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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1056	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello	Nyctimene aello		[MSW2] Does not include celaeno; see Hill (1983).; [MSW3] aello species group. Includes celaeno, see Flannery (1995a, b) and Bergmans (2001). Also see Bonaccorso (1998).; [HMW] Cephalotes aello Thomas, 1900 , “Milne Bay [Milne Bay Province], S.E. [Papua] New Guinea . Nyctimene aello is the only member of the aello species group, which is one of four groups currently recognized in Nyctimene ( aello , cyclotis , albiventer , and cephalotes ). Species groups of Nyctimene here do not necessarily reflect genetic relationships but do reflect current morphological similarities. Full-scale revision of the entire genus is currently underway that will completely rework species recognition and evolutionary relationships. Thus, taxonomy of Nyctimene presented here should not be presented as authoritative and is based on taxonomy presented in Mammal Species of the World in 2005, with a few alterations to avoid additional confusion to the already convoluted taxonomy of Nyctimene . Nyctimene celaeno has been considered a distinct subspecies of N. aello or a distinct species, but it is tentatively considered a synonym of N. aello here. Monotypic.; [batnames2022]  aello species group. Includes celaeno, see Flannery (1995a, b) and Bergmans (2001). Also see Bonaccorso (1998).; [IUCN] The eastern and western distribution on the island of New Guinea might represent subspecies, but this has not been tested (Flannery 1995).; [batnames2023]  aello species group. Includes celaeno, see Flannery (1995a, b) and Bergmans (2001). Also see Bonaccorso (1998).; [batnames2025_1.7] aellospecies group. Includes celaeno, see Flannery (1995a, b) and Bergmans (2001). Also see Bonaccorso (1998).									celaeno		celaeno	aello 	aello - celaeno	aello, celaeno	The eastern and western distribution on the island of New Guinea might represent subspecies, but this has not been tested (Flannery 1995).	aello 	aello - celaeno	aello, celaeno	aello, celaeno	aello 	aello - celaeno	aello (O. Thomas, 1900)|celaeno O. Thomas, 1922		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Broad-striped tube-nosed	Halmahera, New Guinea bat	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nyctimene aello	Papua New Guinea, Milne Bay Prov., Milne Bay.	Thomas	1900	Ann. Mag. Nat. Hist., ser. 7, 5:216.	Distribution: Confined to New Guinea (where apparently widely distributed), in cluding Misol (off its western end).		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Broad-striped tube-nosed	New Guinea bat	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1900	Ann. Mag. Nat. Hist., ser. 7, 5:216.	Does not include celaeno; see Hill (1983).	New Guinea.	Papua New Guinea, Milne Bay Prov., Milne Bay.		THOMAS	1900	Dor sal stripe broad (about one third breadth of furred area of back). Premaxillae relatively proclivous. Size large (forearm length, 77-85 mm; maxillary tooth row length, 12.2-14.2 mm). Frontal region of skull relatively depressed. Rostrum not greatly shortened.	Distribution: Confined to New Guinea (where apparently widely distributed), in cluding Misol (off its western end).	No sub species.		37	species	N. aello	THOMAS	1900	Nyctimene	genus	Nyctimene aello				Dor sal stripe broad (about one third breadth of furred area of back). Premaxillae relatively proclivous. Size large (forearm length, 77-85 mm; maxillary tooth row length, 12.2-14.2 mm). Frontal region of skull relatively depressed. Rostrum not greatly shortened.	No sub species.		13. N. aello (THOMAS 1900) [aello group],	13	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Nyctimene aello	Nyctimene		aello	Thomas	y	1900		Ann. Mag. Nat. Hist., ser. 7	5		216		Broad-striped Tube-nosed Fruit Bat	Papua New Guinea, Milne Bay Prov., Milne Bay.	Mainland New Guinea; Kairiru and Admosin Isls (Papua New Guinea); Misool and Salawati Isl (Prov. of Papua, Indonesia).	IUCN/SSC Action Plan (1992) – Rare as N. aello, No Data: Limited Distribution as N. celaeno. IUCN 2003 – Lower Risk (nt) as N. aello; Vulnerable as N. celaeno.	celaeno Thomas, 1922.	aello species group. Includes celaeno, see Flannery (1995a, b) and Bergmans (2001). Also see Bonaccorso (1998).	