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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1027	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator	Nyctalus aviator		[MSW2] Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951:161); but see Corbet (1978c:56).; [MSW3] Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951), but see Corbet (1978c) and Yoshiyuki (1989). Reviewed by Yoshiyuki (1989).; [HMW] Nyctalus aviator Thomas, 1911 , “ Tokyo , Hondo [= Honshu], Japan .” Nyctalus aviator is sometimes included within N. lasiopterus , from which it is morphologically distinct, but to which it is genetically sister. N. aviator and N. lasiopterus are togethersister to N. noctula , all three sharing the same diploid number. Monotypic.; [batnames2022] Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951), but see Corbet (1978 c ) and Yoshiyuki (1989).Reviewed by Yoshiyuki (1989).; [IUCN] This species has previously been considered as a subspecies of lasiopterus .; [batnames2023] Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951), but see Corbet (1978 c ) and Yoshiyuki (1989).Reviewed by Yoshiyuki (1989).; [batnames2025_1.7] Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951), but see Corbet (1978c) and Yoshiyuki (1989).Reviewed by Yoshiyuki (1989).									molossus			aviator 	aviator - molossus	molossus, aviator	This species has previously been considered as a subspecies of lasiopterus .	aviator 	aviator - molossus	molossus, aviator	molossus, aviator	aviator 	aviator - molossus	molossus (Temminck, 1840) [preoccupied]|aviator O. Thomas, 1911		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Korea, Japan	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nyctalus aviator	Japan, Honshu, Tokyo.	Thomas	1911	Ann. Mag. Nat. Hist., ser. 8, 8:380.	Distribution: Confined to Japan, Korea, and northeastern China.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Korea, Japan, E China	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1911	Ann. Mag. Nat. Hist., ser. 8, 8:380.	Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951:161); but see Corbet (1978c:56).	Hokkaido, Shikoku, Kyushu, Tsushima, Iki (Japan); Korea; E China.	Japan, Honshu, Tokyo.		THOMAS	1911	Size fairly large (forearm length, 59 62 mm). Margin of plagiopatagium attached to ankle.	Distribution: Confined to Japan, Korea, and northeastern China.	No subspecies.		118	species	N. aviator	THOMAS	1911	Nyctalus	genus	Nyctalus aviator				Size fairly large (forearm length, 59 62 mm). Margin of plagiopatagium attached to ankle.	No subspecies.		7. N. aviator THOMAS 1911 [noctula group].	7	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Nyctalus aviator	Nyctalus		aviator	Thomas	y	1911		Ann. Mag. Nat. Hist., ser. 8	8		380		Birdlike Noctule	Japan, Honshu, Tokyo.	Hokkaido, Shikoku, Kyushu, Tsushima, Iki (Japan); Korea; E and C China. Possibly occurs in Russian Far East, see Tiunov (1997).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	molossus Temminck, 1840 [not Pallas, 1767].	Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951), but see Corbet (1978c) and Yoshiyuki (1989). Reviewed by Yoshiyuki (1989).	4C3D87E8FFFC6A43FF8F91831880BED0	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	765	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFFC6A43FF8F91831880BED0.xml	Nyctalus aviator	Vespertilionidae	Nyctalus	aviator	Thomas	1911	Noctule aviatrice @fr | Gro RRer Ostasien-Abendsegler @de | Noctulooriental @es | Japanese Large Noctule @en	Nyctalus aviator Thomas, 1911 , “ Tokyo , Hondo [= Honshu], Japan .” Nyctalus aviator is sometimes included within N. lasiopterus , from which it is morphologically distinct, but to which it is genetically sister. N. aviator and N. lasiopterus are togethersister to N. noctula , all three sharing the same diploid number. Monotypic.	NE & EC China ( Heilongjiang , Jilin , Henan , Anhui , and Zhejiang ), Korean Peninsula, and numerous islands in Japan ( Hokkaido , Honshu, Shikoku, Kyushu, Tsushima, Iki, Fukue, and Okinawajima); there are also visual and echolocation records from the Russian Far East, but further investigation is needed.	