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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1032	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus	Nyctalus montanus		[MSW2] Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951:159); but see Corbet (1978c:55).; [MSW3] Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951), but see Gaisler (1970), Corbet (1978c), Corbet and Hill (1992), and Bates and Harrison (1997).; [HMW] Plerygistes montanus Barrett-Hamilton, 1906 , “Mussooree, Northwestern Himalayas [Dehra Dun, Uttar Pradesh , India ], at an altitude of 4500 to 5500 feet [= 1372 to 1676 m ].” The phylogenetic affiliations of N. montanus are uncertain but morphologically it 1s most similar to N. leisler :, in which it has been included as a subspecies. Monotypic.; [batnames2022] Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951), but see Gaisler (1970), Corbet (1978 c ), Corbet andHill (1992), and Bates and Harrison (1997).; [IUCN] Earlier included under Nyctalus leisleri (Kuhl, 1817) (Ellerman and Morrison-Scott 1951), now considered distinct (Bates and Harrison 1997, Simmons 2005). Recently reviewed by Benda and Gaisler (2015).; [batnames2023] Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951), but see Gaisler (1970), Corbet (1978 c ), Corbet andHill (1992), and Bates and Harrison (1997).; [batnames2025_1.7] Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951), but see Gaisler (1970), Corbet (1978c), Corbet andHill (1992), and Bates and Harrison (1997).														montanus	Earlier included under Nyctalus leisleri (Kuhl, 1817) (Ellerman and Morrison-Scott 1951), now considered distinct (Bates and Harrison 1997, Simmons 2005). Recently reviewed by Benda and Gaisler (2015).			montanus	montanus			montanus (Barrett-Hamilton, 1906)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		E Afghanistan – N India	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nyctalus montanus	India, Uttar Pradesh, Dehra Dun, Mussooree.	Barrett-Hamilton	1906	Ann. Mag. Nat. Hist., ser. 7, 17:99.	Distribution: Ranging from Afghanistan to northern India.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		E Afghanistan – N India	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Barrett-Hamilton	1906	Ann. Mag. Nat. Hist., ser. 7, 17:99.	Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951:159); but see Corbet (1978c:55).	Afghanistan, Pakistan, N India, Nepal.	India, Uttar Pradesh, Dehra Dun, Mussooree.		BARRETT-HAMILTON	1906	Size fairly small (forearm length, 42-44 mm). Anterior upper premolar displaced medially and greatly reduced.	Distribution: Ranging from Afghanistan to northern India.	No subspecies.		118	species	N. montanus	BARRETT-HAMILTON	1906	Nyctalus	genus	Nyctalus montanus				Size fairly small (forearm length, 42-44 mm). Anterior upper premolar displaced medially and greatly reduced.	No subspecies.		5. N. montanus (BARRETT-HAMILTON 1906) [noctula group].	5	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Nyctalus montanus	Nyctalus		montanus	Barrett-Hamilton	y	1906		Ann. Mag. Nat. Hist., ser. 7	17		99		Mountain Noctule	India, Uttar Pradesh, Dehra Dun, Mussooree.	E Afghanistan, Pakistan, N India, Nepal.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951), but see Gaisler (1970), Corbet (1978c), Corbet and Hill (1992), and Bates and Harrison (1997).	4C3D87E8FFFC6A4CFA979B421CA0BCB9	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	765	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFFC6A4CFA979B421CA0BCB9.xml	Nyctalus montanus	Vespertilionidae	Nyctalus	montanus	Barrett-Hamilton	1906	Noctule des montagnes @fr | Bergabendsegler @de | Noctulomontano @es | Himalayan Noctule @en	Plerygistes montanus Barrett-Hamilton, 1906 , “Mussooree, Northwestern Himalayas [Dehra Dun, Uttar Pradesh , India ], at an altitude of 4500 to 5500 feet [= 1372 to 1676 m ].” The phylogenetic affiliations of N. montanus are uncertain but morphologically it 1s most similar to N. leisler :, in which it has been included as a subspecies. Monotypic.	WC Afghanistan ( Nangarhar and Paktika provinces) and along the S Himalayas of N India ( Himachal Pradesh and Uttarakhand ) and W Nepal ; it apparently occurs in N Pakistan , but this requires confirmation.	Head-body ¢.60-70 mm, tail ¢. 43 mm , ear c.12-14 mm, forearm 42-5-44 mm. The Mountain Noctule is very similar to Leisler’s Noctule (N. leisleri ). Dorsal pelage is uniformly brown, with ventral pelage very slightly lighter; individual hairs are bicolored, being brown throughout except base, which is black. Face, ears, wings, and uropatagium are darkly pigmented and mostly naked. Ears are short and triangular, with 4-5 folds on outer edge; tragus is club-shaped and expanded distally, being short and stubby with a rounded tip. Muzzle is short with large glands between nostrils and eyes. Tail extends to a few millimeters beyond uropatagium. Calcaris well developed with a postcalcarial lobe divided by a visible T-piece,as in all other Nyctalus . Skull is robust, being larger and stouter than in Leisler’s Noctule; lambdoid crests are relatively developed but sagittal crest is not; P* is very small, about one-quarter the size of that of Leisler’s Noctule; lower molars are nyctalodont.	