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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1062	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis	Nyctimene cyclotis		[MSW2] Formerly included certans, but see Peterson (1991).; [MSW3] cyclotis species group. Apparently does not include certans; see Peterson (1991) and Flannery (1995a, b), though also see Bonaccorso (1998) and comments under certans.; [HMW] Nyctimene cyclotis K. Andersen, 1910 , “Arfak Mts. [Manokwari Division], N.W. New Guinea ,” Indonesia . Nyctimene cyclotis 1s currently in the cyclotis species group along with N. certans and N. wrightae . Nyctimene certans is sometimes considered a subspecies of N. cyclotis , but morphological studies have confirmed their specific status. Distribution is often recorded encompassing much of New Guinea , but this is now recognized as a distinct species, N. wrightae . Specimens collected in Mansuar and Waigeo islands have been recently attributed to N. cyclotis based on morphological grounds, but these island populations are currently under revision. They are tentatively included under N. cyclotis here until they are thoroughly investigated. Monotypic.; [batnames2022]  cyclotis species group. Does not include certans; see Peterson (1991), Flannery (1995a, b),and Irwin (2017); but see also Bonaccorso (1998) and comments under certans. Revised by Irwin (2017).; [IUCN] A recent taxonomic study by Irwin (2017) resolved the taxonomic status of four similar-sized Nyctimene that occur on the island of New Guinea and provided revised diagnoses for three of them. The distinct species status of N. cyclotis was supported by morphological analyses, and it is not synonymous with N. certans .; [batnames2023]  cyclotis species group. Does not include certans; see Peterson (1991), Flannery (1995a, b),and Irwin (2017); but see also Bonaccorso (1998) and comments under certans. Revised by Irwin (2017).; [batnames2025_1.7] cyclotisspecies group. Does not include certans; see Peterson (1991), Flannery (1995a, b),and Irwin (2017); but see also Bonaccorso (1998) and comments under certans. Revised by Irwin (2017).							cyclotis, certans							cyclotis	A recent taxonomic study by Irwin (2017) resolved the taxonomic status of four similar-sized Nyctimene that occur on the island of New Guinea and provided revised diagnoses for three of them. The distinct species status of N. cyclotis was supported by morphological analyses, and it is not synonymous with N. certans .			cyclotis 	cyclotis 			cyclotis Andersen, 1910		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Round-eared tube-nosed	New Guinea bat	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nyctimene cyclotis	Indonesia, Irian Jaya, Manokwari Div., Arfak Mtns.	K. Andersen	1910	Ann. Mag. Nat. Hist., ser. 7, 6:623.	Distribution: Confined to the moun tains of New Guinea and New Britain.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Round-eared tube-nosed bat	New Guinea; E New Britain	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	K. Andersen	1910	Ann. Mag. Nat. Hist., ser. 7, 6:623.	Formerly included certans, but see Peterson (1991).	New Guinea; New Britain (Bismarck Arch.).	Indonesia, Irian Jaya, Manokwari Div., Arfak Mtns.		ANDERSEN	1910	Dorsal stripe narrow (much less than one third breadth of furred area of back). Premaxillae not proclivous. Size relatively small (forearm length, 53-58 mm; maxillary tooth row length, 8.5-9.8 mm). Upper molar clearly smaller than last upper premolar. Ears unusually broad and rounded.	Distribution: Confined to the moun tains of New Guinea and New Britain.	Two sub species:	N. c. cyclotis (extreme northwestern New Guinea), N. c. certans (remainder of New Guinea and, presumeably, New Britain).	36	species	N. cyclotis	ANDERSEN	1910	Nyctimene	genus	Nyctimene cyclotis				Dorsal stripe narrow (much less than one third breadth of furred area of back). Premaxillae not proclivous. Size relatively small (forearm length, 53-58 mm; maxillary tooth row length, 8.5-9.8 mm). Upper molar clearly smaller than last upper premolar. Ears unusually broad and rounded.	Two sub species:		4. N. cyclotis ANDERSEN 1910 [cyclotis group],	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Nyctimene cyclotis	Nyctimene		cyclotis	K. Andersen		1910		Ann. Mag. Nat. Hist., ser. 7	6		623		Round-eared Tube-nosed Fruit Bat	Indonesia, Prov. of Papua, Manokwari Div., Arfak Mtns.	