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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1038	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis	Nycteris grandis		[MSW3] hispida species group. Reviewed by Van Cakenberghe and De Vree (1993b). N. marica is sometimes recognized as a distinct savanna subspecies, but this does not seem justified based on morphology; see Van Cakenberghe and De Vree (1993b). See Hickey and Dunlop (2000).; [HMW] Nycteris grandis Peters, 1865 , “ Guinea .” Nycteris grandis belongs to the hispida group. Taxonomic relationships between forestand savanna-dwelling populations need investigation. Monotypic.; [batnames2022]  hispida species group. Reviewed by Van Cakenberghe and De Vree (1993b). N. marica is sometimes recognized as a distinct savanna subspecies, but this does not seem justified based on morphology; see Van Cakenberghe and De Vree (1993b). See Hickey and Dunlop (2000).; [IUCN]  N. marica Kershaw, 1923, is the available name for the southern savanna species if it is recognised as distinct from this species.; [batnames2023]  hispida species group. Reviewed by Van Cakenberghe and De Vree (1993b). N. marica is sometimes recognized as a distinct savanna subspecies, but this does not seem justified based on morphology; see Van Cakenberghe and De Vree (1993b). See Hickey and Dunlop (2000).; [batnames2025_1.7] hispida species group. Reviewed by Van Cakenberghe and De Vree (1993b). N. marica is sometimes recognized as a distinct savanna subspecies, but this does not seem justified based on morphology; see Van Cakenberghe and De Vree (1993b). See Hickey and Dunlop (2000).						baikii, marica, proxima.	marica, grandis		baikii, marica, proxima			grandis 	grandis - baikii, marica, proxima	grandis, baikii, marica, proxima	 N. marica Kershaw, 1923, is the available name for the southern savanna species if it is recognised as distinct from this species.	grandis 	grandis - baikii, marica, proxima	grandis, baikii, marica, proxima	grandis, baikii, marica, proxima	grandis 	grandis - baikii, marica, proxima	grandis W. C. H. Peters, 1865|baikii W. C. H. Peters, 1867 [nomen nudum | not used as valid]|marica P. S. Kershaw, 1923|proxima LÃ¶nnberg & Gyldenstolpe, 1925		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Large slit-faced bat	Guinea – Tanzania, Mozambique, Rhodesia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nycteris grandis	"Guinea".	Peters	1865	Monatsb. Preuss. Akad. Wiss. Berlin, p. 358.	Distribution: Confined to forest and woodland areas of tropical Africa from Senegal to Kenya and south to Zimbabwe and Mozambique, including Pemba and Zanzibar is lands off the east coast.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Large slit-faced bat	Senegal – Kenya, Tanzania, Mozambique, Zimbabwe; ref. 4.29	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1865	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1865:358.		Senegal to Kenya, Zimbabwe, and Mozambique; Zanzibar and Pemba.	"Guinea".		PETERS	1871	Pos terior lower premolar reduced, its crown rising scarcely above cingula of anterior lower premolar and first lower molar. Upper incisors definitely tricuspid. Tragus more or less semilunate with no well marked concavity in its anterior margin. Size large (forearm length, 51-66 mm; condylocanine length, 20-25 mm; maxillary toothrow length, 7.9-9.7 mm).	Distribution: Confined to forest and woodland areas of tropical Africa from Senegal to Kenya and south to Zimbabwe and Mozambique, including Pemba and Zanzibar is lands off the east coast.	Two subspecies:	N. g. marica (southeastern Kenya south to Zimbabwe and Mozambique, including Pemba and Zanzibar). N. g. grandis (= proxima) (remaining range).	