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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1028	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Nyctalus leisleri [synonym of]	Nyctalus azoreum	Nyctalus azoreum	N/A	Nyctalus azoreum	Nyctalus azoreum	Nyctalus azoreum	Nyctalus azoreum	Nyctalus azoreum	Nyctalus azoreum	Nyctalus azoreum	Nyctalus azoreum	Nyctalus azoreum	Nyctalus azoreum		[MSW2] Listed as a subspecies of N. leisleri by Corbet (1978c), but see Palmeirim (1991).; [MSW3] Listed as a subspecies of leisleri by Corbet (1978c), but see Palmeirim (1991) and Horácek et al. (2000).; [HMW] Pterygistes azoreum Thomas, 1901 , “St. Michael’s, Azores ,” Portugal . Nyctalus azoreum is morphologically and ecologically distinct from N. leisleri but is genetically not particularly divergent. However, there is considerable genetic divergence between different island populations of N. azoreum , indicating that the species has been present on the islands for a long time, and that there is limited breeding between islands. Monotypic.; [batnames2022] Listed as a subspecies of leisleri by Corbet (1978 c ), but see Palmeirim (1991) and HorÃ¡cek et al. (2000).; [IUCN] There is marked genetic differentiation between populations on central and eastern islands in the archipelago, indicating that they have evolved in relative isolation for a long time (Salgueiro et al. 2004).; [batnames2023] Listed as a subspecies of leisleri by Corbet (1978 c ), but see Palmeirim (1991) and HorÃ¡cek et al. (2000).; [batnames2025_1.7] Listed as a subspecies of leisleri by Corbet (1978c), but see Palmeirim (1991) and HorÃ¡cek et al. (2000).														azoreum	There is marked genetic differentiation between populations on central and eastern islands in the archipelago, indicating that they have evolved in relative isolation for a long time (Salgueiro et al. 2004).			azoreum 	azoreum 			azoreum (O. Thomas, 1901)						N/A							Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Azores Is; ref. 4.75	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1901	Ann. Mag. Nat. Hist., ser. 7, 8:34.	Listed as a subspecies of N. leisleri by Corbet (1978c), but see Palmeirim (1991).	Azores (Portugal).	Portugal, Azores, St. Michael.																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Nyctalus azoreum	Nyctalus		azoreum	Thomas	y	1901		Ann. Mag. Nat. Hist., ser. 7	8		34		Azores Noctule	Portugal, Azores, St. Michael.	Azores Isls (Portugal).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Listed as a subspecies of leisleri by Corbet (1978c), but see Palmeirim (1991) and Horácek et al. (2000).	4C3D87E8FFF16A4EFA5297501962B1F7	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	768	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFF16A4EFA5297501962B1F7.xml	Nyctalus azoreum	Vespertilionidae	Nyctalus	azoreum		1901	Noctule des Acores @fr | Azoren-Abendsegler @de | Noctulode las Azores @es | Azorean Bat @en	Pterygistes azoreum Thomas, 1901 , “St. Michael’s, Azores ,” Portugal . Nyctalus azoreum is morphologically and ecologically distinct from N. leisleri but is genetically not particularly divergent. However, there is considerable genetic divergence between different island populations of N. azoreum , indicating that the species has been present on the islands for a long time, and that there is limited breeding between islands. Monotypic.	The Azores Archipelago on Faial, Pico, Sao Jorge , Graciosa, Terceira, Sao Miguel , and Santa Maria Is.	Head-body 51-57 mm, tail 42-43 mm, ear 12 mm , forearm 35-42 mm; weight 6-15 g. The Azores Noctule is externally very similar to Leisler’s Noctule ( Nyctalus leisleri ) but smaller and darker. Dorsal pelage is dark brown with a yellowish hue, while ventral pelage is yellowish. Face, ears, and membranes are darkly pigmented;tail extends a few millimeters past uropatagium. Muzzle is short, with large glands between nostrils and eyes; ears are short and triangular, with 4-5 folds on outer edge. Tragus is very short and rounded, mushroom-shaped, as is characteristic of the genus. Skull is robust and lacks a defined forehead; skull and dentition are similar to Leisler’s Noctule overall; lower molars are nyctalodont.	Forages in a variety of habitats across the islands. The Azores Noctule favors natural and semi-natural dry forest and heathland, although it is frequently found feeding around agricultural fields and villages. Elevation range is 0-600 m.	Insectivorous. Azores Noctules feed largely during the day, and stay high in the canopy and over the vegetation in open areas when foraging. Some individuals have been observed foraging around streetlights at night.	