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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1049	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata	Nycteris tragata		[MSW2] Clearly distinct from javanica; see Ellerman and Morrison-Scott (1955:9).; [MSW3] javanica species group. Distinct from javanica; see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993a), but also see Corbet and Hill (1992).; [HMW] Petalia tragata K. Andersen, 1912 , “Bidi Caves, Sarawak ,” Borneo. Nycteris tragata belongs to the javanica group. Monotypic.; [batnames2022]  hispida species group; see Demos et al. (2019). Distinct from javanica; see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993a), but also see Corbet and Hill (1992). For chromosome morphology see Volleth et al. (2019).; [IUCN] This species is often considered to be a subspecies of Nycteris javanica (Corbet and Hill 1992). However, it is recognised as a distinct species by Simmons (2005), also see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993).; [batnames2023]  hispida species group; see Demos et al. (2019). Distinct from javanica; see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993a), but also see Corbet and Hill (1992). For chromosome morphology see Volleth et al. (2019).; [batnames2025_1.7] hispida species group; see Demos et al. (2019). Distinct from javanica; see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993a), but also see Corbet and Hill (1992). For chromosome morphology see Volleth et al. (2019).														tragata	This species is often considered to be a subspecies of Nycteris javanica (Corbet and Hill 1992). However, it is recognised as a distinct species by Simmons (2005), also see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993).			tragata	tragata			tragata (Andersen, 1912)					Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nycteris tragata	Malaysia, Sarawak, Bidi caves.	K. Andersen	1912	Ann. Mag. Nat. Hist., ser. 8, 10:546.	Distribution: Ranging from ex treme southern Burma and Thailand through Malaya and Sumatra to Borneo and perhaps Celebes.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		S Burma – Sumatra, Borneo	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	K. Andersen	1912	Ann. Mag. Nat. Hist., ser. 8, 10:546.	Clearly distinct from javanica; see Ellerman and Morrison-Scott (1955:9).	Burma, Thailand, W Malaysia, Sumatra, Borneo.	Malaysia, Sarawak, Bidi caves.		ANDERSEN	1912	Upper incisors more or less bicuspid. Size fairly large (forearm length, 46-53 mm; condylocanine length, 17-20 mm; maxillary toothrow length, 7.1-8.8 mm).	Distribution: Ranging from ex treme southern Burma and Thailand through Malaya and Sumatra to Borneo and perhaps Celebes.	No subspecies.		49	species	N. tragata	ANDERSEN	1912	Nycteris	genus	Nycteris tragata				Upper incisors more or less bicuspid. Size fairly large (forearm length, 46-53 mm; condylocanine length, 17-20 mm; maxillary toothrow length, 7.1-8.8 mm).	No subspecies.		2. N. tragata (ANDERSEN 1912) [javanica group].	2	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Nycteridae			Nycteris tragata	Nycteris		tragata	K. Andersen	y	1912		Ann. Mag. Nat. Hist., ser. 8	10		546		Malayan Slit-faced Bat	Malaysia, Sarawak, Bidi caves.	Burma, Thailand, W Malaysia, Sumatra, Borneo.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		javanica species group. Distinct from javanica; see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993a), but also see Corbet and Hill (1992).	038D264C754BD70F85B5FCDB5765F77A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Nycteridae_374.pdf.imf	hash://md5/ffb45e34754fd70b851affef527effbd	384	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/8D/26/038D264C754BD70F85B5FCDB5765F77A.xml	Nycteris tragata	Nycteridae	Nycteris	tragata	K. Andersen	1912	Nyctere de Malaisie @fr | Malayische Schlitznase @de | Nicterio malayo @es | Malayan Hollow-faced Bat @en	Petalia tragata K. Andersen, 1912 , “Bidi Caves, Sarawak ,” Borneo. Nycteris tragata belongs to the javanica group. Monotypic.	Patchily distributed in Malay Peninsula, Sumatra, and Borneo.	Head-body 68-75 mm, tail 70-73 mm, car 31-33 mm, hindfoot 10 mm , forearm 46-55 mm; weight 13-22 g. The Malayan Slit-faced Bat is small, with short rounded wings. Longitudinal cleft runs along top of muzzle, covering noseleaves. Fur is long and fluffy, pale reddish brown to grayish brown dorsally, and slightly lighter ventrally. Ears are very long. Wing membranes are dark brown. There are no obvious sexual differences in pelage.	Variety of tropical lowland forests and regenerating forests but apparently not rubber plantations.	No information.	