49	species	N. grandis	PETERS	1871	Nycteris	genus	Nycteris grandis				Pos terior lower premolar reduced, its crown rising scarcely above cingula of anterior lower premolar and first lower molar. Upper incisors definitely tricuspid. Tragus more or less semilunate with no well marked concavity in its anterior margin. Size large (forearm length, 51-66 mm; condylocanine length, 20-25 mm; maxillary toothrow length, 7.9-9.7 mm).	Two subspecies:		7. N. grandis PETERS 1871 [hispida group].	7	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Nycteridae			Nycteris grandis	Nycteris		grandis	Peters		1865		Monatsb. K. Preuss. Akad. Wiss. Berlin	1865		358		Large Slit-faced Bat	"Guinea".	Senegal to Dem. Rep. Congo, Kenya, Zimbabwe, Malawi, and Mozambique; Zanzibar and Pemba.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	baikii Gray, 1866; marica Kershaw, 1923; proxima Lönnberg and Gyldenstolpe, 1925.	hispida species group. Reviewed by Van Cakenberghe and De Vree (1993b). N. marica is sometimes recognized as a distinct savanna subspecies, but this does not seem justified based on morphology; see Van Cakenberghe and De Vree (1993b). See Hickey and Dunlop (2000).	038D264C754BD70E80B3F9A3531CFE13	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Nycteridae_374.pdf.imf	hash://md5/ffb45e34754fd70b851affef527effbd	384	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/8D/26/038D264C754BD70E80B3F9A3531CFE13.xml	Nycteris grandis	Nycteridae	Nycteris	grandis	Peters	1865	Grand Nyctere @fr | Grof Re Schlitznase @de | Nicterio grande @es	Nycteris grandis Peters, 1865 , “ Guinea .” Nycteris grandis belongs to the hispida group. Taxonomic relationships between forestand savanna-dwelling populations need investigation. Monotypic.	Widely distributed in tropical forest belt of Africa extending from Senegal in W to E DR Congo in the FE; there is a separate population restricted to coastal Kenya , Tanzania (including Pemba and Unguja Is in Zanzibar Archipelago), and NE Mozambique , with scattered records from Zambia , S Malawi , N Zimbabwe , and WC Mozambique .	Head-body 70-90 mm, tail 60-84 mm, ear 24-35 mm, hindfoot 14-17 mm, forearm 52-65 mm; weight 21-43 g. The Large Slit-faced Bat is mediumsized, with short rounded wings. Longitudinal cleft runs along top of muzzle, covering noseleaves. Fur is long and fluffy, dark reddish brown to dark brown (in forest) or grayish brown (in savanna) dorsally, and slightly lighter ventrally. Ears are very long but relatively short for a slitfaced bat. Wing membranes are dark brown and free of hairs except along margins of forelimbs. There are no obvious sexual differences in pelage.	Variety of tropical lowland rainforest habitats including swamp, secondary, and riparian forests. In East Africa, the Large Slit-faced Bat is restricted to coastal forests, and in southern Africa, it occurs in riverine forests in woodlands.	The Large Slit-faced Bat is the only carnivorous bat in Africa, feeding on various vertebrates including small mammals (other species of bats) and birds but mainly frogs and fishes (Ptychadena and Tilapia). It also preys on various invertebrates, particularly katydids and crickets. During one study, each bat consumed on average of two frogs per night.	Pregnant and lactating Large Slit-faced Bats have been captured at various times of the year in West and Central Africa, but it is not known if breeding is seasonal. In southern Africa, it would appear that young are born during the wet season. Litter size 1s one.	The Large Slit-faced Bat roosts during the day in sheltered structures including hollow trees, caves, and road culverts. It leaves day roost after sunset. In woodland habitat, it appears to have two distinct foraging strategies: perch-hunting and gleaning prey in flight. It regularly returns to the same dayroost.	Social organization of the Large Slit-faced Bat appears to differ in forests and woodlands; in forests,it typically roosts singly or in pairs, but in woodlands, it can occur in groups of up to several individuals. Individuals forage more than 2 km from roosting sites. Echolocation call is multiharmonic and involves a steep FM sweep.	