Arfak Mtns. (New Guinea). Specimens from Mansuar Isl. (Prov. of Papua, Indonesia) may also represent cyclotis (Meinig, 2002). Specimens from New Britain formerly assigned to this species apparently represent vizcaccia (Bonaccorso, 1998).	IUCN/SSC Action Plan (1992) – No Data. IUCN 2003 – Lower Risk (nt).		cyclotis species group. Apparently does not include certans; see Peterson (1991) and Flannery (1995a, b), though also see Bonaccorso (1998) and comments under certans.	03AD87FAFFFFF6168C673329FE08FD05	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	117	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFFFF6168C673329FE08FD05.xml	Nyctimene cyclotis	Pteropodidae	Nyctimene	cyclotis	K. Andersen	1910	Nyctimene a oreilles rondes @fr | Rundohr Réhrennasenflughund @de | Nyctimeno de orejas redondas @es | Round-eared Tube-nosed Bat @en	Nyctimene cyclotis K. Andersen, 1910 , “Arfak Mts. [Manokwari Division], N.W. New Guinea ,” Indonesia . Nyctimene cyclotis 1s currently in the cyclotis species group along with N. certans and N. wrightae . Nyctimene certans is sometimes considered a subspecies of N. cyclotis , but morphological studies have confirmed their specific status. Distribution is often recorded encompassing much of New Guinea , but this is now recognized as a distinct species, N. wrightae . Specimens collected in Mansuar and Waigeo islands have been recently attributed to N. cyclotis based on morphological grounds, but these island populations are currently under revision. They are tentatively included under N. cyclotis here until they are thoroughly investigated. Monotypic.	NW New Guinea (Arfak Mts); also tentatively on Raja Ampat Is (Waigeo and Mansuar).	Ear 13-5-13- 7 mm , hindfoot 12-14- 6 mm , forearm 54- 2-56 mm . Rostrum of the Round-eared Tube-nosed Fruit Bat is short, with long tubular divergent nostrils. Ears are rounded, with blunt tips, and thickened along tops; eyes are large, with dark reddish-brown irises. Dorsal pelage is grizzled grayish brown and browner on head, with almost indistinguishable ill-defined, short, thin dorsal stripe extending from mid-back to rump. Ventral pelage is paler dull cream. Wings and ears are dark brown; ears, tube nostrils, and wing digits have some yellow spotting. Second digit of wing is shorter in the Round-eared Tube-nosed Fruit Bat than in the Mountain Tube-nosed Fruit Bat (N. certans ); second digit of wing has a claw, and wing attaches at second digit of foot. Tail is short, black, and wrinkled, and narrow uropatagium connects at base and stretches to calcar at ankles. Claws are black. Skull and mandible are robust, and rostrum is short. Single lower incisor is completely deciduous, falling out before adulthood; lower molars are broad and rounded in dorsal view; C, replaces incisors and is long and powerful; P,is elongated and longer than P and P,; they have narrow and rounded dental arcade; and it differs from the Mountain Tube-nosed Fruit Bat by having relatively longer upper post-canine tooth row.	No specific information but probably montane tropical forests. All ecological accounts on the Round-eared Tube-nosed Fruit Bat are based on specimens that are now primarily attributed to the Happy Tube-nosed Fruit Bat (N. wrightae ).	No information.	No information.	No information.	No information.	Classified as Data Deficient on The IUCN Red List. The Round-eared Tube-nosed Fruit Bat is currently known from two male specimens collected more than 100 years ago from the Arfak Mountains. Virtually nothing is known about its ecology and threats.	Donnellan et al. (1995) | Helgen (2007a) | Hutson, Suyanto, Helgen & Bonaccorso (2008a) | Irwin (2017) | Wiantoro (2011)		100. Round-eared Tube-nosed Fruit Bat Nyctimene cyclotis French: Nyctimene a oreilles rondes / German: RundohrRéhrennasenflughund / Spanish: Nyctimeno de orejas redondas Other common names: Round-eared Tube-nosed Bat Taxonomy. Nyctimene cyclotis K. Andersen, 1910 , “Arfak Mts. [Manokwari Division], N.W. New Guinea ,” Indonesia . Nyctimene cyclotis 1s currently in the cyclotis species group along with N. certans and N. wrightae . Nyctimene certans is sometimes considered a subspecies of N. cyclotis , but morphological studies have confirmed their specific status. Distribution is often recorded encompassing much of New Guinea , but this is now recognized as a distinct species, N. wrightae . Specimens collected in Mansuar and Waigeo islands have been recently attributed