Occurs primarily in riparian habitats and arid floodplains, and has been recorded from elevations of 680-2100 m along the southern edge of the Himalayas.	The Mountain Noctule is insectivorous but there are apparent records of the species feeding on fish near riverbanks.	No information.	Mountain Noctules roost around rocky cliffs and crevices, in rocks and overhanging vegetation. One individual was recorded roosting in the roof of a bungalow in Shimla, northern India .	No information. Mountain Noctules probably roost like other species of Nyctalus in smallto medium-sized colonies.	Classified as Least Concern on The IUCN Red List. The Mountain Noctule has a relatively restricted distribution along the Himalayas that may be threatened by deforestation due to logging and agricultural expansion, as well as mining activities, and hunting for medicinal purposes.	Bates & Harrison (1997) | Benda & Gaisler (2015) | Corbet (1978) | Corbet & Hill (1992) | Gaisler (1970) | Molur & Srinivasulu (2008c) | Molur et al. (2002) | Roberts (1997) | Saikia et al. (2011) | Srinivasulu, C. & Srinivasulu (2012) | Srinivasulu, C. et al. (2010)	https://zenodo.org/record/6397792/files/figure.png	13. Mountain Noctule Nyctalus montanus French: Noctule des montagnes / German: Bergabendsegler / Spanish: Noctulo montano Other common names: Himalayan Noctule Taxonomy. Plerygistes montanus Barrett-Hamilton, 1906 , “Mussooree, Northwestern Himalayas [Dehra Dun, Uttar Pradesh , India ], at an altitude of 4500 to 5500 feet [= 1372 to 1676 m ].” The phylogenetic affiliations of N. montanus are uncertain but morphologically it 1s most similar to N. leisler :, in which it has been included as a subspecies. Monotypic. Distribution. WC Afghanistan ( Nangarhar and Paktika provinces) and along the S Himalayas of N India ( Himachal Pradesh and Uttarakhand ) and W Nepal ; it apparently occurs in N Pakistan , but this requires confirmation. Descriptive notes. Head-body ¢.60-70 mm, tail ¢. 43 mm , ear c.12-14 mm, forearm 42-5-44 mm. The Mountain Noctule is very similar to Leisler’s Noctule (N. leisleri ). Dorsal pelage is uniformly brown, with ventral pelage very slightly lighter; individual hairs are bicolored, being brown throughout except base, which is black. Face, ears, wings, and uropatagium are darkly pigmented and mostly naked. Ears are short and triangular, with 4-5 folds on outer edge; tragus is club-shaped and expanded distally, being short and stubby with a rounded tip. Muzzle is short with large glands between nostrils and eyes. Tail extends to a few millimeters beyond uropatagium. Calcaris well developed with a postcalcarial lobe divided by a visible T-piece,as in all other Nyctalus . Skull is robust, being larger and stouter than in Leisler’s Noctule; lambdoid crests are relatively developed but sagittal crest is not; P* is very small, about one-quarter the size of that of Leisler’s Noctule; lower molars are nyctalodont. Habitat. Occurs primarily in riparian habitats and arid floodplains, and has been recorded from elevations of 680-2100 m along the southern edge of the Himalayas. Food and Feeding. The Mountain Noctule is insectivorous but there are apparent records of the species feeding on fish near riverbanks. Breeding. No information. Activity patterns. Mountain Noctules roost around rocky cliffs and crevices, in rocks and overhanging vegetation. One individual was recorded roosting in the roof of a bungalow in Shimla, northern India . Movements, Home range and Social organization. No information. Mountain Noctules probably roost like other species of Nyctalus in smallto medium-sized colonies. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Mountain Noctule has a relatively restricted distribution along the Himalayas that may be threatened by deforestation due to logging and agricultural expansion, as well as mining activities, and hunting for medicinal purposes. Bibliography. Bates & Harrison (1997), Benda & Gaisler (2015), Corbet (1978), Corbet & Hill (1992), Gaisler (1970), Molur & Srinivasulu (2008c), Molur et al. (2002), Roberts (1997), Saikia et al. (2011), Srinivasulu, C. & Srinivasulu (2012), Srinivasulu, C. et al. (2010).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Nyctalus montanus	Nyctalus		montanus	Barrett-Hamilton	1906	1	Ann. Mag. Nat. Hist.	ser. 7, 17: 99	Mountain Noctule	None.	India, Uttar Pradesh, Dehra Dun, Mussooree.	E Afghanistan, Pakistan, N India, Nepal.	Not listed.	