Classified as Least Concern on The IUCN Red List.	Ansell (1986) | Brosset (1966b) | Cockle et al. (1998) | Fenton, Cumming et al. (1987) | Fenton, Gaudet & Leonard (1983) | Fenton, Rautenbach et al. (1993) | Fenton, Swanepoel et al. (1990) | Fenton, Thomas & Sasseen (1981) | Griffiths (1994, 1997) | Happold, M. (2013ae) | Hickey & Dunlop (2000) | Kingdon (1974) | Monadjem, Richards & Denys (2016) | Monadjem, Schoeman et al. (2010) | Monadjem, Taylor et al. (2010) | Rosevear (1965) | Smithers & Wilson (1979) | Thomas et al. (1994) | Van Cakenberghe & De Vree (1993a) | Wolton et al. (1982)	https://zenodo.org/record/6576960/files/figure.png	8. Large Slitfaced Bat Nycteris grandis French: Grand Nyctere / German: GrofRe Schlitznase / Spanish: Nicterio grande Taxonomy. Nycteris grandis Peters, 1865 , “ Guinea .” Nycteris grandis belongs to the hispida group. Taxonomic relationships between forestand savanna-dwelling populations need investigation. Monotypic. Distribution. Widely distributed in tropical forest belt of Africa extending from Senegal in W to E DR Congo in the FE; there is a separate population restricted to coastal Kenya , Tanzania (including Pemba and Unguja Is in Zanzibar Archipelago), and NE Mozambique , with scattered records from Zambia , S Malawi , N Zimbabwe , and WC Mozambique . Descriptive notes. Head-body 70-90 mm, tail 60-84 mm, ear 24-35 mm, hindfoot 14-17 mm, forearm 52-65 mm; weight 21-43 g. The Large Slit-faced Bat is mediumsized, with short rounded wings. Longitudinal cleft runs along top of muzzle, covering noseleaves. Fur is long and fluffy, dark reddish brown to dark brown (in forest) or grayish brown (in savanna) dorsally, and slightly lighter ventrally. Ears are very long but relatively short for a slitfaced bat. Wing membranes are dark brown and free of hairs except along margins of forelimbs. There are no obvious sexual differences in pelage. Habitat. Variety of tropical lowland rainforest habitats including swamp, secondary, and riparian forests. In East Africa, the Large Slit-faced Bat is restricted to coastal forests, and in southern Africa, it occurs in riverine forests in woodlands. Food and Feeding. The Large Slit-faced Bat is the only carnivorous bat in Africa, feeding on various vertebrates including small mammals (other species of bats) and birds but mainly frogs and fishes (Ptychadena and Tilapia). It also preys on various invertebrates, particularly katydids and crickets. During one study, each bat consumed on average of two frogs per night. Breeding. Pregnant and lactating Large Slit-faced Bats have been captured at various times of the year in West and Central Africa, but it is not known if breeding is seasonal. In southern Africa, it would appear that young are born during the wet season. Litter size 1s one. Activity patterns. The Large Slit-faced Bat roosts during the day in sheltered structures including hollow trees, caves, and road culverts. It leaves day roost after sunset. In woodland habitat, it appears to have two distinct foraging strategies: perch-hunting and gleaning prey in flight. It regularly returns to the same dayroost. Movements, Home range and Social organization. Social organization of the Large Slit-faced Bat appears to differ in forests and woodlands; in forests,it typically roosts singly or in pairs, but in woodlands, it can occur in groups of up to several individuals. Individuals forage more than 2 km from roosting sites. Echolocation call is multiharmonic and involves a steep FM sweep. Status and Conservation. Classified as Least Concern on The IUCN Red List. Bibliography. Ansell (1986), Brosset (1966b), Cockle et al. (1998), Fenton, Cumming et al. (1987), Fenton, Gaudet & Leonard (1983), Fenton, Rautenbach et al. (1993), Fenton, Swanepoel et al. (1990), Fenton, Thomas & Sasseen (1981), Griffiths (1994, 1997), Happold, M. (2013ae), Hickey & Dunlop (2000), Kingdon (1974), Monadjem, Richards & Denys (2016), Monadjem, Schoeman et al. (2010), Monadjem, Taylor et al. (2010), Rosevear (1965), Smithers & Wilson (1979), Thomas et al. (1994), Van Cakenberghe & De Vree (1993a), Wolton et al. (1982).