Most births occur in mid-June to July; litter size is 1-2 young. Young begin to fly in about mid-July. Like many temperate bats, Azores Noctules probably exhibit delayed fertilization and mate earlier in the year.	Roosts are most often found in buildings, rock crevices in cliffs, and holes in trees. Unlike most noctules and bats in general, the Azores Noctule often forages diurnally, leaving its roosts before sunset. This may be because of the lack of predation or competition with birds on the islands. Calls are similar to those of Leisler’s Noctule but at a higher maximum energy frequency (32-1 kHz).	The Azores Noctule lives in colonies of individuals that roost together; when leaving the roost, they cluster in small groups, as do continental bats. Females create maternity colonies with only females and young from April until September or October, while males are solitary or in small groups during breeding season, although outside this period both males and females live solitarily or in small groups. Mean number of individuals in a roost, as indicated by M. Leonardo and F. M. Medeiros in 2011, was five, with a range of 1-684 individuals. Maternal roosts were up to 684 individuals in buildings, while in trees cavities they were generally 20-50 individuals, depending on the size of the cavity.	Classified as Vulnerable on The IUCN Red List. The Azores Noctule is restricted to the Azores , where it is relatively common on Sao Miguel , Faial, Terceira, and Sao Jorge . However,it is rare on Graciosa and extremely rare on Santa Maria, and it may be declining throughout its entire range due to habitat degradation, roost destruction, and other forms of human persecution. Total population is estimated to be 2000-5000 individuals, with less than 1000 individuals on Sao Miguel , where the species is most abundant. Because the species flies during the day, it is especially vulnerable to persecution by people, as colonies are easier to find. The Azores Noctule is the only mammal endemic to the Azores ; the only other land mammal native to the islandsis the Greater Myotis ( Myotis myotis ).	Boston et al. (2015) | Irwin & Speakman (2003) | Leonardo & Medeiros (2011) | Moore (1975) | Palmeirim (1991) | Piraccini (2016a) | Rainho et al. (2002) | Salgueiro, Coelho et al. (2004) | Salgueiro, Palmeirim & Coelho (2010) | Salgueiro, Palmeirim, Ruedi & Coelho (2008) | Salgueiro, Ruedi,Coelho & Palmeirim (2007) | Skiba (2003) | Speakman & Webb (1993)	https://zenodo.org/record/6397806/files/figure.png	17. Azores Noctule Nyctalus azoreum French: Noctule des Acores / German: Azoren-Abendsegler / Spanish: Noctulo de las Azores Other common names: Azorean Bat Taxonomy. Pterygistes azoreum Thomas, 1901 , “St. Michael’s, Azores ,” Portugal . Nyctalus azoreum is morphologically and ecologically distinct from N. leisleri but is genetically not particularly divergent. However, there is considerable genetic divergence between different island populations of N. azoreum , indicating that the species has been present on the islands for a long time, and that there is limited breeding between islands. Monotypic. Distribution. The Azores Archipelago on Faial, Pico, Sao Jorge , Graciosa, Terceira, Sao Miguel , and Santa Maria Is. Descriptive notes. Head-body 51-57 mm, tail 42-43 mm, ear 12 mm , forearm 35-42 mm; weight 6-15 g. The Azores Noctule is externally very similar to Leisler’s Noctule ( Nyctalus leisleri ) but smaller and darker. Dorsal pelage is dark brown with a yellowish hue, while ventral pelage is yellowish. Face, ears, and membranes are darkly pigmented;tail extends a few millimeters past uropatagium. Muzzle is short, with large glands between nostrils and eyes; ears are short and triangular, with 4-5 folds on outer edge. Tragus is very short and rounded, mushroom-shaped, as is characteristic of the genus. Skull is robust and lacks a defined forehead; skull and dentition are similar to Leisler’s Noctule overall; lower molars are nyctalodont. Habitat. Forages in a variety of habitats across the islands. The Azores Noctule favors natural and semi-natural dry forest and heathland, although it is frequently found feeding around agricultural fields and villages. Elevation range is 0-600 m. Food and Feeding. Insectivorous. Azores Noctules feed largely during the day, and stay high in the canopy and over the vegetation in open areas when foraging. Some individuals have been observed foraging around streetlights at night. Breeding. Most births occur in mid-June to July; litter size is 1-2 young. Young begin to fly in about mid-July. Like many temperate bats, Azores Noctules probably exhibit delayed fertilization and mate earlier in