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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1031	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri	Nyctalus leisleri		[MSW2] Includes verrucosus, see Corbet (1978c:55), who also included azoreum; but see Palmeirim (1991).; [MSW3] Includes verrucosus, see Corbet (1978c), who also included azoreum; but see Palmeirim (1991). Reviewed in part by Bates and Harrison (1997). For discussion of correct spelling (leisleri) see Bogdanowicz and Kock (1998).; [HMW] Vespertilio leisler: Kuhl, 1817 , Hanau, Hessen , Germany . Nyctalus leisleri 1s sister to N. azorewm, with very limited genetic distinction, although N. azorewm 1s morphologically and ecologically distinctive and has been isolated on the Azores for enough time to permit some genetic variation between differ ent island populations. The insular subspecies verrucosus has sometimes been considered a distinct species but is here included in N. leisleri , based on cranial morphometrics and genetic data. Two subspecies recognized.; [batnames2022] Includes verrucosus , see Corbet (1978 c ), who also included azoreum ; but see Palmeirim (1991). Reviewed in part byBates and Harrison (1997). For discussion of correct spelling ( leisleri ) see Bogdanowicz and Kock (1998).; [batnames2023] Includes verrucosus , see Corbet (1978 c ), who also included azoreum ; but see Palmeirim (1991). Reviewed in part byBates and Harrison (1997). For discussion of correct spelling ( leisleri ) see Bogdanowicz and Kock (1998).; [batnames2025_1.7] Includes verrucosus, see Corbet (1978c), who also included azoreum; but see Palmeirim (1991). Reviewed in part byBates and Harrison (1997). For discussion of correct spelling (leisleri) see Bogdanowicz and Kock (1998).				azoreum, verrucosus		dasykarpos, madeirae, pachygnathus, verrucosus.	azoreum, verrucosus, leisleri	leisleri, verrucosus	dasykarpos, pachygnathus; verrucosus - madeirae	leisleri, verrucosus		leisleri, verrucosus	leisleri - dasykarpos, pachygnathus; verrucosus - madeirae	leisleri, dasykarpos, verrucosus, pachygnathus, madeirae		leisleri, verrucosus	leisleri - dasykarpos, pachygnathus; verrucosus - madeirae	leisleri, dasykarpos, verrucosus, pachygnathus, madeirae	dasykarpos, leisleri, dasycarpus, verrucosus, lesleria, pachygnathus, leislerii, madeirae	leisleri, verrucosus 	leisleri - dasykarpos, pachygnathus; verrucosus - madeirae	dasykarpos (Kuhl, 1817) [not used as valid | nomen novum]|leisleri (Kuhl, 1817)|dasycarpus (A. G. Desmarest, 1819)|verrucosus Bowdich, 1825|lesleria (S.D.W., 1836) [unjustified emendation]|pachygnathus (Michahelles in J. A. Wagner, 1839)|leislerii (Temminck, 1840) [incorrect subsequent spelling]|madeirae (Barrett-Hamilton, 1906)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Lesser noctule	Madeira, Azores, W Europe – N India	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nyctalus leisleri	Germany, Hessen, Hanau.	Kuhl	1818	Ann. Wetterau Ges. Naturk., 4(1):46.	Distribution: Ranging from western Europe and extreme northern Africa to Pakistan, also Madeira and the Azores.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Lesser noctule (Leisler's bat)	Madeira, Canary Is, N Africa, W Europe – Kashmir	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Kuhl	1817	Die Deutschen Fledermause. Hanau, p. 14, 46.	Includes verrucosus, see Corbet (1978c:55), who also included azoreum; but see Palmeirim (1991).	W Europe to Urals and Caucasus; Britain; Ireland; Madeira Isl (Morocco); Azores (Portugal); W Himalayas; E Afghanistan; NW Africa.	Germany, Hessen, Hanau.		KUHL	1817	Size fairly small (forearm length, 35-46 mm). Rostrum of medium length and width. Supraorbital tubercle poorly developed. Accessory cusp on upper canine poorly developed. Anterior upper premolar displaced medially but not greatly reduced.	