Head-body 80-106 mm, tail 45-62 mm, ear 16-23 mm, hindfoot 12-17 mm, forearm 58-64 mm; weight 26-61 g. Pelage of the Bird-like Noctule is velvety and deep yellowish brown throughout, being slightly lighter ventrally. Ears, face, wings, and uropatagium are mid-brown. Thumbs and ears are short, and tragus short and stubby, with a rounded tip; antitragus is long and low. Muzzle is short, with large glands between nostrils and eyes, and ears are short and triangular, with 4-5 folds on outer edge. Calcar is keeled, and wing membraneis attached to ankle. Skull is broad and robust; postorbital process is weak. I? is longer than I’; teeth are large with low cusps; lower molars are nyctalodont. Chromosomal complement has 2n = 42 and FN = 50.	Found primarily in mountainous regions with temperate deciduous forests.	Insectivorous and carnivorous. The Bird-like Noctule is a large, fast aerial hawker that feeds on a variety of insects, as well on occasion as nocturnally migrating birds. In fecal samples collected in Japan , bird bones and feathers were detected, although these bats have never been directly observed hunting birds. The species is known to feed on a variety of insects, including various mayflies, dragonflies, damselflies, grasshoppers, hemipterans, beetles, ichneumonids, winged ants, syrphid flies, chironomid flies, trichopterans, and lepidopterans. In Japan , Coleoptera made up a large portion of the diet, comprising 40-51-2% of arthropod fragments in fecal samples, although Lepidoptera were also significant, ranging 11-34-1% of fragments throughout the year.	Mating takes place in autumn, up to early October, before hibernation; ovulation and fertilization are delayed, and sperm is stored until next spring. Births generally occurin lateJune and earlyJuly;litter size is 1-2 (normally two). Young begin to fly at ¢.40-45 days; females generally start breeding in their first autumn.	The Bird-like Noctule is primarily nocturnal throughout its active months, although it is known to forage during the day when energy demands are high, right after leaving hibernation (March to early April) and just before entering hibernation (October to late November); hibernates in colonies. Diurnal roosts are often found in tree cavities, buildings, bird boxes, and mineshafts. Bats leave the roost in early evening and spend the night foraging high in the forest canopy. Calls are FM for the scanning phase and long duration QCF in the search phase; they have a general frequency range of c¢.17-30 kHz,start frequency range of 27-1-54-1 kHz, and end frequency range of 17-6-22-6 kHz; call durations are 2-2-17-8 milliseconds. Bird-like Noctules are known tolive for at leastsix years. In Japan , the gyrfalcon (Falco rusticolus) has been reported as a predator.	Bird-like Noctules roost in small colonies and probably migrate between breeding and hibernating areas during autumn and spring. During summer and the breeding season, females form large maternal colonies of tens to over 100 individuals, while males form small male-exclusive colonies of 1-10 individuals. During and a little before and after hibernation, however, the species forms colonies consisting of several to tens of individuals of both sexes (October to May).	Classified as Near Threatened on The IUCN Red List. The Bird-like Noctule is relatively widespread and moderately common, although it is threatened by deforestation and other forms of human disturbance.	