03AD87FAFFFFF61189693EF4F6D4F977	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	117	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFFFF61189693EF4F6D4F977.xml	Nyctimene aello	Pteropodidae	Nyctimene	aello	Thomas	1900	Grand Nyctimene @fr | Breitstreifen-Réhrennasenflughund @de | Nyctimeno grande @es | Greater Tube-nosed Bat @en | Dark Tube-nosed Fruit Bat (celaeno) @en	Cephalotes aello Thomas, 1900 , “Milne Bay [Milne Bay Province], S.E. [Papua] New Guinea . Nyctimene aello is the only member of the aello species group, which is one of four groups currently recognized in Nyctimene ( aello , cyclotis , albiventer , and cephalotes ). Species groups of Nyctimene here do not necessarily reflect genetic relationships but do reflect current morphological similarities. Full-scale revision of the entire genus is currently underway that will completely rework species recognition and evolutionary relationships. Thus, taxonomy of Nyctimene presented here should not be presented as authoritative and is based on taxonomy presented in Mammal Species of the World in 2005, with a few alterations to avoid additional confusion to the already convoluted taxonomy of Nyctimene . Nyctimene celaeno has been considered a distinct subspecies of N. aello or a distinct species, but it is tentatively considered a synonym of N. aello here. Monotypic.	[Lowland New Guinea along with Raja Ampat Is (Salawati and Misool) and Kairiru and Admosin Is, off N coast.	Head-body 103-144 mm , tail 14- 1-31 mm , ear 13- 4-24 mm , hindfoot 16-22- 7 mm , forearm 77-7-88- 1 mm ; weight 70- 3-93 g . Rostrum of the Broadstriped Tube-nosed Fruit Bat is short, with long tubular divergent nostrils that curve outward at rims. Ears are rounded, with bluntly pointed tips; eyes are large, with dark reddish-brown irises. Dorsal and ventral pelage is uniformly pale grayish brown, and there is thick ( 8-14 mm across) dark brown dorsal stripe. Wings and ears are light brown; ears generally have some yellow spotting, and wings have some darker brown and lighter patches throughout. Second digit of wing has a claw, and wing attaches at second digit of foot. Tail is very short, black, and wrinkled, and narrow uropatagium connects at base and stretches to calcar at ankles. Claws are black. Skull and mandible are robust, and rostrum is short. Dental formula for all species of Nyctimene is11/0,C1/1,P3/3,M 1/2 (x2) = 24.	Primary rainforest (but one collection in secondary rainforest) from sea level up to elevations of ¢. 1000 m but extremely rare above 200 m .	Broad-striped Tube-nosed Fruit Bats are frugivorous and likely a Ficus ( Moraceae ) specialist. They have been observed picking ripe figs after briefly hovering near clusters offigs.	Pregnant Broad-striped Tube-nosed Fruit Bats have been captured in January—February and a lactating female in September.	Broad-striped Tube-nosed Fruit Bats are nocturnal.	The Broad-striped Tube-nosed Fruit Bat appears to occur at low densities.	Classified as Least Concern on The IUCN Red List. The Broad-striped Tube-nosed Fruit Bat is widespread and common throughout its distribution. It faces no known major threats, but very little is known about its ecology and threats.	Bergmans (2001) | Bonaccorso (1998) | Bonaccorso & Helgen (2008b) | Flannery (1995a, 1995b) | Giannini & Simmons (2007a) | Helgen (2007a) | Irwin (2017) | McKean (1972) | Simmons (2005)	https://zenodo.org/record/6448871/files/figure.png	99. Broad-striped Tube-nosed Fruit Bat Nyctimene aello French: Grand Nyctimene / German: Breitstreifen-Réhrennasenflughund / Spanish: Nyctimeno grande Other common names: Greater Tube-nosed Bat , Dark Tube-nosed Fruit Bat (celaeno) Taxonomy. Cephalotes aello Thomas, 1900 , “Milne Bay [Milne Bay Province], S.E. [Papua] New Guinea . Nyctimene aello is the only member of the aello species group, which is one of four groups currently recognized in Nyctimene ( aello , cyclotis , albiventer , and cephalotes ). Species groups of Nyctimene here do not necessarily reflect genetic relationships but do reflect current morphological similarities. Full-scale revision of the entire genus is currently underway that will completely rework species recognition and evolutionary relationships. Thus, taxonomy of Nyctimene presented here should not be presented as authoritative and is based on taxonomy presented in Mammal Species of the World in 2005, with a few alterations to avoid additional