Distribution: Ranging from western Europe and extreme northern Africa to Pakistan, also Madeira and the Azores.	Three subspecies are recognized:	N. I. azoreum (Azores), N. I. verrucosus (Madeira), N. I. leisleri (remainder of range).	118	species	N. leisleri	KUHL	1817	Nyctalus	genus	Nyctalus leisleri				Size fairly small (forearm length, 35-46 mm). Rostrum of medium length and width. Supraorbital tubercle poorly developed. Accessory cusp on upper canine poorly developed. Anterior upper premolar displaced medially but not greatly reduced.	Three subspecies are recognized:		4. N. leisleri (KUHL 1817) [noctula group],	4	_N. l. leisleri_ (Kuhl, 1817) (synonyms: _dasycarpus_ (Desmarest, 1819), _dasykarpos_ (Kuhl, 1817), _pachygnathus_ (Michahelles, 1839)); _N. l. verrucosus_ Bowdich, 1825 (synonyms: _madeirae_ (Barrett-Hamilton, 1906))			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Nyctalus leisleri	Nyctalus		leisleri	Kuhl	y	1817		Die Deutschen Fledermäuse. Hanau			14, 46		Leisler's Noctule	Germany, Hessen, Hanau.	W Europe to Urals, Caucasus, and Turkey; Britain and Ireland; Sweden, S Finland, Baltic states; Madeira Isl; W Himalayas, Pakistan, E Afghanistan; NW Africa.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	dasykarpos Kuhl, 1819; pachygnathus Michahelles, 1839; verrucosus Bowditch, 1825; madeirae Barrett-Hamilton, 1906.	Includes verrucosus, see Corbet (1978c), who also included azoreum; but see Palmeirim (1991). Reviewed in part by Bates and Harrison (1997). For discussion of correct spelling (leisleri) see Bogdanowicz and Kock (1998).	4C3D87E8FFF26A4EFA8A90281FFAB1F6	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	767	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFF26A4EFA8A90281FFAB1F6.xml	Nyctalus leisleri	Vespertilionidae	Nyctalus	leisleri		1817	Noctule de Leisler @fr | Kleiner Abendsegler @de | Néctulopequeno @es | Leisler’s Bat @en | Lesser Noctule @en	Vespertilio leisler: Kuhl, 1817 , Hanau, Hessen , Germany . Nyctalus leisleri 1s sister to N. azorewm, with very limited genetic distinction, although N. azorewm 1s morphologically and ecologically distinctive and has been isolated on the Azores for enough time to permit some genetic variation between differ ent island populations. The insular subspecies verrucosus has sometimes been considered a distinct species but is here included in N. leisleri , based on cranial morphometrics and genetic data. Two subspecies recognized.	N.l.leisleriKuhl,1817—throughoutEuropefromIreland,GreatBritain,andIberianPeninsulaEtoSWRussia,Caucasus,Turkey,andNIran,andanisolatedpopulationinSSweden,aswellasCorsica,Cyprus,W&NCrete(recordsmaybeaccidentalmigrants),andCanaryIs(TenerifeandLaPalma),NAfricainNMorocco,NAlgeria,andarecordfromNELibya,andSCAsiainNEAfghanistan,NPakistan,andNWIndia;possiblyalsofoundonRhodesIoffWTurkeybutrecordthereisprobablyavagrantfrommainlandpopulations. N. l. verrucosus Bowditch, 1825 — Madeira I.	Head—body 48-72 mm, tail 35-48 mm, ear 11-16 mm, hindfoot 9-10 mm, forearm 35-46 mm; weight 8-20 g. Dorsal pelage of Leisler’s Noctule is golden yellowish brown (hairs dark brown at base, becoming golden brown toward tip), ventral pelage is a lighter yellowish (hairs blackish brown with brown tip). Face, ears, and membranes are dark brown, and tail extends a few millimeters past uropatagium. Muzzle is short, with large glands between nostrils and eyes, and ears are short and triangular, with 4-5 folds on outer edge. Tragus is very short and rounded, mushroom-shaped,as is characteristic of the genus. Wings attach at ankle, and calcar reaches halfway to tail. Postcalcarial lobe is wide with visible T-shaped piece of cartilage. Skull is small, and rostrum is broad; lambdoid crest is weakly developed, and there is no sagittal crest; dentition is moderately weak; lower molars are nyctalodont. Chromosomal complement has 2n = 46 and FNa = 50.	