Practically nothing is known about breeding biology of the Malayan Slitfaced Bat, except that pregnant and lactating females have been collected throughout the year, suggesting that breeding is not seasonal. Littersize is one.	Malayan Slit-faced Bats roost during the day mostly in hollow trees and caves, with a single record from an abandoned building. Echolocation call is multiharmonic and involves a steep FM sweep with peak energy at 84 kHz.	Malayan Slit-faced Bats roost singly or in small groups of 2-5 individuals.	Classified as Near Threatened on The IUCN Red List. The Malayan Slit-faced Batis primarily restricted to undisturbed forest and does not enter rubber plantations. Natural lowland forest is being cleared at a rapid rate in this region, and this is cause for conservation concern.	Griffiths (1994, 1997) | Kingston, Francis et al. (2003) | Medway (1983) | Mohd-Hanif et al. (2015) | Phommexay et al. (2011) | Pottie et al. (2005) | Thomas et al. (1994) | Van Cakenberghe & De Vree (1993b)	https://zenodo.org/record/6576952/files/figure.png	5. Malayan Slit-faced Bat Nycteris tragata French: Nyctere de Malaisie / German: Malayische Schlitznase / Spanish: Nicterio malayo Other common names: Malayan Hollow-faced Bat Taxonomy. Petalia tragata K. Andersen, 1912 , “Bidi Caves, Sarawak ,” Borneo. Nycteris tragata belongs to the javanica group. Monotypic. Distribution. Patchily distributed in Malay Peninsula, Sumatra, and Borneo. Descriptive notes. Head-body 68-75 mm, tail 70-73 mm, car 31-33 mm, hindfoot 10 mm , forearm 46-55 mm; weight 13-22 g. The Malayan Slit-faced Bat is small, with short rounded wings. Longitudinal cleft runs along top of muzzle, covering noseleaves. Fur is long and fluffy, pale reddish brown to grayish brown dorsally, and slightly lighter ventrally. Ears are very long. Wing membranes are dark brown. There are no obvious sexual differences in pelage. Habitat. Variety of tropical lowland forests and regenerating forests but apparently not rubber plantations. Food and Feeding. No information. Breeding. Practically nothing is known about breeding biology of the Malayan Slitfaced Bat, except that pregnant and lactating females have been collected throughout the year, suggesting that breeding is not seasonal. Littersize is one. Activity patterns. Malayan Slit-faced Bats roost during the day mostly in hollow trees and caves, with a single record from an abandoned building. Echolocation call is multiharmonic and involves a steep FM sweep with peak energy at 84 kHz. Movements, Home range and Social organization. Malayan Slit-faced Bats roost singly or in small groups of 2-5 individuals. Status and Conservation. Classified as Near Threatened on The IUCN Red List. The Malayan Slit-faced Batis primarily restricted to undisturbed forest and does not enter rubber plantations. Natural lowland forest is being cleared at a rapid rate in this region, and this is cause for conservation concern. Bibliography. Griffiths (1994, 1997), Kingston, Francis et al. (2003), Medway (1983), Mohd-Hanif et al. (2015), Phommexay et al. (2011), Pottie et al. (2005), Thomas et al. (1994), Van Cakenberghe & De Vree (1993b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Nycteridae	Nycteris tragata	Nycteris	Nycteris	tragata	K. Andersen	1912	1	Ann. Mag. Nat. Hist.	ser. 8, 10: 546	Malayan Slit-faced Bat	None.	Malaysia, Sarawak, Bidi caves.	Burma, Thailand, W Malaysia, Sumatra, Borneo.	Not listed.	Near Threatened	 hispida species group; see Demos et al. (2019). Distinct from javanica; see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993a), but also see Corbet and Hill (1992). For chromosome morphology see Volleth et al. (2019).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nycteris tragata	23	Malayan Slit-faced Bat	Malayan Hollow-faced Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	NYCTERIDAE	NA	NA	Nycteris	NA	tragata	K. Andersen	1912	1	Petalia_tragata	Andersen, K. (1912). Brief diagnoses of eight new Petalia, with a list of the known forms of the genus. Annals and Magazine of Natural History, ser. 8, 10, 546.	https://www.biodiversitylibrary.org/item/53333#page/578/mode/1up	BM 1903.3.31.1		"Bidi Caves, Sarawak," Borneo.			tragata (K. Andersen, 1912)	NA	NA	Myanmar|Thailand|Malaysia|Singapore|Indonesia|Brunei	Asia	Indomalaya	NT	0	0	0	Nycteris_tragata	0	sciname match	Nycteris_tragata	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14937	Nycteris tragata	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	NYCTERIDAE	Nycteris	tragata	(K. Andersen, 1912)	This species is often considered to be a subspecies of Nycteris javanica (Corbet and Hill 1992). However, it is recognised as a distinct species by Simmons (2005), also see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993).	