confusion to the already convoluted taxonomy of Nyctimene . Nyctimene celaeno has been considered a distinct subspecies of N. aello or a distinct species, but it is tentatively considered a synonym of N. aello here. Monotypic. Distribution. [Lowland New Guinea along with Raja Ampat Is (Salawati and Misool) and Kairiru and Admosin Is, off N coast. Descriptive notes. Head-body 103-144 mm , tail 14- 1-31 mm , ear 13- 4-24 mm , hindfoot 16-22- 7 mm , forearm 77-7-88- 1 mm ; weight 70- 3-93 g . Rostrum of the Broadstriped Tube-nosed Fruit Bat is short, with long tubular divergent nostrils that curve outward at rims. Ears are rounded, with bluntly pointed tips; eyes are large, with dark reddish-brown irises. Dorsal and ventral pelage is uniformly pale grayish brown, and there is thick ( 8-14 mm across) dark brown dorsal stripe. Wings and ears are light brown; ears generally have some yellow spotting, and wings have some darker brown and lighter patches throughout. Second digit of wing has a claw, and wing attaches at second digit of foot. Tail is very short, black, and wrinkled, and narrow uropatagium connects at base and stretches to calcar at ankles. Claws are black. Skull and mandible are robust, and rostrum is short. Dental formula for all species of Nyctimene is11/0,C1/1,P3/3,M 1/2 (x2) = 24. Single lowerincisor is completely deciduous, falling out before adulthood; lower molars are broad and rounded in dorsal view; C, replaces incisors and is long and powerful; and P, is elongated and longer than P and P,, Habitat. Primary rainforest (but one collection in secondary rainforest) from sea level up to elevations of ¢. 1000 m but extremely rare above 200 m . Food and Feeding. Broad-striped Tube-nosed Fruit Bats are frugivorous and likely a Ficus ( Moraceae ) specialist. They have been observed picking ripe figs after briefly hovering near clusters offigs. Breeding. Pregnant Broad-striped Tube-nosed Fruit Bats have been captured in January—February and a lactating female in September. Activity patterns. Broad-striped Tube-nosed Fruit Bats are nocturnal. Movements, Home range and Social organization. The Broad-striped Tube-nosed Fruit Bat appears to occur at low densities. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Broad-striped Tube-nosed Fruit Bat is widespread and common throughout its distribution. It faces no known major threats, but very little is known about its ecology and threats. Bibliography. Bergmans (2001), Bonaccorso (1998), Bonaccorso & Helgen (2008b), Flannery (1995a, 1995b), Giannini & Simmons (2007a), Helgen (2007a), Irwin (2017), McKean (1972), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Nyctimene aello	Nyctimene		aello	Thomas	1900	1	Ann. Mag. Nat. Hist.	ser. 7, 5: 216	Broad-striped Tube-nosed Fruit Bat	 celaeno Thomas, 1922.	Papua New Guinea, Milne Bay Prov., Milne Bay.	Mainland New Guinea; Kairiru and Admosin Isls (Papua New Guinea); Misool and Salawati Isl (Prov. of Papua, Indonesia).	Not listed.	Least Concern	 aello species group. Includes celaeno, see Flannery (1995a, b) and Bergmans (2001). Also see Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctimene aello	23	Broad-striped Tube-nosed Fruit Bat	Greater Tube-nosed Bat|Dark Tube-nosed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	NYCTIMENINAE	NA	Nyctimene	NA	aello	O. Thomas	1900	1						"Milne Bay [Milne Bay Province], S.E. [Papua] New Guinea.			aello (O. Thomas, 1900)|celaeno O. Thomas, 1922	NA	NA	Indonesia|Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Nyctimene_aello	0	sciname match	Nyctimene_aello	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14954	Nyctimene aello	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Nyctimene	aello	(Thomas, 1900)	The eastern and western distribution on the island of New Guinea might represent subspecies, but this has not been tested (Flannery 1995).	20000000	Nyctimene aello	Least Concern		2021	2020-11-22 00:00:00 UTC	3.1	English	This species is assessed as Least Concern given its wide distribution, use of a broad range of forested habitats, presumed large population size, occurrence in protected areas, and the absence of evidence for a decline.	Nyctimene aello has been recorded from primary and secondary rainforest, swamp forest, and rural gardens. Animals roost in foliage, either singly or in small groups. It is a frugivorous species, likely to be a fig specialist. Pregnant females have been recorded in January and February (Flannery 1995, Bonaccorso 1998).	