A variety of habitats including temperate deciduous and coniferous forests, agricultural regions, and river valleys. In North Africa, the species is found mainly in oak forests (as well as juniper-cypress forests) at elevations of 400-900 m. Leisler’s Noctules have been found at elevations from sea level to ¢. 2400 m throughout their range.	Insectivorous. Leisler’s Noctules feed on a variety of medium-sized flying insects, their diet consisting primarily of lepidopterans and flies (particularly Nematocera) in most parts of their range, although this is variable. They are also known to feed on a variety of other insects, including Trichoptera, Neuroptera , Coleoptera , Ephemeroptera , and Hymenoptera . Leisler’s Noctule is a fast aerial hawker, generally forages above or along the canopy of forests and along vegetation or above river valleys and other open areas.	Mating occurs in late summer before hibernation in late August to September and the species exhibits delayed fertilization. Parturition occurs after hibernation. Young are born around mid-summer, in June and July; lactation can occur until mid-July, with the first volant young usually recorded in July, although births have been recorded in May, with volant young in June in Morocco . Litter size is 1-2 (two is commoner). Leisler’s Noctules can apparently live up to nine years.	Roosts during summer are found in tree holes and rock crevices, as well as in buildings and other man-made structures, whereas in winter they are mainly in tree holes and occasionally in underground sites (e.g. tunnels and animal holes) or buildings. Leisler’s Noctule is largely nocturnal, leaving the roost after dusk and foraging throughout most of the night, going back to the roost ¢.30 minutes before dawn. It forages in two periods, one just after dusk and the other before dawn. It uses temporary night roosts between the two foraging bouts throughout the night; these are often in buildings. Leisler’s Noctules have been observed flying during the day, but this is probably only out of necessity. They are active throughout summer but enter hibernation, usually starting around mid-August to September and ending during spring, around March or April. In Morocco , the hibernation period is shorter, and individuals leave hibernation earlier. Activity peaks before and after hibernation, which is also when migrations generally occur. There are two types of search-phase calls, a QCF call shape with start frequencies of 22-26 kHz and call durations of 15-25 milliseconds, and a shallow FM call shape with start frequencies of 26-30 kHz, end frequencies of 25-26 kHz, peak frequencies of 25-26 kHz, call durations of 7-15 milliseconds, and intercall intervals of 220-550 milliseconds. Predators include diurnal raptors such as peregrine falcons (Falco peregrinus), Eurasian hobbies (F subbuteo), and common kestrels (F tinnunculus).	Most populations of Leisler’s Noctule in central and eastern continental Europe are highly migratory and will travel hundreds to over a thousand kilometers. Most bats banded in Germany moved in a southwest direction. Many island and southern populations, where winter is less severe, are typically non-migratory, but there are records of individuals migrating within and between Ireland and Great Britain . Migration generally occurs after leaving hibernation in spring (March—May) and before entering hibernation in late summer and early autumn (August—early October); young are often born in the north, with hibernation generally occurring in the south. Some island records may be attributed to vagrants (e.g. on Rhodes