20000000	Nycteris tragata	Near Threatened	A3c	2020	2018-08-03 00:00:00 UTC	3.1	English	This species is listed as Near Threatened (nearly meets criterion A3c). Although it is widespread, there is significant deforestation occurring at a rapid rate throughout its range. It seems to be a naturally uncommon species, and its preference for primary forest means it is particularly susceptible to loss of its habitat. Its global population is projected to decline by 25-30% over the next 15 years (three generations; generation length = 5 years, Pacifici et al. 2013) due to habitat loss making it close to qualifying for Vulnerable under criterion A2c.	the species has been collected from a wide variety of forests including primary dipterocarp forest, secondary forests, kerangas, limestone areas and peat swamp forest (Anwarali et al.  2008; Soisook 2011; Jayaraj et al.  2011, 2013; Huang et al.  2014). It roosts in small groups in fallen tree hollows and similar man-made hollows such as culverts as well as crevices of large boulders and in limestone caves.	The major threat for this species is habitat loss due to deforestation for logging, plantations, agriculture, as well forest fire.	Nycteris tragata is a naturally uncommon species, and its preference for primary forest means it is particularly susceptible to loss of its habitat. Its global population is projected to decline by 25-30% over the next 15 years (three generations; generation length = 5 years, Pacifici ;et al. ;2013) ;due to forest loss and degradation.	Decreasing	This species' distribution includes Myanmar, Thailand, Peninsular Malaysia, Sumatra (Indonesia) and throughout Borneo (Simmons 2005, Shazali et al. 2018).The type locality for this species is Bidi caves, Sarawak, Borneo Malaysia. In Sabah this species is found in Bohay Dulang, Semporna, Sepilok, Tawau HP, Monggis Sub Station.		Terrestrial	The species range includes several protected areas.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Nycteridae	Nycteris	Nycteris	tragata	K. Andersen	1912	1	Ann. Mag. Nat. Hist.	ser. 8, 10: 546	Malayan Slit-faced Bat	None.	Malaysia, Sarawak, Bidi caves.	Burma, Thailand, W Malaysia, Sumatra, Borneo.	Not listed.	Near Threatened	 hispida species group; see Demos et al. (2019). Distinct from javanica; see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993a), but also see Corbet and Hill (1992). For chromosome morphology see Volleth et al. (2019).	Nycteris tragata	1004846	23	Malayan Slit-faced Bat	Malayan Hollow-faced Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Nycteridae	NA	NA	Nycteris	NA	tragata	K. Andersen	1912	1	Petalia_tragata	Andersen, K. (1912). Brief diagnoses of eight new Petalia, with a list of the known forms of the genus. Annals and Magazine of Natural History, ser. 8, 10, 546.	https://www.biodiversitylibrary.org/item/53333#page/578/mode/1up	BM 1903.3.31.1		"Bidi Caves, Sarawak," Borneo.			tragata (K. Andersen, 1912)	NA	NA				Myanmar|Thailand|Malaysia|Singapore|Indonesia|Brunei	Asia	Indomalaya	NT	0	0	0	Nycteris_tragata	0	sciname match	Nycteris_tragata	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nycteris_tragata	1004846	23	Malayan Slit-faced Bat	Malayan Hollow-faced Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Nycteridae	NA	NA	Nycteris	NA	tragata	Andersen	1	Petalia tragata	Andersen, K.C. 1912-11-01. Brief diagnoses of eight new _Petalia_, with a list of the known forms of the genus. Annals and Magazine of Natural History (8)10(59):546-550.	https://www.biodiversitylibrary.org/page/18640617	BMNH:Mamm:1903.3.31.1	holotype	https://data.nhm.ac.uk/object/298de46b-fa58-4749-85c5-f766180e61f6	"Bidi Caves, Sarawak," Borneo.			NA	NA				Myanmar|Thailand|Malaysia|Singapore|Indonesia|Brunei	Asia	Indomalaya	NT	0	0	0	Nycteris_tragata	0	sciname match	Nycteris_tragata	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Nycteridae	Nycteris	Nycteris	tragata	K. Andersen	1912	1	Ann. Mag. Nat. Hist.	ser. 8, 10: 546	Malayan Slit-faced Bat	None.	Malaysia, Sarawak, Bidi caves.	Burma, Thailand, W Malaysia, Sumatra, Borneo.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14937/22014643/' target='_blank'>Near Threatened</a>	hispida species group; see Demos et al. (2019). Distinct from javanica; see Ellerman and Morrison-Scott (1955) and Van Cakenberghe and De Vree (1993a), but also see Corbet and Hill (1992). For chromosome morphology see Volleth et al. (2019).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nycteris tragata; Nycteris tragata; Nycteris tragata; Nycteris tragata; Nycteris tragata; Nycteris tragata; tragata; Nyctere de Malaisie; Malayische Schlitznase; Nicterio malayo; Malayan Hollow-faced Bat; Malayan Slit-faced Bat; Malayan Hollow-faced Bat; Malayan Slit-faced Bat; Malayan Slit-faced Bat; N. tragata