the year. Activity patterns. Roosts are most often found in buildings, rock crevices in cliffs, and holes in trees. Unlike most noctules and bats in general, the Azores Noctule often forages diurnally, leaving its roosts before sunset. This may be because of the lack of predation or competition with birds on the islands. Calls are similar to those of Leisler’s Noctule but at a higher maximum energy frequency (32-1 kHz). Movements, Home range and Social organization. The Azores Noctule lives in colonies of individuals that roost together; when leaving the roost, they cluster in small groups, as do continental bats. Females create maternity colonies with only females and young from April until September or October, while males are solitary or in small groups during breeding season, although outside this period both males and females live solitarily or in small groups. Mean number of individuals in a roost, as indicated by M. Leonardo and F. M. Medeiros in 2011, was five, with a range of 1-684 individuals. Maternal roosts were up to 684 individuals in buildings, while in trees cavities they were generally 20-50 individuals, depending on the size of the cavity. Status and Conservation. Classified as Vulnerable on The IUCN Red List. The Azores Noctule is restricted to the Azores , where it is relatively common on Sao Miguel , Faial, Terceira, and Sao Jorge . However,it is rare on Graciosa and extremely rare on Santa Maria, and it may be declining throughout its entire range due to habitat degradation, roost destruction, and other forms of human persecution. Total population is estimated to be 2000-5000 individuals, with less than 1000 individuals on Sao Miguel , where the species is most abundant. Because the species flies during the day, it is especially vulnerable to persecution by people, as colonies are easier to find. The Azores Noctule is the only mammal endemic to the Azores ; the only other land mammal native to the islandsis the Greater Myotis ( Myotis myotis ). Bibliography. Boston et al. (2015), Irwin & Speakman (2003), Leonardo & Medeiros (2011), Moore (1975), Palmeirim (1991), Piraccini (2016a), Rainho et al. (2002), Salgueiro, Coelho et al. (2004), Salgueiro, Palmeirim & Coelho (2010), Salgueiro, Palmeirim, Ruedi & Coelho (2008), Salgueiro, Ruedi,Coelho & Palmeirim (2007), Skiba (2003), Speakman & Webb (1993).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Nyctalus azoreum	Nyctalus		azoreum	Thomas	1901	1	Ann. Mag. Nat. Hist.	ser. 7, 8: 34	Azores Noctule	None.	Portugal, Azores, St. Michael.	Azores Isls (Portugal).	Not listed.	Vulnerable	Listed as a subspecies of leisleri by Corbet (1978 c ), but see Palmeirim (1991) and HorÃ¡cek et al. (2000).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctalus azoreum	23	Azores Noctule	Azorean Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Nyctalus	NA	azoreum	O. Thomas	1901	1	Pterygistes_azoreum	Thomas, O. (1901). Some new African Bats (including one from the Azores) and a new Galago. Annals and Magazine of Natural History, ser. 7, 8, 33.	https://www.biodiversitylibrary.org/item/94923#page/53/mode/1up	BM 1865.10.2.1		"St. Michael's, Azores," Portugal.			azoreum (O. Thomas, 1901)	NA	NA	Azores	Europe	Palearctic	VU	0	0	0	Nyctalus_azoreum	0	sciname match	Nyctalus_azoreum	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14922	Nyctalus azoreum	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Nyctalus	azoreum	(Thomas, 1901)	There is marked genetic differentiation between populations on central and eastern islands in the archipelago, indicating that they have evolved in relative isolation for a long time (Salgueiro et al. 2004).	200000000	Nyctalus azoreum	Vulnerable	C2a(i)	2022	2016-04-25 00:00:00 UTC	3.1	English	This species' population has been estimated at 2,000-5,000 individuals and the most abundant subpopulation numbers less than 1,000 individuals. For these reasons this species meets the criteria for being assessed as Vulnerable (VU C2a(i)). This species is endemic to the Azores, where it has a small population and range. It is suspected to be declining as a result of disturbance and destruction of colonies, loss and degradation of natural and semi-natural habitats and pesticide use. There is no direct information on dispersal between islands, but genetic research suggests that dispersal is restricted, and so the population is precautionarily regarded as being severely fragmented; further research would be required to confirm this. For these reasons, it may be approaching the Endangered category.	This bat forages over a variety of habitats on the islands, favouring natural and semi-natural habitats. It frequently feeds around artificial lighting (e.g. streetlamps). Most maternity colonies are probably located in buildings, trees and rock crevices.	