and Crete). The longest recorded movement was of a female which traveled 1567 km from Germany to Spain . Average foraging area is ¢.7-4 km? but can be up to c.18-4 km*in southern UK . Although notterritorial throughout much of the year, males become very territorial during the mating season, when they form small mating harems in roosts of up to nine females. Roosts in summer consist mostly of maternity colonies with females and young, and bachelor colonies for males. Maternity colonies generally comprise c.10-50 individuals, but roosts of up to ¢.1000 individuals have been reported in Ireland . Males and immatures roost either alone or together in smaller roosts.	Classified as Least Concern on The IUCN Red List. Leisler’s Noctule is widespread and common throughout parts ofits distribution (particularly Ireland ), although it is considered rare in many regions, and has become locally extinct in parts of central European Russia . The species may be threatened by the destruction of roosts in trees and buildings, as well as general loss of habitat. In Italy , introduced rose-ringed parakeets ( Psittacula krameri) have been recorded killing a roosting Leisler’s Noctule.	Aulagnier (2013i) | Beck (1995) | Benda & Gaisler (2015) | Benda, Faizolahi et al. (2012) | Benda, Georgiakakis et al. (2008) | Benda, Hanék et al. (2007) | Benda, Spitzenberger et al. (2014) | Boston (2008) | Boston, Montgomery et al. (2015) | Boston, Roué et al. (2012) | Dondini & Vergari (2009) | Juste & Paunovié (2016a) | Kafhuch, Kristin & Kristofik (2005) | Menchetti, Scalera & Mori (2014) | Mikula et al. (2016) | Ohlendorf et al. (2000) | Palmeirim (1991) | Ruczynski et al. (2010) | Russ, Briffa & Montgomery (2003) | Salgueiro, Coelho et al. (2004) | Salgueiro, Palmeirim & Coelho (2010) | Salgueiro, Ruedi et al. (2007) | Shiel & Fairley (1998) | Shiel, Duvergé et al. (1998) | Shiel, Shiel & Fairley (1999) | Spada et al. (2008) | Speakman & Webb (1993) | Stebbings & Griffith (1986) | Sullivan et al. (1993) | Waters, Jones & Furlong (1999) | Waters, Rydell & Jones (1995) | Zingg (1988)	https://zenodo.org/record/6397802/files/figure.png	16. Leisler’s Noctule Nyctalus leisleri French: Noctule de Leisler / German: Kleiner Abendsegler / Spanish: Néctulo pequeno Other common names: Leisler’s Bat , Lesser Noctule Taxonomy. Vespertilio leisler: Kuhl, 1817 , Hanau, Hessen , Germany . Nyctalus leisleri 1s sister to N. azorewm, with very limited genetic distinction, although N. azorewm 1s morphologically and ecologically distinctive and has been isolated on the Azores for enough time to permit some genetic variation between differ ent island populations. The insular subspecies verrucosus has sometimes been considered a distinct species but is here included in N. leisleri , based on cranial morphometrics and genetic data. Two subspecies recognized. Subspecies and Distribution. N.l.leisleriKuhl,1817—throughoutEuropefromIreland,GreatBritain,andIberianPeninsulaEtoSWRussia,Caucasus,Turkey,andNIran,andanisolatedpopulationinSSweden,aswellasCorsica,Cyprus,W&NCrete(recordsmaybeaccidentalmigrants),andCanaryIs(TenerifeandLaPalma),NAfricainNMorocco,NAlgeria,andarecordfromNELibya,andSCAsiainNEAfghanistan,NPakistan,andNWIndia;possiblyalsofoundonRhodesIoffWTurkeybutrecordthereisprobablyavagrantfrommainlandpopulations. N. l. verrucosus Bowditch, 1825 — Madeira I. Descriptive notes. Head—body 48-72 mm, tail 35-48 mm, ear 11-16 mm, hindfoot 9-10 mm, forearm 35-46 mm; weight 8-20 g. Dorsal pelage of Leisler’s Noctule is golden yellowish brown (hairs dark brown at base, becoming golden brown toward tip), ventral pelage is a lighter yellowish (hairs blackish brown with brown tip). Face, ears, and membranes are dark brown, and tail extends a few millimeters past uropatagium. Muzzle