Abe et al. (2005) | Brazil & Sasaki (2004) | Datzmann et al. (2012) | Fukui, Agetsuma & Hill (2004) | Fukui, Dewa et al. (2013) | Harada, Uchida et al. (1982) | Jo Yeong-Seok et al. (2018) | Maeda (1983) | Mikula et al. (2016) | Ohdachi et al. (2009) | Osawa et al. (2017) | Salgueiro et al. (2007) | Smith & Xie Yan (2008) | Tsytsulina (2008a) | Yoshiyuki (1989)	https://zenodo.org/record/6397788/files/figure.png	11. Bird-like Noctule Nyctalus aviator French: Noctule aviatrice / German: GroRRer Ostasien-Abendsegler / Spanish: Noctulo oriental Other common names: Japanese Large Noctule Taxonomy. Nyctalus aviator Thomas, 1911 , “ Tokyo , Hondo [= Honshu], Japan .” Nyctalus aviator is sometimes included within N. lasiopterus , from which it is morphologically distinct, but to which it is genetically sister. N. aviator and N. lasiopterus are togethersister to N. noctula , all three sharing the same diploid number. Monotypic. Distribution. NE & EC China ( Heilongjiang , Jilin , Henan , Anhui , and Zhejiang ), Korean Peninsula, and numerous islands in Japan ( Hokkaido , Honshu, Shikoku, Kyushu, Tsushima, Iki, Fukue, and Okinawajima); there are also visual and echolocation records from the Russian Far East, but further investigation is needed. Descriptive notes. Head-body 80-106 mm, tail 45-62 mm, ear 16-23 mm, hindfoot 12-17 mm, forearm 58-64 mm; weight 26-61 g. Pelage of the Bird-like Noctule is velvety and deep yellowish brown throughout, being slightly lighter ventrally. Ears, face, wings, and uropatagium are mid-brown. Thumbs and ears are short, and tragus short and stubby, with a rounded tip; antitragus is long and low. Muzzle is short, with large glands between nostrils and eyes, and ears are short and triangular, with 4-5 folds on outer edge. Calcar is keeled, and wing membraneis attached to ankle. Skull is broad and robust; postorbital process is weak. I? is longer than I’; teeth are large with low cusps; lower molars are nyctalodont. Chromosomal complement has 2n = 42 and FN = 50. Habitat. Found primarily in mountainous regions with temperate deciduous forests. Food and Feeding. Insectivorous and carnivorous. The Bird-like Noctule is a large, fast aerial hawker that feeds on a variety of insects, as well on occasion as nocturnally migrating birds. In fecal samples collected in Japan , bird bones and feathers were detected, although these bats have never been directly observed hunting birds. The species is known to feed on a variety of insects, including various mayflies, dragonflies, damselflies, grasshoppers, hemipterans, beetles, ichneumonids, winged ants, syrphid flies, chironomid flies, trichopterans, and lepidopterans. In Japan , Coleoptera made up a large portion of the diet, comprising 40-51-2% of arthropod fragments in fecal samples, although Lepidoptera were also significant, ranging 11-34-1% of fragments throughout the year. Breeding. Mating takes place in autumn, up to early October, before hibernation; ovulation and fertilization are delayed, and sperm is stored until next spring. Births generally occurin lateJune and earlyJuly;litter size is 1-2 (normally two). Young begin to fly at ¢.40-45 days; females generally start breeding in their first autumn. Activity patterns. The Bird-like Noctule is primarily nocturnal throughout its active months, although it is known to forage during the day when energy demands are high, right after leaving hibernation (March to early April) and just before entering hibernation (October to late November); hibernates in colonies. Diurnal roosts are often found in tree cavities, buildings, bird boxes, and mineshafts. Bats leave the roost in early evening and spend the night foraging high in the forest canopy. Calls are FM for the scanning phase and long duration QCF in the search phase; they have a general frequency range of c¢.17-30 kHz,start frequency range of 27-1-54-1 kHz, and end frequency range of 17-6-22-6 kHz; call durations are 2-2-17-8 milliseconds. Bird-like Noctules are known tolive for at leastsix years. In Japan , the gyrfalcon (Falco rusticolus) has been reported as a predator. Movements, Home range and Social organization. Bird-like Noctules roost in small colonies and probably migrate between breeding and hibernating areas during autumn and spring. During summer and the breeding season, females form large maternal colonies of tens to over 100 individuals, while males form small male-exclusive colonies of 1-10 individuals. During and a little before and after hibernation, however, the species forms colonies consisting of several to tens of individuals of both sexes (October to May). Status and Conservation. Classified as Near Threatened on The IUCN Red List. The Bird-like Noctule is relatively widespread and moderately common, although it is threatened by deforestation and other forms of human disturbance. Bibliography. Abe et al. (2005), Brazil & Sasaki (2004), Datzmann et al. (2012), Fukui, Agetsuma & Hill (2004), Fukui, Dewa et al. (2013), Harada, Uchida et al. (1982), Jo Yeong-Seok et al. (2018), Maeda (1983), Mikula et al. (2016), Ohdachi et al. (2009), Osawa et al. (2017), Salgueiro et al. (2007), Smith & Xie Yan (2008), Tsytsulina (2008a), Yoshiyuki (1989).