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Nycteridae	Nycteris grandis	Nycteris	Nycteris	grandis	Peters	1865	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1870:58:00	Large Slit-faced Bat	 baikii Gray, 1866; marica Kershaw, 1923; proxima L&ouml;nnberg and Gyldenstolpe, 1925.	"Guinea".	Senegal to Dem. Rep. Congo, Kenya, Zimbabwe, Malawi, and Mozambique; Zanzibar and Pemba.	Not listed.	Least Concern	 hispida species group. Reviewed by Van Cakenberghe and De Vree (1993b). N. marica is sometimes recognized as a distinct savanna subspecies, but this does not seem justified based on morphology; see Van Cakenberghe and De Vree (1993b). See Hickey and Dunlop (2000).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nycteris grandis	23	Large Slit-faced Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	NYCTERIDAE	NA	NA	Nycteris	NA	grandis	W. Peters	1865	0	Nycteris_grandis	Peters, W. C. H. (1865). Ãœber Flederthiere (Vespertilio soricinus Pallas, Choeronycteris Lichtenst., Rhinophylla pumilio nov. gen., Artibeus fallax nov. sp., A. concolor nov. sp., Dermanura quadrivittatum nov. sp., Nycteris grandis n. sp.). Monatsberichte der KÃ¶niglichen Preussische Akademie des Wissenschaften zu Berlin, 1865, 358.	https://www.biodiversitylibrary.org/item/109287#page/376/mode/1up	RMNH MAM.27348		"Guinea."			grandis W. Peters, 1865|baikii J. E. Gray, 1866|marica Kershaw, 1923|proxima LÃ¶nnberg & Gyldenstolpe, 1925	NA	NA	Senegal|Guinea-Bissau|Guinea|Sierra Leone|Liberia|CÃ´te d'Ivoire|Ghana|Togo|Benin|Nigeria|Cameroon|Equatorial Guinea|Central African Republic|Gabon|Republic of the Congo|Democratic Republic of the Congo|South Sudan|Kenya|Tanzania|Malawi|Zambia|Mozambique|Zimbabwe	Africa	Afrotropic	LC	0	0	0	Nycteris_grandis	0	sciname match	Nycteris_grandis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14929	Nycteris grandis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	NYCTERIDAE	Nycteris	grandis	Peters, 1865	 N. marica Kershaw, 1923, is the available name for the southern savanna species if it is recognised as distinct from this species.	20000000	Nycteris grandis	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species has been recorded from a variety of lowland habitats, ranging from lowland tropical moist forest (often found near to swampy sites) to drier savanna areas and Miombo woodland (Rosevear 1965; Fenton et al. 1990; Hickey and Dunlop 2000; Skinner and Chimimba 2005). It generally roosts in hollow trees, but is also found in man made structures such as houses (Fenton et al. 1990, 1993), disused water towers (Fenton et al. 1990, 1993) and culverts (Rosevear, 1965). Animals might also roost in hollow fallen logs, and holes or small caverns in rocks (Rosevear, 1965; Hickey and Dunlop 2000).	In general there appear to be no major threats to this species as a whole. It is threatened in parts of its range by habitat loss, particularly the logging of large trees used for roosting. Some populations may be threatened by overharvesting for subsistence food.	It appears to be a rare, or rarely recorded, species. The species occurs in small colonies, but is usually found only as single animals or in pairs.	Decreasing	This species is broadly distributed in sub-Saharan Africa. It ranges from Senegal, through West and Central Africa, to southern Sudan, southeastern Kenya and eastern Tanzania, with scattered records as far south as Zambia, Zimbabwe and Mozambique. It is a lowland species.		Terrestrial	In view of the species wide range, it is presumably present in a number of protected areas. No direct conservation measures are currently needed for this species as a whole.