is short, with large glands between nostrils and eyes, and ears are short and triangular, with 4-5 folds on outer edge. Tragus is very short and rounded, mushroom-shaped,as is characteristic of the genus. Wings attach at ankle, and calcar reaches halfway to tail. Postcalcarial lobe is wide with visible T-shaped piece of cartilage. Skull is small, and rostrum is broad; lambdoid crest is weakly developed, and there is no sagittal crest; dentition is moderately weak; lower molars are nyctalodont. Chromosomal complement has 2n = 46 and FNa = 50. Habitat. A variety of habitats including temperate deciduous and coniferous forests, agricultural regions, and river valleys. In North Africa, the species is found mainly in oak forests (as well as juniper-cypress forests) at elevations of 400-900 m. Leisler’s Noctules have been found at elevations from sea level to ¢. 2400 m throughout their range. Food and Feeding. Insectivorous. Leisler’s Noctules feed on a variety of medium-sized flying insects, their diet consisting primarily of lepidopterans and flies (particularly Nematocera) in most parts of their range, although this is variable. They are also known to feed on a variety of other insects, including Trichoptera, Neuroptera , Coleoptera , Ephemeroptera , and Hymenoptera . Leisler’s Noctule is a fast aerial hawker, generally forages above or along the canopy of forests and along vegetation or above river valleys and other open areas. Breeding. Mating occurs in late summer before hibernation in late August to September and the species exhibits delayed fertilization. Parturition occurs after hibernation. Young are born around mid-summer, in June and July; lactation can occur until mid-July, with the first volant young usually recorded in July, although births have been recorded in May, with volant young in June in Morocco . Litter size is 1-2 (two is commoner). Leisler’s Noctules can apparently live up to nine years. Activity patterns. Roosts during summer are found in tree holes and rock crevices, as well as in buildings and other man-made structures, whereas in winter they are mainly in tree holes and occasionally in underground sites (e.g. tunnels and animal holes) or buildings. Leisler’s Noctule is largely nocturnal, leaving the roost after dusk and foraging throughout most of the night, going back to the roost ¢.30 minutes before dawn. It forages in two periods, one just after dusk and the other before dawn. It uses temporary night roosts between the two foraging bouts throughout the night; these are often in buildings. Leisler’s Noctules have been observed flying during the day, but this is probably only out of necessity. They are active throughout summer but enter hibernation, usually starting around mid-August to September and ending during spring, around March or April. In Morocco , the hibernation period is shorter, and individuals leave hibernation earlier. Activity peaks before and after hibernation, which is also when migrations generally occur. There are two types of search-phase calls, a QCF call shape with start frequencies of 22-26 kHz and call durations of 15-25 milliseconds, and a shallow FM call shape with start frequencies of 26-30 kHz, end frequencies of 25-26 kHz, peak frequencies of 25-26 kHz, call durations of 7-15 milliseconds, and intercall intervals of 220-550 milliseconds. Predators include diurnal raptors such as peregrine falcons (Falco peregrinus), Eurasian hobbies (F subbuteo), and common kestrels (F tinnunculus). Movements, Home range and Social organization. Most populations of Leisler’s Noctule in central and eastern continental Europe are highly migratory and will travel hundreds to over a thousand kilometers. Most bats banded in Germany moved in a southwest direction. Many island