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Nyctalus aviator	Nyctalus		aviator	Thomas	1911	1	Ann. Mag. Nat. Hist.	ser.8, 8: 380	Birdlike Noctule	 molossus Temminck, 1840 [not Pallas, 1767].	Japan, Honshu, Tokyo.	Hokkaido, Shikoku, Kyushu, Tsushima, Iki (Japan); Korea; E and C China. Possibly occurs in Russian Far East, see Tiunov (1997).	Not listed.	Near Threatened	Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951), but see Corbet (1978 c ) and Yoshiyuki (1989).Reviewed by Yoshiyuki (1989).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nyctalus aviator	23	Bird-like Noctule	Japanese Large Noctule	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Nyctalus	NA	aviator	O. Thomas	1911	0	Nyctalus_aviator	Thomas, O. (1911). Two new eastern bats. Annals and Magazine of Natural History, ser. 8, 8, 380.	https://www.biodiversitylibrary.org/item/53331#page/406/mode/1up	BM 1905.1.4.5		"Tokyo, Hondo [= Honshu], Japan."			molossus (Temminck, 1840) [preoccupied]|aviator O. Thomas, 1911	NA	NA	China|North Korea|South Korea|Russia?|Japan	Asia	Palearctic	NT	0	0	0	Nyctalus_aviator	0	sciname match	Nyctalus_aviator	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14921	Nyctalus aviator	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Nyctalus	aviator	(Thomas, 1911)	This species has previously been considered as a subspecies of lasiopterus .	20000000	Nyctalus aviator	Near Threatened	A2c	2019	2018-07-23 00:00:00 UTC	3.1	English	The species is listed as Near Threatened (and is approaching criterion A2c) as it is suspected to be declining at a rate 20-30% over the next 18 years (3 generations based on estimated generation length of 6 years; Maeda 1974) across it ranges as a result of habitat loss and human disturbance of sensitive tree-cavity roosts. It is a naturally rare species that is dependent on forests. It forms large nursery colonies that are highly vulnerable to disturbance. An additional assessment is warranted once better population trends and generation lengths are quantified.	Roosts mainly in tree cavities, but occasionally uses slits under elevated bridges, buildings and bird boxes (Fukui 2015). Environments around roosts vary from forest to urban areas. Feeding habitats are unknown. It is nocturnal but often shows day-time feeding mainly in early winter. It mainly feeds on Coleoptera, Lepidoptera, Trichoptera, Ephemeroptera and Hemiptera (Fukui et al.    2013). In spring and from autumn to early winter, this species regularly preys on small birds of Passeriformes (Fukui et al.    2013). From June to August, females form maternity colonies (Maeda 1974). The size of a maternity colony varies from tens to more than a hundred individuals. During the period when females are involved in maternity colonies, males roost in solitary or males form small colonies consisting of several to tens of individuals (Maeda 1974). Parturition occurs during late June to early July in Iwate and Hokkaido (Maeda 1972). Litter size is sometimes 1 but usually 2. Forearm length at birth and its ratio to mother are 22.1 mm (range = 20.0–27.0; n = 10) and 0.36 (n = 10) on average respectively; body weight at birth is 6.3 g on average (5.4–8.3; n = 9), and increases to 25 g up to 4 weeks of age; young begin to fly at about 40–45 days of age (Maeda 1972). Yearling females copulate in the first autumn but yearling males do not; the testes of adult males reach a maximum size in July and this state continues until the beginning of October (Maeda 1974).	