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Nycteridae	Nycteris	Nycteris	grandis	Peters	1865	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1870:58:00	Large Slit-faced Bat	 baikii Gray, 1866; marica Kershaw, 1923; proxima L&ouml;nnberg and Gyldenstolpe, 1925.	"Guinea".	Senegal to Dem. Rep. Congo, Kenya, Zimbabwe, Malawi, and Mozambique; Zanzibar and Pemba.	Not listed.	Least Concern	 hispida species group. Reviewed by Van Cakenberghe and De Vree (1993b). N. marica is sometimes recognized as a distinct savanna subspecies, but this does not seem justified based on morphology; see Van Cakenberghe and De Vree (1993b). See Hickey and Dunlop (2000).	Nycteris grandis	1004836	23	Large Slit-faced Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Nycteridae	NA	NA	Nycteris	NA	grandis	W. Peters	1865	0	Nycteris_grandis	Peters, W. C. H. (1865). Ãœber Flederthiere (Vespertilio soricinus Pallas, Choeronycteris Lichtenst., Rhinophylla pumilio nov. gen., Artibeus fallax nov. sp., A. concolor nov. sp., Dermanura quadrivittatum nov. sp., Nycteris grandis n. sp.). Monatsberichte der KÃ¶niglichen Preussische Akademie des Wissenschaften zu Berlin, 1865, 358.	https://www.biodiversitylibrary.org/item/109287#page/376/mode/1up	RMNH MAM.27348		"Guinea."			grandis W. Peters, 1865|baikii J. E. Gray, 1866|marica Kershaw, 1923|proxima LÃ¶nnberg & Gyldenstolpe, 1925	NA	NA				Senegal|Guinea-Bissau|Guinea|Sierra Leone|Liberia|CÃ´te d'Ivoire|Ghana|Togo|Benin|Nigeria|Cameroon|Equatorial Guinea|Central African Republic|Gabon|Republic of the Congo|Democratic Republic of the Congo|South Sudan|Kenya|Tanzania|Malawi|Zambia|Mozambique|Zimbabwe	Africa	Afrotropic	LC	0	0	0	Nycteris_grandis	0	sciname match	Nycteris_grandis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nycteris_grandis	1004836	23	Large Slit-faced Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Nycteridae	NA	NA	Nycteris	NA	grandis	W. C. H. Peters	0	Nycteris grandis	Peters, W.C.H. 1865. Ãœber Flederthiere (_Vespertilio soricinus_ Pallas, _Choeronycteris_ Lichtenst., _Rhinophylla pumilio_ nov. gen., _Artibeus fallax_ nov. sp., _A. concolor_ nov. sp., _Dermanura quadrivittatum_ nov. sp., _Nycteris grandis_ n. sp.). Monatsberichte der KÃ¶niglichen Preussischen Akademie der Wissenschaften zu Berlin 1865:351-359.	https://www.biodiversitylibrary.org/page/35168801	RMNH.MAM.27348	holotype	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.27348	"Guinea."			NA	NA				Senegal|Guinea-Bissau|Guinea|Sierra Leone|Liberia|Cote d'Ivoire|Ghana|Togo|Benin|Nigeria|Cameroon|Equatorial Guinea|Central African Republic|Gabon|Republic of the Congo|Democratic Republic of the Congo|South Sudan|Kenya|Tanzania|Malawi|Zambia|Mozambique|Zimbabwe	Africa	Afrotropic	LC	0	0	0	Nycteris_grandis	0	sciname match	Nycteris_grandis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Nycteridae	Nycteris	Nycteris	grandis	Peters	1865	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1870:58:00	Large Slit-faced Bat	baikii Gray, 1866; marica Kershaw, 1923; proxima L&ouml;nnberg and Gyldenstolpe, 1925.	"Guinea".	Senegal to Dem. Rep. Congo, Kenya, Zimbabwe, Malawi, and Mozambique; Zanzibar and Pemba.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14929/22012638/' target='_blank'>Least Concern</a>	hispida species group. Reviewed by Van Cakenberghe and De Vree (1993b). N. marica is sometimes recognized as a distinct savanna subspecies, but this does not seem justified based on morphology; see Van Cakenberghe and De Vree (1993b). See Hickey and Dunlop (2000).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nycteris grandis; Nycteris grandis; Nycteris grandis; Nycteris grandis; Nycteris grandis; Nycteris grandis; baikii; marica; proxima; baikii; marica; proxima; grandis; baikii; marica; proxima; Grand Nyctere; Grof Re Schlitznase; Nicterio grande; Large Slit-faced Bat; Large Slit-faced Bat; Large Slit-faced Bat; N. grandis