and southern populations, where winter is less severe, are typically non-migratory, but there are records of individuals migrating within and between Ireland and Great Britain . Migration generally occurs after leaving hibernation in spring (March—May) and before entering hibernation in late summer and early autumn (August—early October); young are often born in the north, with hibernation generally occurring in the south. Some island records may be attributed to vagrants (e.g. on Rhodes and Crete). The longest recorded movement was of a female which traveled 1567 km from Germany to Spain . Average foraging area is ¢.7-4 km? but can be up to c.18-4 km*in southern UK . Although notterritorial throughout much of the year, males become very territorial during the mating season, when they form small mating harems in roosts of up to nine females. Roosts in summer consist mostly of maternity colonies with females and young, and bachelor colonies for males. Maternity colonies generally comprise c.10-50 individuals, but roosts of up to ¢.1000 individuals have been reported in Ireland . Males and immatures roost either alone or together in smaller roosts. Status and Conservation. Classified as Least Concern on The IUCN Red List. Leisler’s Noctule is widespread and common throughout parts ofits distribution (particularly Ireland ), although it is considered rare in many regions, and has become locally extinct in parts of central European Russia . The species may be threatened by the destruction of roosts in trees and buildings, as well as general loss of habitat. In Italy , introduced rose-ringed parakeets ( Psittacula krameri) have been recorded killing a roosting Leisler’s Noctule. Bibliography. Aulagnier (2013i), Beck (1995), Benda & Gaisler (2015), Benda, Faizolahi et al. (2012), Benda, Georgiakakis et al. (2008), Benda, Hanék et al. (2007), Benda, Spitzenberger et al. (2014), Boston (2008), Boston, Montgomery et al. (2015), Boston, Roué et al. (2012), Dondini & Vergari (2009), Juste & Paunovié (2016a), Kafhuch, Kristin & Kristofik (2005), Menchetti, Scalera & Mori (2014), Mikula et al. (2016), Ohlendorf et al. (2000), Palmeirim (1991), Ruczynski et al. (2010), Russ, Briffa & Montgomery (2003), Salgueiro, Coelho et al. (2004), Salgueiro, Palmeirim & Coelho (2010), Salgueiro, Ruedi et al. (2007), Shiel & Fairley (1998), Shiel, Duvergé et al. (1998), Shiel, Shiel & Fairley (1999), Spada et al. (2008), Speakman & Webb (1993), Stebbings & Griffith (1986), Sullivan et al. (1993), Waters, Jones & Furlong (1999), Waters, Rydell & Jones (1995), Zingg (1988).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Nyctalus leisleri	Nyctalus		leisleri	Kuhl	1817	1	Die Deutschen Flederm&auml;use. Hanau	p. 14, 46	Leisler's Noctule	 dasykarpos Kuhl, 1819; pachygnathus Michahelles, 1839; <b>verrucosus</b> Bowditch, 1825; madeirae Barrett-Hamilton, 1906.	Germany, Hessen, Hanau.	W Europe to Urals, Caucasus, and Turkey; Britain and Ireland; Sweden, S Finland, Baltic states; Madeira Isl; W Himalayas, Pakistan, E Afghanistan; NW Africa.	Not listed.	Least Concern	Includes verrucosus , see Corbet (1978 c ), who also included azoreum ; but see Palmeirim (1991). Reviewed in part byBates and Harrison (1997). For discussion of correct spelling ( leisleri ) see Bogdanowicz and Kock (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctalus leisleri	23	Leisler's Noctule	Leisler's Bat|Lesser Noctule	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Nyctalus	NA	leisleri	Kuhl	1817	1	Vespertilio_Leisleri	Kuhl, H. (1817). Die Deutschen Fledermäuse. Hanau, 6.		ZMB 471, ZMB 472 [syntypes]		Hanau, Hessen, Germany.			