Deforestation and human disturbance are considered the major threats in Japan.	Most of the records over the last 30 years in Japan have been reported from Hokkaido, Honshu, Iki Island, Fukue Island, and Okinawa-jima Island. Population in NE China cannot be estimated even approximately, due to poor knowledge about bats of that area. The species is suspected to be declining at a rate of 20-30% over the next 18 years (three generations based on estimated generation length of 6 years; Maeda 1974) across it ranges as a result of habitat loss and human disturbance of sensitive tree-cavity roosts.	Decreasing	Palaearctic, marginally Oriental. This species is known from eastern China (provinces of Heilongjiang, Jilin, Henan, Zhejiang, and Anhui (Smith and Xie 2013), the Korean Peninsula (Yoon 2010) and Japan, where there are records from Hokkaido, Honshu, Shikoku, Kyushu and Isls of Iki, Fukue, Okinawa-jima and Tsushima (Fukui 2015). In the Russian Far East, it is known from several doubtful observations, there are no significant evidences for presence of this bat in Russian fauna (Tiunov 1997), however, there is high possibility for its occurrence at least as vagrant.		Terrestrial	It is listed as Near Threatened (NT) in the Chinese Red List, Vulnerable (VU) in the Japanese Red List (Ministry of the Environment 2014), and as a Rare Species in Russia.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Nyctalus		aviator	Thomas	1911	1	Ann. Mag. Nat. Hist.	ser.8, 8: 380	Birdlike Noctule	 molossus Temminck, 1840 [not Pallas, 1767].	Japan, Honshu, Tokyo.	Hokkaido, Shikoku, Kyushu, Tsushima, Iki (Japan); Korea; E and C China. Possibly occurs in Russian Far East, see Tiunov (1997).	Not listed.	Near Threatened	Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951), but see Corbet (1978 c ) and Yoshiyuki (1989).Reviewed by Yoshiyuki (1989).	Nyctalus aviator	1005601	23	Bird-like Noctule	Japanese Large Noctule	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Nyctalus	NA	aviator	O. Thomas	1911	0	Nyctalus_aviator	Thomas, O. (1911). Two new eastern bats. Annals and Magazine of Natural History, ser. 8, 8, 380.	https://www.biodiversitylibrary.org/item/53331#page/406/mode/1up	BM 1905.1.4.5		"Tokyo, Hondo [= Honshu], Japan."			molossus (Temminck, 1840) [preoccupied]|aviator O. Thomas, 1911	NA	NA				China|North Korea|South Korea|Russia?|Japan	Asia	Palearctic	NT	0	0	0	Nyctalus_aviator	0	sciname match	Nyctalus_aviator	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nyctalus_aviator	1005601	23	Bird-like Noctule	Japanese Large Noctule	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Nyctalus	NA	aviator	O. Thomas	0	Nyctalus aviator	Thomas, O. 1911-08-29. Two new eastern bats. Annals and Magazine of Natural History (8)8(45):378-380.	https://www.biodiversitylibrary.org/page/22099043	BMNH:Mamm:1905.1.4.5	holotype	https://data.nhm.ac.uk/object/2353a09a-2c63-4c82-ad6f-23492c912c48	"Tokyo, Hondo [= Honshu], Japan."			NA	NA				China|North Korea|South Korea|Russia?|Japan	Asia	Palearctic	NT	0	0	0	Nyctalus_aviator	0	sciname match	Nyctalus_aviator	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Nyctalus		aviator	Thomas	1911	0	Ann. Mag. Nat. Hist.	ser.8, 8: 380	Birdlike Noctule	molossus Temminck, 1840 [not Pallas, 1767].	Japan, Honshu, Tokyo.	Hokkaido, Shikoku, Kyushu, Tsushima, Iki (Japan); Korea; E and C China. Possibly occurs in Russian Far East, see Tiunov (1997).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14921/22016483/' target='_blank'>Near Threatened</a>	Listed as a subspecies of lasiopterus by Ellerman and Morrison-Scott (1951), but see Corbet (1978c) and Yoshiyuki (1989).Reviewed by Yoshiyuki (1989).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Nyctalus aviator; Nyctalus aviator; Nyctalus aviator; Nyctalus aviator; Nyctalus aviator; Nyctalus aviator; molossus; molossus; molossus; aviator; Noctule aviatrice; Gro RRer Ostasien-Abendsegler; Noctulooriental; Japanese Large Noctule; Bird-like Noctule; Japanese Large Noctule; Birdlike Noctule; Birdlike Noctule; N. aviator