Least Concern	Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951), but see Gaisler (1970), Corbet (1978 c ), Corbet andHill (1992), and Bates and Harrison (1997).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctalus montanus	23	Mountain Noctule	Himalayan Noctule	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Nyctalus	NA	montanus	Barrett-Hamilton	1906	1						"Mussooree, North-western Himalayas [Dehra Dun, Uttar Pradesh, India], at an altitude of 4500 to 5500 feet [= 1372 to 1676 m]."			montanus (Barrett-Hamilton, 1906)	NA	NA	Afghanistan|Pakistan?|India|Nepal	Asia	Palearctic	LC	0	0	0	Nyctalus_montanus	0	sciname match	Nyctalus_montanus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14923	Nyctalus montanus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Nyctalus	montanus	(Barrett-Hamilton, 1906)	Earlier included under Nyctalus leisleri (Kuhl, 1817) (Ellerman and Morrison-Scott 1951), now considered distinct (Bates and Harrison 1997, Simmons 2005). Recently reviewed by Benda and Gaisler (2015).	20000000	Nyctalus montanus	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species is found in riparian habitat and arid flood plains with rich undergrowth in the Himalaya. It roosts among rocky cliffs, crevices among rocks and overhanging vegetation (Molur et al. 2002).	This species is threatened by deforestation, generally resulting from logging operations and the conversion of land for agricultural use, mining and quarrying activities. It is also threatened due to hunting for medicinal purposes (Molur et al. 2002).	The abundance, population size and trends for this species are not known (Molur et al. 2002).	Decreasing	This species is endemic to South Asia, where it is presently known from Afghanistan (Nangarhar and Paktika provinces), India (Himachal Pradesh and Uttarakhand), Nepal (Midwestern Nepal) (Srinivasulu and Srinivasulu 2012, Benda and Gaisler 2015). The specimens reported from Pakistan by Roberts (1977) are of N. leisleri. No records of N. montanus is available from Pakistan (Benda and Gaisler 2015). It has been recorded at an altitudinal range of 680 to 1,692 m asl.		Terrestrial	There are no direct conservation measures in place for this species. The species has not been recorded from any protected areas. Detailed surveys, ecological studies, population and habitat monitoring, protection against poaching are the main recommendations (Molur et al . 2002).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Nyctalus		montanus	Barrett-Hamilton	1906	1	Ann. Mag. Nat. Hist.	ser. 7, 17: 99	Mountain Noctule	None.	India, Uttar Pradesh, Dehra Dun, Mussooree.	E Afghanistan, Pakistan, N India, Nepal.	Not listed.	Least Concern	Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951), but see Gaisler (1970), Corbet (1978 c ), Corbet andHill (1992), and Bates and Harrison (1997).	Nyctalus montanus	1005606	23	Mountain Noctule	Himalayan Noctule	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Nyctalus	NA	montanus	Barrett-Hamilton	1906	1						"Mussooree, North-western Himalayas [Dehra Dun, Uttar Pradesh, India], at an altitude of 4500 to 5500 feet [= 1372 to 1676 m]."			montanus (Barrett-Hamilton, 1906)	NA	NA				Afghanistan|Pakistan?|India|Nepal	Asia	Palearctic	LC	0	0	0	Nyctalus_montanus	0	sciname match	Nyctalus_montanus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctalus_montanus	1005606	23	Mountain Noctule	Himalayan Noctule	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Nyctalus	NA	montanus	Barrett-Hamilton	1	Pterygistes montanus	Barrett-Hamilton, G.E.H. 1906-01-01. Descriptions of two new species of _Pterygistes_. Annals and Magazine of Natural History (7)17(97):98-100.	https://www.biodiversitylibrary.org/page/19245317	BMNH:Mamm:1879.11.21.164	holotype	https://data.nhm.ac.uk/object/d973c828-6562-4d73-bde1-e1f620676d13	"Mussooree, North-western Himalayas [Dehra Dun, Uttar Pradesh, India], at an altitude of 4500 to 5500 feet [= 1372 to 1676 m]."			NA	NA				Afghanistan|Pakistan?|India|Nepal	Asia	Palearctic	LC	0	0	0	Nyctalus_montanus	0	sciname match	Nyctalus_montanus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Nyctalus		montanus	Barrett-Hamilton	1906	1	Ann. Mag. Nat. Hist.	ser. 7, 17: 99	Mountain Noctule	None.	India, Uttar Pradesh, Dehra Dun, Mussooree.	E Afghanistan, Pakistan, N India, Nepal.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14923/22016710/' target='_blank'>Least Concern</a>	Listed as a subspecies of leisleri by Ellerman and Morrison-Scott (1951), but see Gaisler (1970), Corbet (1978c), Corbet andHill (1992), and Bates and Harrison (1997).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctalus montanus; Nyctalus montanus; Nyctalus montanus; Nyctalus montanus; Nyctalus montanus; Nyctalus montanus; montanus; Noctule des montagnes; Bergabendsegler; Noctulomontano; Himalayan Noctule; Mountain Noctule; Himalayan Noctule; Mountain Noctule; Mountain Noctule; N. montanus