leisleri (Kuhl, 1817)|dasykarpos (Kuhl, 1819)|verrucosus Bowditch, 1825|pachygnathus (Michahelles, 1839)|madeirae Barrett-Hamilton, 1906	NA	NA	Morocco|Algeria|Canary Islands|Madeira|Ireland|United Kingdom|Portugal|Spain|France|Luxembourg|Belgium|Netherlands|Germany|Sweden|Switzerland|Liechtenstein|Italy|Austria|Slovenia|Czech Republic|Poland|Hungary|Slovakia|Croatia|Bosnia & Herzegovina|Serbia|Kosovo|Albania|Montenegro|North Macedonia|Greece|Bulgaria|Romania|Moldova|Ukraine|Belarus|Lithuania|Latvia|Russia|Kazakhstan?|Turkey|Cyprus|Georgia|Armenia|Azerbaijan|Iran|Afghanistan|Pakistan|India|China?	Africa|Asia|Europe	Palearctic	LC	0	0	0	Nyctalus_leisleri	0	sciname match	Nyctalus_leisleri	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14919	Nyctalus leisleri	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Nyctalus	leisleri	(Kuhl, 1817)		20000000	Nyctalus leisleri	Least Concern		2016	2016-04-25 00:00:00 UTC	3.1	English	The species is widespread and abundant, and there is no evidence of current significant population decline. Consequently it is assessed as Least Concern.	The Lesser Noctule (Nyctalus leisleri ) ;forages over woodland ;<span lang="EN-US">(both Eurosiberian and Mediterranean), pasture, and river valleys, where it feeds on flies (including mosquitos), moths and beetles. It is linked to old trees. Summer nursery roosts are located in tree holes, but also in buildings and bat boxes. Nursery colonies usually number 20-50 females, occasionally up to 1,000 (e.g., in Ireland: Stebbings and Griffith 1986). In winter this bat hibernates mainly in tree holes, or occasionally in underground sites or buildings, often in large groups. Females migrate over distances up to 1,567 km (Ohlendorf et al. 2000).	Threats include disturbance to and destruction of roosts in trees and buildings, and loss or degradation of foraging habitat. However, these are not thought to be major threats at present.	It is widespread although patchily distributed in Europe. Common in parts of range (e.g., Ireland), scarce in other parts (Stebbings and Griffith 1986). Local extinctions have been reported for the central part of Russian Federation (K. Tsytsulina pers. comm. 2005), although the species remains common in other parts of Russia and the Caucasus (S. Kruskop pers. comm. 2005). In North Africa it is limited by the distribution of suitable habitat, but is particularly abundant in Djurdjura, NE Algeria. There is no information about trends.	Unknown	The Lesser Noctule (Nyctalus leisleri ) ; is largely a western Palaearctic species (Europe and north-west Africa), with scattered records in the western parts of the eastern Palaearctic (Pakistan, Afghanistan, the Himalayas). It is widely distributed in Europe from southern Scotland and Ireland approximately along the southern edge of the Baltic Sea south and parts of Mediterranean coast to western Russia. It is present on Crete (Benda et al. 2009), ;Madeira and the Canary Islands (on Tenerife and La Palma only) but absent from southwestern Italy and Sicily, Southeastern Spain, most of Fennoscandia and northern Russia. In North Africa it is recorded from Mediterranean montane Morocco and Algeria and there is one record from Cyrenaica (Libya). It occurs from sea level to 2,400 m.		Terrestrial	It is protected by national legislation in most range states. There are also international legal obligations for its protection through the Bonn Convention (Eurobats) and Bern Convention, in parts of its range where these apply. It is included in Annex IV of EU Habitats and Species Directive, and there is some habitat protection through Natura 2000. It occurs in protected areas throughout its range. No specific conservation actions known.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Nyctalus		leisleri	Kuhl	1817	1	Die Deutschen Flederm&auml;use. Hanau	p. 14, 46	Leisler's Noctule	 dasykarpos Kuhl, 1819; pachygnathus Michahelles, 1839; <b>verrucosus</b> Bowditch, 1825; madeirae Barrett-Hamilton, 1906.	