyctimene minutus 	 albiventer species group. Previously conidered a subspecies of minutus , which appears to be a misidentified albiventer (see that species account). The status of varius remains unclear, but we provisionally recognize it at the specific level pending additional research.	Nyctimene varia	1004426	23	Lesser Tube-nosed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	NYCTIMENINAE	NA	Nyctimene	NA	varia	K. Andersen	1910	0						"Mt. Mada, Buru [Island, Moluccas, Indonesia]."			varia K. Andersen, 1910	previously known under the name minutus; next oldest name for the species previously known as N. minutus (name is now a synonym of N. albiventer since the type specimen likely represents a misidentified specimen of that species)	Wilson, D. E. & Mittermeier, R. A.Handbook of the Mammals of the World. 9. Bats. Lynx Edicions, Barcelona.				Indonesia	Oceania	Australasia/Oceania	VU	0	0	0	Nyctimene_varius	0	oldname match	Nyctimene_minutus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctimene_varia	1004426	23	Lesser Tube-nosed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Nyctimeninae	NA	Nyctimene	NA	varia	Andersen	0	Nyctimene varius	Andersen, K.C. 1910-12-01. Ten new fruit-bats of the genera _Nyctimene_, _Cynopterus_, and _Eonycteris_. Annals and Magazine of Natural History (8)6(36):621-625.	https://www.biodiversitylibrary.org/page/18619863	BMNH:Mamm:1910.11.13.1	holotype	https://data.nhm.ac.uk/object/0d3cdfc2-bd47-4e75-9c06-bbe0690c0e6c	"Mt. Mada, Buru [Island, Moluccas, Indonesia]."			previously known under the name minutus; varia is the next oldest name for the species previously known as N. minutus (name is now a synonym of N. albiventer since the type specimen likely represents a misidentified specimen of that species)	Wilson, D. E. & Mittermeier, R. A.Handbook of the Mammals of the World. 9. Bats. Lynx Edicions, Barcelona.				Indonesia	Oceania (Continent)	Australasia	VU (as Nyctimene varius)	0	0	0	Nyctimene_varius	0	oldname match	Nyctimene_minutus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Nyctimene		varia	K. Andersen	1910	0	Ann. Mag. Nat. Hist.	ser. 8, 6: 622	Lesser Tube-nosed Fruit Bat	None.	Indonesia, Buru, Mount Mada.	Sulawesi, C Moluccas.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14960/22008183/' target='_blank'>Vulnerable as Nyctimene varius</a>	albiventerspecies group. Previously conidered a subspecies of minutus, which appears to be a misidentified albiventer (see that species account). The status of varia remains unclear, but we provisionally recognize it at the specific level pending additional research. We have changed the ending of the species epithet to agree with the feminine genus name Nyctimene; see Zijlstra (2023).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctimene minutus; Nyctimene varius; Nyctimene varius; Nyctimene varius; Nyctimene minutus; Nyctimene varius; minutus; varius; minutus?; varius; Petit Nyctiméne; Kleiner Rohrennasenflughund; Nyctimeno pequeno; Lesser Tube-nosed Fruit Bat; Lesser Tube-nosed Fruit Bat; Lesser Tube-nosed Fruit Bat; N. varia