There appear to be no major threats to this species that could cause widespread or imminent decline, mainly because of the species’ small size, small colony size and broad distribution. It does not appear to be the target of hunters.	There are no population-level data for this relatively common species. It appears to have a low population density but is widespread throughout the island of New Guinea. The species population is assumed to be stable.	Stable	It occurs over most of the island of New Guinea, except for the highland areas of the central cordillera, the Finisterre Range on the Huon Peninsula, and the Arfak Mountains of the Vogelkop Peninsula. It does appear to be distributed in the North Coastal Range. It has been recorded from sea level up to 1,000 m asl. Records also occur on the islands of Misool and Salawati in Indonesia, and Kairiru Island in East Sepik Province, Papua New Guinea.  Identifying this species is more straightforward than some of the smaller species of Nyctimene  because of its larger size and the relative broadness of the dark stripe that extends down the fur on the back. Thus, its distribution is probably better understood than the smaller species that are easier to misidentify.	The species is not known to be hunted, used, or traded.	Terrestrial	It is presumed to be present in most protected areas within its distribution. Broadscale protection of forests will help maintain population size in this species. Additional research into the species ecology, threats, population status and trends will help inform its conservation.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Nyctimene		aello	Thomas	1900	1	Ann. Mag. Nat. Hist.	ser. 7, 5: 216	Broad-striped Tube-nosed Fruit Bat	 celaeno Thomas, 1922.	Papua New Guinea, Milne Bay Prov., Milne Bay.	Mainland New Guinea; Kairiru and Admosin Isls (Papua New Guinea); Misool and Salawati Isl (Prov. of Papua, Indonesia).	Not listed.	Least Concern	 aello species group. Includes celaeno, see Flannery (1995a, b) and Bergmans (2001). Also see Bonaccorso (1998).	Nyctimene aello	1004414	23	Broad-striped Tube-nosed Fruit Bat	Greater Tube-nosed Bat|Dark Tube-nosed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	NYCTIMENINAE	NA	Nyctimene	NA	aello	O. Thomas	1900	1						"Milne Bay [Milne Bay Province], S.E. [Papua] New Guinea.			aello (O. Thomas, 1900)|celaeno O. Thomas, 1922	NA	NA				Indonesia|Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Nyctimene_aello	0	sciname match	Nyctimene_aello	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctimene_aello	1004414	23	Broad-striped Tube-nosed Fruit Bat	Greater Tube-nosed Bat|Dark Tube-nosed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Nyctimeninae	NA	Nyctimene	NA	aello	O. Thomas	1	Cephalotes aello	Thomas, O. 1900-02-01. Description of a new fruit-bat from New Guinea. Annals and Magazine of Natural History (7)5(26):216-217.	https://www.biodiversitylibrary.org/page/24340184	BMNH:Mamm:1899.12.3.1	holotype	https://data.nhm.ac.uk/object/96cbe387-0666-444b-b883-067cc4dcdd43	"Milne Bay [Milne Bay Province], S.E. [Papua] New Guinea.			NA	NA				Indonesia|Papua New Guinea	Oceania (Continent)	Australasia	LC	0	0	0	Nyctimene_aello	0	sciname match	Nyctimene_aello	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Nyctimene		aello	Thomas	1900	1	Ann. Mag. Nat. Hist.	ser. 7, 5: 216	Broad-striped Tube-nosed Fruit Bat	celaeno Thomas, 1922.	Papua New Guinea, Milne Bay Prov., Milne Bay.	Mainland New Guinea; Kairiru and Admosin Isls (Papua New Guinea); Misool and Salawati Isl (Prov. of Papua, Indonesia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14954/22008855/' target='_blank'>Least Concern</a>	aellospecies group. Includes celaeno, see Flannery (1995a, b) and Bergmans (2001). Also see Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctimene aello; Nyctimene aello; Nyctimene aello; Nyctimene aello; Nyctimene aello; Nyctimene aello; celaeno; celaeno; celaeno; aello; celaeno; Grand Nyctimene; Breitstreifen-Réhrennasenflughund; Nyctimeno grande; Greater Tube-nosed Bat; Dark Tube-nosed Fruit Bat (celaeno); Broad-striped Tube-nosed Fruit Bat; Greater Tube-nosed Bat; Dark Tube-nosed Fruit Bat; Broad-striped Tube-nosed Fruit Bat; Broad-striped Tube-nosed Fruit Bat; N. aello