Germany, Hessen, Hanau.	W Europe to Urals, Caucasus, and Turkey; Britain and Ireland; Sweden, S Finland, Baltic states; Madeira Isl; W Himalayas, Pakistan, E Afghanistan; NW Africa.	Not listed.	Least Concern	Includes verrucosus , see Corbet (1978 c ), who also included azoreum ; but see Palmeirim (1991). Reviewed in part byBates and Harrison (1997). For discussion of correct spelling ( leisleri ) see Bogdanowicz and Kock (1998).	Nyctalus leisleri	1005605	23	Leisler's Noctule	Leisler's Bat|Lesser Noctule	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Nyctalus	NA	leisleri	Kuhl	1817	1	Vespertilio_Leisleri	Kuhl, H. (1817). Die Deutschen Fledermäuse. Hanau, 6.		ZMB 471, ZMB 472 [syntypes]		Hanau, Hessen, Germany.			leisleri (Kuhl, 1817)|dasykarpos (Kuhl, 1819)|verrucosus Bowditch, 1825|pachygnathus (Michahelles, 1839)|madeirae Barrett-Hamilton, 1906	NA	NA				Morocco|Algeria|Canary Islands|Madeira|Ireland|United Kingdom|Portugal|Spain|France|Luxembourg|Belgium|Netherlands|Germany|Sweden|Switzerland|Liechtenstein|Italy|Austria|Slovenia|Czech Republic|Poland|Hungary|Slovakia|Croatia|Bosnia & Herzegovina|Serbia|Kosovo|Albania|Montenegro|North Macedonia|Greece|Bulgaria|Romania|Moldova|Ukraine|Belarus|Lithuania|Latvia|Russia|Kazakhstan?|Turkey|Cyprus|Georgia|Armenia|Azerbaijan|Iran|Afghanistan|Pakistan|India|China?	Africa|Asia|Europe	Palearctic	LC	0	0	0	Nyctalus_leisleri	0	sciname match	Nyctalus_leisleri	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctalus_leisleri	1005605	23	Leisler's Noctule	Leisler's Bat|Lesser Noctule	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Nyctalus	NA	leisleri	Kuhl	1	Vespertilio Leisleri	Kuhl, H. 1817. Die deutschen Fledermäuse. None, Hanau, 67 pp.	https://nbn-resolving.org/urn:nbn:de:hebis:30-91692	ZMB 471, ZMB 472	syntypes		Hanau, Hessen, Germany.			NA	NA				Morocco|Algeria|Canary Islands|Madeira|Ireland|United Kingdom|Portugal|Spain|France|Luxembourg|Belgium|Netherlands|Germany|Sweden|Switzerland|Liechtenstein|Italy|Austria|Slovenia|Czech Republic|Poland|Hungary|Slovakia|Croatia|Bosnia and Herzegovina|Serbia|Kosovo|Albania|Montenegro|North Macedonia|Greece|Bulgaria|Romania|Moldova|Ukraine|Belarus|Lithuania|Latvia|Russia|Kazakhstan?|Turkey|Cyprus|Georgia|Armenia|Azerbaijan|Iran|Afghanistan|Pakistan|India|China?	Africa|Asia|Europe	Palearctic	LC	0	0	0	Nyctalus_leisleri	0	sciname match	Nyctalus_leisleri	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Nyctalus		leisleri	Kuhl	1817	1	Die Deutschen Flederm&auml;use. Hanau	p. 6, 38	Leisler's Noctule	dasykarpos Kuhl, 1819; pachygnathus Michahelles, 1839; verrucosus Bowditch, 1825; madeirae Barrett-Hamilton, 1906.	Germany, Hessen, Hanau.	W Europe to Urals, Caucasus, and Turkey; Britain and Ireland; Sweden, S Finland, Baltic states; Madeira Isl; W Himalayas, Pakistan, E Afghanistan; NW Africa.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14919/22016159/' target='_blank'>Least Concern</a>	Includes verrucosus, see Corbet (1978c), who also included azoreum; but see Palmeirim (1991). Reviewed in part byBates and Harrison (1997). For discussion of correct spelling (leisleri) see Bogdanowicz and Kock (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctalus leisleri; Nyctalus leisleri; Nyctalus leisleri; Nyctalus leisleri; Nyctalus leisleri; Nyctalus leisleri; leisleri; verrucosus; dasykarpos; pachygnathus; verrucosus - madeirae; leisleri; verrucosus; verrucosus; dasykarpos; pachygnathus; verrucosus - madeirae; leisleri; dasykarpos; verrucosus; pachygnathus; madeirae; Noctule de Leisler; Kleiner Abendsegler; Néctulopequeno; Leisler’s Bat; Lesser Noctule; Leisler's Noctule; Leisler's Bat; Lesser Noctule; Leisler's Noctule; Leisler's Noctule; N. leisleri