to N. cyclotis based on morphological grounds, but these island populations are currently under revision. They are tentatively included under N. cyclotis here until they are thoroughly investigated. Monotypic. Distribution. NW New Guinea (Arfak Mts); also tentatively on Raja Ampat Is (Waigeo and Mansuar). Descriptive notes. Ear 13-5-13- 7 mm , hindfoot 12-14- 6 mm , forearm 54- 2-56 mm . Rostrum of the Round-eared Tube-nosed Fruit Bat is short, with long tubular divergent nostrils. Ears are rounded, with blunt tips, and thickened along tops; eyes are large, with dark reddish-brown irises. Dorsal pelage is grizzled grayish brown and browner on head, with almost indistinguishable ill-defined, short, thin dorsal stripe extending from mid-back to rump. Ventral pelage is paler dull cream. Wings and ears are dark brown; ears, tube nostrils, and wing digits have some yellow spotting. Second digit of wing is shorter in the Round-eared Tube-nosed Fruit Bat than in the Mountain Tube-nosed Fruit Bat (N. certans ); second digit of wing has a claw, and wing attaches at second digit of foot. Tail is short, black, and wrinkled, and narrow uropatagium connects at base and stretches to calcar at ankles. Claws are black. Skull and mandible are robust, and rostrum is short. Single lower incisor is completely deciduous, falling out before adulthood; lower molars are broad and rounded in dorsal view; C, replaces incisors and is long and powerful; P,is elongated and longer than P and P,; they have narrow and rounded dental arcade; and it differs from the Mountain Tube-nosed Fruit Bat by having relatively longer upper post-canine tooth row. Habitat. No specific information but probably montane tropical forests. All ecological accounts on the Round-eared Tube-nosed Fruit Bat are based on specimens that are now primarily attributed to the Happy Tube-nosed Fruit Bat (N. wrightae ). Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. The Round-eared Tube-nosed Fruit Bat is currently known from two male specimens collected more than 100 years ago from the Arfak Mountains. Virtually nothing is known about its ecology and threats. Bibliography. Donnellan et al. (1995), Helgen (2007a), Hutson, Suyanto, Helgen & Bonaccorso (2008a), Irwin (2017), Wiantoro (2011).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Nyctimene cyclotis	Nyctimene		cyclotis	K. Andersen	1910	0	Ann. Mag. Nat. Hist.	ser. 8, 6: 623	Round-eared Tube-nosed Fruit Bat	None.	Indonesia, Prov. of Papua, Manokwari Div., Arfak Mtns.	Arfak Mtns. (New Guinea). Specimens from Mansuar Isl. (Prov. of Papua, Indonesia) may also represent cyclotis (Meinig, 2002). Specimens from New Britain formerly assigned to this species apparently represent vizcaccia (Bonaccorso, 1998).	Not listed.	Data Deficient	 cyclotis species group. Does not include certans; see Peterson (1991), Flannery (1995a, b),and Irwin (2017); but see also Bonaccorso (1998) and comments under certans. Revised by Irwin (2017).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctimene cyclotis	23	Round-eared Tube-nosed Fruit Bat	Round-eared Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	NYCTIMENINAE	NA	Nyctimene	NA	cyclotis	K. Andersen	1910	0	Nyctimene_cyclotis	Andersen, K. (1910). Ten new fruit-bats of the genera Nyctimene, Cynopterus and Eonycteris. Annals and Magazine of Natural History, ser. 8, 6, 623.	https://www.biodiversitylibrary.org/item/61726#page/637/mode/1up	BM 1910.7.16.9		"Arfak Mts. [Manokwari Division], N.W. New Guinea," Indonesia.			cyclotis K. Andersen, 1910	NA	NA	Indonesia	Oceania	Australasia/Oceania	DD	0	0	0	Nyctimene_cyclotis	0	sciname match	Nyctimene_cyclotis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14955	Nyctimene cyclotis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Nyctimene	cyclotis	Andersen, 1910	A recent taxonomic study by Irwin (2017) resolved the taxonomic status of four similar-sized Nyctimene that occur on the island of New Guinea and provided revised diagnoses for three of them. The distinct species status of N. cyclotis was supported by morphological analyses, and it is not synonymous with N. certans .	20000000	Nyctimene cyclotis	Data Deficient		2021	2020-11-22 00:00:00 UTC	3.1	English	This species is assessed as Data Deficient because of a lack of information about its distribution and population size.	