Human persecution, and the destruction of roost sites, are likely to be the main threats (A. Rainho pers. comm. 2006). Habitat loss and degradation, use of pesticides and the spread of exotic plant species may also have a detrimental effect on the species. It is suspected that the extreme scarcity of the species on Santa Maria is attributable to habitat loss and degradation. This species is particularly vulnerable to persecution because it flies during the day, making colonies obvious and easy to find (A. Rainho pers. comm. 2006).	Surveys in 2002, 2003 and 2004 found the Azorean Bat (Nyctalus azoreum ) quite abundant on San Miguel, Faial, Terceira and San Jorge, but rare on Graciosa and extremely rare on Santa Maria. It is absent from Flores and Corvo. There is no quantitative information on population trend, but it is suspected that the species may be declining as a result of habitat degradation, destruction of or exclusion from roosts (both in trees and buildings), and human persecution. Local environmental groups report that numerous colonies have disappeared (A. Rainho pers. comm. 2006). The total population is estimated at 2,000-5,000 individuals, and there are fewer than 1,000 individuals on San Miguel, where the species is most abundant (Cabral et al. 2005, J.M. Palmeirim, A. Rainho and L. Rodrigues pers. comm. 2006).	Decreasing	The Azorean Bat (Nyctalus azoreum ) is restricted to the Azores Archipelago (Portugal), where it occurs on Faial, Pico, San Jorge, Graciosa, Terceira, San Miguel and Santa Maria islands, from sea level to 600 m (A. Rainho pers. comm. 2006). Its Area of Occurrence is estimated at ca ; 2,200 km<sup>2</sup>.		Terrestrial	There is no specific national legislation. It is protected under Bern Convention and included in Annex IV of EU Habitats and Species Directive. There are proposals for protection and monitoring of roosts, public awareness (with special reference to roosts), reduction of adverse agricultural practices, preservation and restoration of natural habitat, use of lights that attract insects (e.g. mercury), and further studies of the species' biology (Rainho et al. 2002). ;<span lang="EN-US">A bat monitoring program, based on annual acoustic surveys, is currently under development (A. Rainho pers. comm.).	Palearctic		FALSE	FALSE	Global & Europe	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Nyctalus		azoreum	Thomas	1901	1	Ann. Mag. Nat. Hist.	ser. 7, 8(43): 33	Azores Noctule	None.	Portugal, Azores, St. Michael	Azores Isls (Portugal)	Not listed.	Vulnerable	Listed as a subspecies of leisleri by Corbet (1978 c ), but see Palmeirim (1991) and HorÃ¡cek et al. (2000).	Nyctalus azoreum	1005602	23	Azores Noctule	Azorean Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Nyctalus	NA	azoreum	O. Thomas	1901	1	Pterygistes_azoreum	Thomas, O. (1901). Some new African Bats (including one from the Azores) and a new Galago. Annals and Magazine of Natural History, ser. 7, 8, 33.	https://www.biodiversitylibrary.org/item/94923#page/53/mode/1up	BM 1865.10.2.1		"St. Michael's, Azores," Portugal.			azoreum (O. Thomas, 1901)	NA	NA				Azores	Europe	Palearctic	VU	0	0	0	Nyctalus_azoreum	0	sciname match	Nyctalus_azoreum	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctalus_azoreum	1005602	23	Azores Noctule	Azorean Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Nyctalus	NA	azoreum	O. Thomas	1	Pterygistes azoreum	Thomas, O. 1901-07-01. Some new African bats (including one from the Azores) and a new _Galago_. Annals and Magazine of Natural History (7)8(43):27-34.	https://www.biodiversitylibrary.org/page/29980035	BMNH:Mamm:1865.10.2.1	holotype	https://data.nhm.ac.uk/object/93074729-a4dd-4827-97bd-67e68988b616	"St. Michael's, Azores," Portugal.			NA	NA				Azores	Europe	Palearctic	VU	0	0	0	Nyctalus_azoreum	0	sciname match	Nyctalus_azoreum	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Nyctalus		azoreum	Thomas	1901	1	Ann. Mag. Nat. Hist.	ser. 7, 8(43): 33	Azores Noctule	None.	Portugal, Azores, St. Michael	Azores Isls (Portugal)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14922/211008291/' target='_blank'>Vulnerable</a>	Listed as a subspecies of leisleri by Corbet (1978c), but see Palmeirim (1991) and HorÃ¡cek et al. (2000).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctalus azoreum; Nyctalus azoreum; Nyctalus azoreum; Nyctalus azoreum; Nyctalus azoreum; Nyctalus azoreum; azoreum; Noctule des Acores; Azoren-Abendsegler; Noctulode las Azores; Azorean Bat; Azores Noctule; Azorean Bat; Azores Noctule; Azores Noctule; N. azoreum