It is presumed to have similar habitat requirements to N. certans , and occurs in primary and secondary forest, forest edge, and rural gardens in hill forests and at lower montane elevations. It is also presumed to occur in moss forest at mid-montane elevations. Meinig (2002) reports captures on Mansuar Island at 5 m asl.	No major threats to this species have been identified that could cause widespread or imminent decline.	The population size of this species is not known. Further survey effort is required to determine the extent of the distribution.	Unknown	This species is known from the Arfak Mountains on the Vogelkop Peninsula in western New Guinea. It has also been reported by Meinig (2002) as occurring on Mansuar Island, Indonesia, but this should be confirmed following the information provided by Irwin (2017) who did not examine representatives from there. The precise altitudinal range is not known.	The species is not known to be hunted, traded, or used.	Terrestrial	It is presumed to be present in most protected areas within the species range (e.g., the Pegunungan Arfak Nature Reserve). Broadscale protection of forests will help maintain population size in this species. Comprehensive research into the species ecology, distribution, threats, and population status and trends are needed.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Nyctimene		cyclotis	K. Andersen	1910	0	Ann. Mag. Nat. Hist.	ser. 8, 6: 623	Round-eared Tube-nosed Fruit Bat	None.	Indonesia, Prov. of Papua, Manokwari Div., Arfak Mtns.	Arfak Mtns. (New Guinea). Specimens from Mansuar Isl. (Prov. of Papua, Indonesia) may also represent cyclotis (Meinig, 2002). Specimens from New Britain formerly assigned to this species apparently represent vizcaccia (Bonaccorso, 1998).	Not listed.	Data Deficient	 cyclotis species group. Does not include certans; see Peterson (1991), Flannery (1995a, b),and Irwin (2017); but see also Bonaccorso (1998) and comments under certans. Revised by Irwin (2017).	Nyctimene cyclotis	1004418	23	Round-eared Tube-nosed Fruit Bat	Round-eared Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	NYCTIMENINAE	NA	Nyctimene	NA	cyclotis	K. Andersen	1910	0	Nyctimene_cyclotis	Andersen, K. (1910). Ten new fruit-bats of the genera Nyctimene, Cynopterus and Eonycteris. Annals and Magazine of Natural History, ser. 8, 6, 623.	https://www.biodiversitylibrary.org/item/61726#page/637/mode/1up	BM 1910.7.16.9		"Arfak Mts. [Manokwari Division], N.W. New Guinea," Indonesia.			cyclotis K. Andersen, 1910	NA	NA				Indonesia	Oceania	Australasia/Oceania	DD	0	0	0	Nyctimene_cyclotis	0	sciname match	Nyctimene_cyclotis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctimene_cyclotis	1004418	23	Round-eared Tube-nosed Fruit Bat	Round-eared Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Nyctimeninae	NA	Nyctimene	NA	cyclotis	Andersen	0	Nyctimene cyclotis	Andersen, K.C. 1910-12-01. Ten new fruit-bats of the genera _Nyctimene_, _Cynopterus_, and _Eonycteris_. Annals and Magazine of Natural History (8)6(36):621-625.	https://www.biodiversitylibrary.org/page/18619864	BMNH:Mamm:1910.7.16.9	holotype	https://data.nhm.ac.uk/object/91a6c085-8f78-469c-be34-53e785dba6f7	"Arfak Mts. [Manokwari Division], N.W. New Guinea," Indonesia.			NA	NA				Indonesia	Oceania (Continent)	Australasia	DD	0	0	0	Nyctimene_cyclotis	0	sciname match	Nyctimene_cyclotis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Nyctimene		cyclotis	K. Andersen	1910	0	Ann. Mag. Nat. Hist.	ser. 8, 6: 623	Round-eared Tube-nosed Fruit Bat	None.	Indonesia, Prov. of Papua, Manokwari Div., Arfak Mtns.	Arfak Mtns. (New Guinea). Specimens from Mansuar Isl. (Prov. of Papua, Indonesia) may also represent cyclotis (Meinig, 2002). Specimens from New Britain formerly assigned to this species apparently represent vizcaccia (Bonaccorso, 1998).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14955/22008406/' target='_blank'>Data Deficient</a>	cyclotisspecies group. Does not include certans; see Peterson (1991), Flannery (1995a, b),and Irwin (2017); but see also Bonaccorso (1998) and comments under certans. Revised by Irwin (2017).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctimene cyclotis; Nyctimene cyclotis; Nyctimene cyclotis; Nyctimene cyclotis; Nyctimene cyclotis; Nyctimene cyclotis; cyclotis; Nyctimene a oreilles rondes; Rundohr Réhrennasenflughund; Nyctimeno de orejas redondas; Round-eared Tube-nosed Bat; Round-eared Tube-nosed Fruit Bat; Round-eared Tube-nosed Bat; Round-eared Tube-nosed Fruit Bat; Round-eared Tube-nosed Fruit Bat; N. cyclotis